Family Schizaeaceae, genus Anemia;
world species list.
By Michael Hassler and Brian Swale.
A list of the species of the genus Anemia, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.
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005.00 Class Filicopsida
Trivial names: Ferns
Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species
[11501 species and 454 hybrids listed.
015.0000 Family Schizaeaceae Kaulfuss; Das Wesen d. Farrnkr. 119 (1827)
Trivial names: Curly-grass family
Statistics: 5 genera, 184-190 spp. [195 species and 9 hybrids listed]
Revision date: 20.07.2001
= Lygodiaceae C. Presl (1845)
= Anemiaceae Link (1833)
= Mohriaceae (C. Presl) Reed (1948)
Revision date: 20.07.2001
015.0003 Anemia Sw.; Syn. Fil. 6: 155 (1806) [nom. cons.
Trivial names: Flowering Ferns
Statistics: 110-115 spp., of them 70 Brazil, 20 Mexico, few in Africa/Madagascar,
1 species in peninsular India [116 species and 6 hybrids listed]
Revision date: 20.07.2001
= Aneimia Kaulf.; Enum. 51 (1824)
= Aneimiaebotrys Fée; Crypt. vasc. Brésil 1: 267 (1869)
= Anemiaebotrys Fée; Crypt. Vasc. Bresil 1: 267 (1869)
= Anemidictyon J. Sm.; Hook., Gen. Fil. t. 103 (1842)
= Anemirhiza J. Sm.; Seem., Bot. Voy. Herald 243 (1854)
= Coptophyllum Gardner (1842)
= Hemianeimia Prantl; Schiz. 86 (1881)
= Hemianemia (Prantl) Reed; Bol. Soc. Broter. II, 21: 154 (1947)
= Ornithopteris Bernh.; Neues Jour. Bot. (Schrad.) 1 (2): 40 (1806)
= Spathepteris Presl; Suppl. Tent. Pterid. 95 (1845)
= Trochopteris Gardner; Lond. Jour. Bot. 1: 74 (1842)
Anemia abbottii Maxon; Proc. Biol. Soc. Wash. 35. 48 (1922)
Remarks: Subgen. Anemirhiza
Anemia adiantifolia (L.) Swartz; Syn. Fil. 157 (1806)
Trivial names: Pine Fern
Remarks: Subgen. Anemirhiza
Anemia affinis Bak.; Syn. 525 (1874)
Remarks: Subgen. Anemia
Anemia alfredi-rohrii Brade; Bradea 1: 27, t.2 (1970)
Anemia alternifolia Mickel; Brittonia 33(3): 424 (1981)
Anemia angolensis Alston; Mendonca, Estud. Ens. Docum. Junta Invest. Ultramar 12: 9. t.2,
3(A) (1954)
Anemia antrorsa Mickel; Amer. Fern J. 75(2): 33 (1985)
Anemia aspera (Fée) Bak.; Journ. Linn. Soc. 14. 27 (1873)
Remarks: Subgen. Coptophyllum
Anemia aurita Sw.; Syn. 157 (1806)
Remarks: Subgen. Anemirhiza
Anemia ayacuchensis Mickel; Amer. Fern J. 75(2): 36 (1985)
Anemia barbatula Christ; Denkschr. Akad. Wien 79. 40 (1907)
Anemia bartlettii Mickel; lowa State J. Sci. 36: 420, t.26B (1962)
Anemia blackii Brade; Arq. Jard. Bot. Rio de Janeiro 18: 25 (1965)
Anemia blechnoides Smith in Rees; Cyclop. 39: Anemia no. 3 (1819)
Anemia brandegeea Dav.; Fern Bull. 13. 20 c. tab (1905)
Anemia buniifolia (Gardn.) Moore; Ind. CXVI (1857)
Anemia cajalbanica Borhidi & Mz.
Anemia candidoi Brade; Arq. Jard. Bot. Rio Janeiro 11: 34. t.13 (1951)
Anemia cicutaria Kze.; Spreng., Syst. 4. 31 (1827)
Anemia cipoensis Sehnem; Fl. Ilustr. Catar. 1 (Esquizeáceas), 40 (1974)
Anemia clinata Mickel; Amer. Fern. J. 56: 58, t. 4 (1966)
Anemia colimensis Mickel; Iowa State J. Sci. 36: 434, t.32B (1962)
Remarks: Subgen. Anemirhiza
Anemia collina Raddi; Opusc. sci. Bot. 3. 282 (1819)
Anemia coriacea Gris.; Cat. Pl. Cub. 272 (1866)
ssp. coriacea Gris.; Cat. Pl. Cub. 272 (1866)
f. nipeensis (Benedict) Mickel; Brittonia 33(3): 426 (1981)
var. immeensis Duek; Feddes Repert. 86(9-10): 523 (1975)
ssp. moaensis Borhidi & Mz.
Anemia costata Sehnem; Fl. Ilustr. Catar. 1 (Esquizeáceas), 59 (1974)
Anemia cuneata Kze.; Spreng. Syst. 4. 32 (1827)
Anemia damazii Christ; Bull. Boiss. II. 7. 792 (1907)
Anemia dardanoi Brade in A. Lima; Bol. Técn. Inst. Agron. Norte 36: 127 (1959)
Anemia dimorphostachys Bak.; Engl. Jahrb. 17. 522 (1893)
Anemia distans Fée; 9 mem. 41 (33) (1857)
Anemia diversifolia Schrad.; Gött. gel. Anz. 864 (1824)
Anemia donnell-smithii Maxon; N. Amer. Fl. 16. 43 (1909)
Anemia dregeana Kze.; Linn. 10. 493 (1836)
Anemia duartei Brade; Bradea 1: 27, t.1 (1979)
Anemia elaphoglossoides Mickel; Amer. Fern J. 74(4): 111 (1984)
Anemia elegans (Gardn.) Presl; Suppl. 81 (1845)
Remarks: Subgen. Coptophyllum
Anemia eximia Taubert; Engl. Jahrb. 21. 422 (1896)
Anemia familiaris Mickel; Brittonia 34(4): 399 (1982)
Anemia ferruginea H.B.K.; Nov. Gen. & Sp. 1. 32 (1815)
var. ahenobarba (Christ) Mickel
Anemia flexuosa (Sav.) Sw.; Syn. 156 (1806)
Remarks: Subgen. Coptophyllum
Anemia gardneri Hook.; lc. pl. t. 190 (1837)
Anemia glareosa Gardn.; Field. & Gardn., Sert. pl. t. 70 (1844)
Remarks: Subgen. Coptophyllum
Anemia gomesii Christ; Bull. Boiss. II. 7. 791 (1907)
Anemia guatemalensis Maxon; N. Amer. Fl. 16. 46 (1909)
Remarks: Subgen. Coptophyllum
Anemia hatschbachii Sehnem; Fl. Ilustr. Catar. 1 (Esquizeáceas), 27 (1974)
Anemia herzogii Rosenst.; Meded. Rijks Herb. Leiden nr. 19. 24 (1913)
Anemia heterodoxa Christ; Ann. Cons. Jard. bot. Geneve 3. 45 (1899)
Anemia hirsuta (L.) Sw.; Syn. 156 (1806)
Remarks: Subgen. Anemia
Anemia hirta (L.) Sw.; Syn. 155 (1806)
Remarks: Subgen. Anemia
Anemia imbricata Sturm; Fl. bras. 1(2). 205 (1859)
Anemia incisa Schrad.; Gött. gel. Anz. 865 (1824)
Anemia intermedia Copel. ex M.E.Jones; Contr. West. Bot. 16: 123 (1929)
Remarks: unverified name
Anemia jaliscana Maxon; N. Amer. Fl. 16. 44 (1909)
Anemia karwinskyana (Presl) Prantl; Unters. Morph. Gefäßkrypt. 2: 99 (1881)
Remarks: Subgen. Coptophyllum
Anemia lancea Christ; Bull. Boiss. II. 7. 791 (1907)
Anemia lanipes C. Chr.; Cat. Pl. Mad. Pter. 65 (1931) (nomen), Dansk Bot. Ark. 7. 177 t. 71 (1932)
Anemia lanuginosa Bongard; Sturm, Fl. Bras. 12. 210 (1859)
Anemia laxa Lind.; Ark. för Bot. 1. 261 t. 13 (1903)
Anemia luetzelburgii Rosenst.; Fedde Rep. 20. 94 (1924)
Anemia madagascariensis C. Chr.; Arch. Bot. (Caen) 2 Bull. mens. 216 (1928)
Anemia mandioccana Raddi; Opusc. sci. Bol. 8. 282 (1819)
Remarks: Subgen. Anemia
Anemia marginata Mickel; Amer. Fern J. 74(4): 111 (1984)
Anemia mexicana Klotzsch; Linnaea 18: 526 (1844)
Trivial names: Mexican Flowering Fern
Remarks: Subgen. Anemirhiza
var. makrinii (Maxon) Mickel; Brittonia 33(3): 421 (1981)
Anemia millefolia (Gardner) C. Presl; Suppl. 80 (1845)
Anemia mirabilis Brade; Sellowia 17: 53, f. 2 (1965)
Anemia multiplex Mickel; Brittonia 34(4): 405 (1982)
Anemia munchii Christ; Bull. Boiss. II. 7. 792 (1907)
Anemia myriophylla Christ; Bull. Boiss. II. 7. 793 (1907)
Remarks: Subgen. Coptophyllum
Anemia nana Bak.; Engl. Jahrb. 17. 522 (1893)
Anemia nervosa Pohl; Sturm, Fl. bras. 12. 193 (1859)
Anemia nigerica Alston; Bol. Soc. Broter. ser. 2. 30: 6 (1956)
Anemia oblongifolia (Cav.) Sw.; Syn. 156 (1806)
Remarks: Subgen. Anemia
Anemia obovata (Underw.) Maxon; N. Amer. Fl. 16. 42 (1909)
Anemia organensis Rosenst.; Fedde Rep. 20. 95 (1924)
Remarks: Subgen. Coptophyllum
Anemia ouropretana Christ; Schwacke, Pl. Nov. Mineiras 2. 36 (1900)
Remarks: Subgen. Anemia
Anemia pallida Gardn.; Field. & Gardn., Sert. pl. ad t. 70 (1844)
Anemia palmarum Liebm.; Ark. för Bot. 1. 261 t. 14 (1903)
Anemia pastinacaria Moritz ex Prantl; Schiz. 110 (1881)
Anemia perrieriana C. Chr.; Cat. Pl. Mad. Pter. 65 (1931) [nomen
Anemia phyllitidis (L.) Sw.; Syn. 155 (1806)
Remarks: Subgen. Anemia
var. tweediana (Hook.) Hassl.
Anemia pinnata Sehnem; Fl. Illustr. Catar. 1 (Esquizeáceas), 21 (1974)
Anemia pohliana Sturm; Fl. bras. 1(2): 195 (1859)
Anemia porrecta Mickel; Amer. Fern J. 75(2): 36 (1985)
Anemia portoricensis Maxon; N. Amer. Fl. 16. 48 (1909)
Remarks: Subgen. Anemirhiza
Anemia proxima C. Chr.; Kungl. Svenska Vetens.-akad. Handl. ser. 3. 16(2): 86.
t.20(1-2) (1937)
Anemia pulchra Pohl; Prantl, Schiz. 109 (1881)
Anemia pumilio Mickel; Brittonia 33(3): 428 (1981)
Anemia pyrenaea Taubert; Engl. Jahrb. 21: 422 (1896)
Anemia radicans Raddi; Opusc. sci. Bol. 3: 282 (1819)
Anemia retroflexa Brade; An. Prim. Reun. Sul-Amer. Bot. 2: 9. t.5 (1940)
Anemia rigida Sehnem; Fl. Ilustr. Catar. 1 (Esquizeáceas), 36 (1974)
Anemia rotundifolia Schrad.; Gött. gel. Anz. 865 (1824)
Anemia rutifolia Mart.; Ic. Cr. Bras. 112, t. 55 (1834)
Anemia salvadorensis Mickel & Seiler; Amer. Fern J. 74(4): 113 (1984)
Anemia sanctae-martae Christ; Bull. Boiss. II. 7. 791 (1907)
Anemia schaeferi Sehnem; Fl. Ilustr. Catar. 1 (Esquizeáceas), 26 (1974)
Anemia schimperiana Presl; Suppl. 84 (1845)
Anemia semihirsuta Mickel (1982)
Anemia sessilis (Jeanpert) C. Chr.; Fedde Repert. 9. 371 (1911)
Anemia simii Tardieu; Notul. Syst. [Paris] 14: 208 (1952)
Anemia simplicior (Christ) Mickel; Iowa State J. Sci. 36: 418, t.25B (1962)
Remarks: Subgen. Coptophyllum
Anemia smithii Brade; Bol. Mus. Nac. Rio Janeiro 6. 95 t. 3 (1929)
Anemia speciosa Presl; Suppl. 89 (1845)
Anemia spicantoides Mabb.; Taxon 32(1): 81 (1983) [nom. nov.
Anemia tenella (Cav.) Sw.; Syn. 156 (1806)
Remarks: Subgen. Anemia
Anemia tenera Pohl; Sturm, Fl. bras. 1(2): 208 (1859)
Anemia tomentosa (Sav.) Sw.; Syn. 157 (1806)
var. australis Mickel
var. anthriscifolia (Schrad.) Mickel
var. mexicana (Presl) Mickel; Iowa State College J. Sci. 36: 427 (1962)
Anemia trichorhiza Gardn.; Hook., Ic. pl. t. 876 (1852)
Anemia tripinnata Copel.; Univ. Calif. Publ. Bot. 17. 24 t. 1 (1932)
Anemia ulei Christ; Schwacke, Pl. Nov. Mineiras 2: 36 (1900)
Anemia underwoodiana Maxon; N. Amer. Fl. 16. 40 (1909)
Anemia villosa Humb. & Bonpl. ex Willd.; Sp. 6. 92 (1810)
Remarks: Subgen. Coptophyllum
var. karwinskyana Presl; Suppl. Tent. Pterid. 83 (1845)
Anemia voerkeliana Duek; Feddes Repert. 86(9-10): 521 (1975)
Anemia warmingii Prantl; Schiz. 113 (1881)
Anemia wettsteinii Christ; Denkschr. Akad. Wien 79. 48 t. 9 f. 3-6 (1907)
Anemia wrightii Baker; Hook. & Baker, Syn. Fil. 10: 435 (1868)
Trivial names: Wright's Flowering Fern
Remarks: Subgen. Anemirhiza
Anemia x didicusana L. D. Gómez; Bresnia 18. 156 (1980)
Remarks: Anemia hirsuta x phyllitidis
Anemia x espiritosantensis Brade; Rodriguesia 10(21): 27. t.1 (1948)
Remarks: Anemia collina x tenella
Anemia x paraphyllitidis Mickel; Brittonia 34(4): 407 (1982)
Remarks: Anemia phyllitidis x semihirsuta
Anemia x recondita Mickel; Brittonia 34(4): 409 (1982)
Remarks: Anemia hirsuta x jaliscana
Anemia x semihirsuta Mickel; Brittonia 34(4): 410 (1982)
Remarks: Anemia hirsuta x phyllitidis
Anemia x zanonii Mickel; Moscosoa 3: 84 (1984)
Remarks: Anemia hirsuta x underwoodiana
This page is Copyright ©, 2001, 2002 Brian Swale, Michael Hassler. All rights reserved.
For more information, go to © Copyright page
For each species, information on its distribution by country and sometimes by province,
region or state within a country; conservation status (where applicable), and all synonyms with publishing
source (other latin names by which the species has been known) that we could find, and where applicable
whether or not the given name has been verified; are available on the CDROM version of this page.
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