Family Lomariopsidaceae, genus Arachniodes;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Arachniodes, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

005.00 Class Filicopsida
		Trivial names: Ferns
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
047.000 Order Dryopteridales
		Statistics: 3 families, 30 genera, 1421-1752 spp. 
		[1989 species and 158 hybrids listed]
049.0000 Family Dryopteridaceae Ching; Acta Phytotax. Sin. 10: 1 (1965) s. str.
		Statistics: 21 genera, 672-939 spp. 
		[1162 species and 147 hybrids listed]
		Revision date: 20.07.2001
		= Aspidiaceae Frank; Leunis, Syn. Pflanzenfam. ed. 2, 3: 1469 [nom. illeg.
		= Dryopteridaceae tribus Stenolepieae H.S. Kung; Proc. Int. Symp. Syst. Pteridol. 
		(eds. K.H. Shing & K.U. Kramer): 172 (1989)
049.0200 Tribus Dryopterideae

049.0207 Arachniodes Blume; Enum. Pl. Javae 2: 241 (1828) Statistics: 100-140 spp. mostly China and E-Asia, 3-4 New World [149 species and 11 hybrids listed]. Many of the Chinese species are probably invalid. Revision date: 20.07.2001 = Byrsopteris Morton; Amer. Fern Journ. 60: 149 (1960) = Leptorumohra H. Ito; Nakai & Honda, Fl. Nov. Jap. 4: 101 (1939) = Lithostegia Ching; Sinensia 4: 2 (1933) [fide Fraser-Jenkins 1997 = Polystichopsis (J. Smith) Holtt.; Amer. Fern Jour. 50: 147-148 (1947) = Polystichopsis J. Sm. emend. C. Chr.; Vid. Selsk. Skr. VIII. 6. 101 (1920) = Rumohra sect. Leptorumohra H. Ito; Nakai & Honda, Fl. Nov. Jap. 11: 579 (1935) Arachniodes acuminata Ching & Wang; Acta Phytotax. Sin. 9: 367 (1964) Arachniodes ailaoshanensis Ching; Bull. Bot. Res., Harbin 6(3): 60 (1986) Arachniodes amabilis (BI.) Tindale; Contr. N. S. Wales Nation Herb. 3 90 (1961) var. fimbriata K. lwats.; Acta Phytotax. Geobot. 30(1-3): 42 (1979) var. okinawensis (Nakaike) Seriz.; J. Phytogeogr. Taxon. 29(1): 24 (1981) Arachniodes amoena (Ching) Ching; Acta Bot. Sin. 10: 256 (1962) Arachniodes anshunensis Ching & Y.T.Hsieh; Bull. Bot. Res., Harbin 6(3): 67 (1986) Arachniodes argillicola (Proctor) Proctor; Amer. Fern J. 72(4): 109 (1982) Arachniodes aristata (Forst. f.) Tindale; Contr. N. S. Wales Nation Herb. 3 89 (1961) Trivial names: Prickly Shield fern, East Indian holly fern Arachniodes aristatissima Ching; Bull. Bot. Res., Harbin 6(3): 1 (1986) Arachniodes assamica (Kuhn) Ohwi; J. Jap. Bot. 37: 76 (1962) Arachniodes attenuata Ching; Bull. Bot. Res., Harbin 6(3): 2 (1986) Arachniodes austroyunnanensis Ching; Bull. Bot. Res., Harbin 6(3): 3 (1986) Arachniodes baiseensis Ching; Bull. Bot. Res., Harbin 6(3): 25 (1986) Arachniodes basipinnata (Ching) Ching ex Y.T.Hsieh; Bull. Bot. Res., Harbin 11(3): 27 (1991) Arachniodes bella (C. Chr.) Ching; Acta Bot. Sin. 10: 261 (1962) Arachniodes borealis Seriz.; J. Jap. Bot. 61(2): 48 (1986) f. ciliata (Nakaike) Seriz.; J. Jap. Bot. 61(2): 52 (1986) Arachniodes cantilenae Sa. Kurata; J. Geobot. 25(4): 98 (1978) Arachniodes caudata Ching; Acta Phytotax. Sin. 9: 384 (1964) Arachniodes caudifolia Ching & Y.T.Hsieh; Bull. Bot. Res., Harbin 4(2): 104 (1984) Arachniodes cavalerii (Christ) Ohwi; J. Jap. Bot. 37: 76 (1962) Arachniodes centrochinensis Ching; Fl. Tsinlingensis 2: 229 (1974) Arachniodes chaerophylloides (Poir.) Proctor; Amer. Fern J. 72(4): 109 (1982) Arachniodes chinensis (Rosenstock) Ching; Acta Bot. Sin. 10: 257 (1962) Arachniodes chingii Y.T.Hsieh; Bull. Bot. Res., Harbin 6(4): 4 (1986) Arachniodes coniifolia (Moore) Ching; Acta Bot. Sin. 10: 257 (1962) Arachniodes cornopteris Ching; Bull. Bot. Res., Harbin 6(3): 4 (1986) Arachniodes costulisora Ching; Bull. Bot. Res., Harbin 6(3): 62 (1986) Arachniodes cyrtomifolia Ching; Bull. Bot. Res., Harbin 6(3): 31 (1986) Arachniodes damiaoshanensis Y.T.Hsieh; Bull. Bot. Res., Harbin 6(4): 6 (1986) Arachniodes davalliaeformis (Christ) Nakaike; New Fl. Japan, Pterid.: 839 (1992) Arachniodes dayaoensis Y.T.Hsieh; Acta Bot. Yunnan. 5(1): 57 (1983) Arachniodes decomposita Ching; Bull. Bot. Res., Harbin 6(3): 49 (1986) Arachniodes denticulata (Sw.) Ching; Acta Bot. Sin. 10: 260 (1962) ssp. denticulata (Sw.) Ching; Acta Bot. Sin. 10: 260 (1962) ssp. laetum (Sw.) comb. ined. Arachniodes diffracta (Baker) Ching; Acta Bot. Sin. 10: 257 (1962) Arachniodes dimorphophylla (Hayata) Ching; Acta Bot. Sin. 10: 257 (1962) Arachniodes elevatus Ching; Bull. Bot. Res., Harbin 6(3): 40 (1986) Arachniodes emeiensis Ching; Bull. Bot. Res., Harbin 6(3): 5 (1986) Arachniodes erythrosora Ching; Bull. Bot. Res., Harbin 6(3): 42 (1986) Arachniodes falcata Ching; Bull. Bot. Res., Harbin 6(3): 7 (1986) Arachniodes fengii Ching; Bull. Bot. Res., Harbin 6(3): 8 (1986) Arachniodes festina (Hance) Ching; Acta Bot. Sin. 10: 257 (1962) Arachniodes foeniculacea Ching; Bull. Bot. Res., Harbin 6(3): 45 (1986) Arachniodes formosa (Fée) Ching; Acta Bot. Sin. 10: 260 (1962) Arachniodes futeshanensis Y.T.Hsieh; Bull. Bot. Res., Harbin 6(4): 5 (1986) Arachniodes gigantea Ching; Bull. Bot. Res., Harbin 6(3): 66 (1986) Arachniodes gijiangensis Ching; Bull. Bot. Res., Harbin 6(3): 33 (1986) Arachniodes gizushanensis Ching; Bull. Bot. Res., Harbin 6(3): 41 (1986) Arachniodes globisora (Hayata) Ching; Acta Phytotax. Sin. 9: 383 (1964) Arachniodes gongshanensis Ching & Y.T.Hsieh; Bull. Bot. Res. Harbin 6(3): 68 (1986) Arachniodes gradata Ching; Bull. Bot. Res., Harbin 6(3): 39 (1986) Arachniodes grossa (Tard. & C. Chr.) Ching; Acta Bot. Sin. 10: 257 (1962) Arachniodes guangnanensis Y.T.Hsieh; Bull. Bot. Res., Harbin 4(2): 106 (1984) Arachniodes guangtongensis Ching; Bull. Bot. Res., Harbin 6(3): 58 (1986) Arachniodes guangxiensis Ching; Bull. Bot. Res., Harbin 6(3): 27 (1986) Arachniodes guanxianensis Ching; Bull. Bot. Res., Harbin 6(3): 50 (1986) Arachniodes hainanensis (Ching) Ching; Acta Bot. Sin. 10: 258 (1962) Arachniodes haniffii (Holtt.) Ching; Acta Bot. Sin. 10: 258 (1962) Arachniodes hekiana Kurata; J. Geobot. 13: 99 (1965) Arachniodes hekouensis Ching; Bull. Bot. Res., Harbin 6(3): 57 (1986) Arachniodes henryi (Christ) Ching; Acta Bot. Si.n. 10: 258 (1962) Arachniodes hiugana Sa.Kurata; J. Geobot. 25(4): 97 (1978) Arachniodes huapingensis Ching & P.S.Chiu; Bull. Bot. Res., Harbin 6(3): 53 (1986) Arachniodes hunanensis Ching; Bull. Bot. Res., Harbin 6(3): 10 (1986) Arachniodes hupingshanensis S.F.Wu; Keys Vasc. Pl. Wuling Mount. (ed. W.T.Wang & al.): 572 (1995) Arachniodes insularis W.H.Wagner; Contr. Univ. Michigan Herb. 19: 63 (1993) Arachniodes ishingensis Ching & Y.T.Xie; Y.T.Xie, Acta Phytotax. Sin. 22(2): 161 (1984) Arachniodes jiangxiensis Ching; Bull. Bot. Res., Harbin 6(3): 43 (1986) Arachniodes jinfushanensis Ching; Bull. Bot. Res., Harbin 6(3): 11 (1986) Arachniodes jingdungensis Ching; Bull. Bot. Res., Harbin 6(3): 64 (1986) Arachniodes jinpingensis Y.T.Hsieh; Acta Bot. Yunnan. 5(1): 55 (1983) Arachniodes kansuensis (Ching) Y.T.Hsieh; Bull. Bot. Res., Harbin 4(2): 109 (1984) Arachniodes lanceolata Y.T.Hsieh; Bull. Bot. Res., Harbin 11(2): 4 (1991) Arachniodes leuconeura Ching; Bull. Bot. Res., Harbin 6(3): 12 (1986) Arachniodes leucostegioides (C. Chr.) Ching; Acta Bot. Sin. 10: 260 (1962) Arachniodes liyangensis Ching & Y.C.Lan; Jiang su zhi wu zhi [Rec. Fl. Jiangsu (Kiangsu) Prov.] 1: 466 (1977) Arachniodes longipinna Ching; Bull. Bot. Res., Harbin 6(3): 13 (1986) Arachniodes lurida (Jenman ex Underw. & Maxon) Proctor; Amer. Fern J. 72(4): 109 (1982) Arachniodes lushuiensis Y.T.Hsieh; Bull. Bot. Res., Harbin 4(2): 108 (1984) Arachniodes macrostegia (Hook.) R. M. Tryon & D.S.Conant; Acta Amazon. 5(1): 29 (1975) Arachniodes maguanensis Ching & Y.T.Hsieh; Bull. Bot. Res., Harbin 6(4): 2 (1986) Arachniodes maoshanensis Ching; Bull. Bot. Res., Harbin 6(3): 54 (1986) Arachniodes maxima (Baker) Brownlie; Nova Hedwigia Beih. 55 [Pterid. Fl. Fiji]: 280 (1977) Arachniodes menglianensis Y.T.Hsieh; Bull. Bot. Res., Harbin 11(2): 3 (1991) Arachniodes mengziensis Ching; Bull. Bot. Res., Harbin 6(3): 14 (1986) Arachniodes michelii (H.Lev.) Ching ex Y.T.Hsieh; Bull. Bot. Res., Harbin 11(3): 27 (1991) Arachniodes miqueliana (Maxim. ex Franchet & Savat.) Ohwi; J. Jap. Bot. 37: 76 (1962) Arachniodes multifida Ching; Bull. Bot. Res., Harbin 6(3): 15 (1986) Arachniodes mutica (Franchet & Savat.) Ohwi; J. Jap. Bot. 37: 76 (1962) Arachniodes nanchuanensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 21 (1984) Arachniodes nanqingensis Ching; Bull. Bot. Res., Harbin 6(3): 38 (1986) Arachniodes neoaristata Ching; Bull. Bot. Res., Harbin 6(3): 34 (1986) Arachniodes nibashanensis Y.T.Hsieh; Bull. Bot. Res., Harbin 6(4): 7 (1986) Arachniodes nigrospinosa (Ching) Ching; Acta Bot. Sin. 10: 258 (1962) Arachniodes nipponica (Rosenst.) Ohwi; Jap. Bot. 37: 76 (1962) Arachniodes nitidula Ching; Bull. Bot. Res., Harbin 6(3): 59 (1986) Arachniodes obtusiloba Ching & Wang; Acta Phytotax. Sin. 9: 369 (1964) Arachniodes obtusipinnula Ching & Y.T.Yie; Acta Phytotax. Sin. 22(2): 160 (1984) Arachniodes ochropteroides (Baker) Lellinger; Amer. Fern J. 77: 101 (1987) Arachniodes oosorae H.Itô; J. Jap. Bot. 66(4): 191 (1991) Arachniodes pianmaensis Ching; Bull. Bot. Res., Harbin 6(3): 65 (1986) Arachniodes pseudoassamica Ching; Bull. Bot. Res., Harbin 6(3): 16 (1996) Arachniodes pseudolongipinna Ching; Bull. Bot. Res., Harbin 6(3): 17 (1986) Arachniodes pseudorepens Nakaike; J. Geobot. 18, 36 (1970) Arachniodes pseudosimplicior Ching; Bull. Bot. Res., Harbin 6(3): 47 (1986) Arachniodes pubescens (L.) Proctor; Amer. Fern J. 72(4): 109 (1982) Arachniodes puncticulata (Alderw.) Ching; Acta Bot. Sirk. 10: 259 (1962) Arachniodes quadripinnata (Hayata) Seriz.; J. Jap. Bot. 61(2): 53 (1986) ssp. quadripinnata (Hayata) Seriz.; J. Jap. Bot. 61(2): 53 (1986) ssp. fimbriata (Koidz.) Seriz.; J. Jap. Bot. 61(2): 53 (1986) Arachniodes reducta Y.T.Hsieh & Y.P.Wu; Bull. Bot. Res., Harbin 4(2): 105 (1984) Arachniodes repens Kurata; J. Geobot. 16: 98 (1968) Arachniodes rhomboidea (Wallich ex Mett.) Ching; Acta Phytotax. Sin. 9: 383 (1964) var. sinica Ching (1964) Arachniodes rigidissima (Hook.) Ching; Acta Bot. Sin. 10: 260 (1962) Arachniodes semifertilis Ching; Bull. Bot. Res., Harbin 6(3): 18 (1986) Arachniodes setifera Ching; Bull. Bot. Res., Harbin 6(3): 52 (1986) Arachniodes shuangbaiensis Ching; Bull. Bot. Res., Harbin 6(3): 21 (1986) Arachniodes sichuanensis Ching; Bull. Bot. Res., Harbin 6(3): 36 (1986) Arachniodes simplicior (Makino) Ohwi; J. Jap. Bot. 37: 76 (1962) Trivial names: Simpler East Indian Holly Fern, Variegated Shield fern var. major (Tagawa) Ohwi (1962) Arachniodes sinoaristata Ching; Bull. Bot. Res., Harbin 6(3): 20 (1986) Arachniodes sinomiqueliana (Ching) Ohwi; J. Jap. Bot. 37: 76 (1962) Arachniodes sinorhomboidea Ching; Bull. Bot. Res., Harbin 6(3): 55 (1986) Arachniodes sparsa Ching; Bull. Bot. Res., Harbin 6(3): 22 (1986) Arachniodes speciosa (D. Don) Ching; Acta Bot. Sin. 10: 259 (1962) Arachniodes spectabilis (Ching) Ching; Acta Bot. Sin, 10: 259 (1962) Arachniodes spinoserrulata Ching; Bull. Bot. Res., Harbin 6(3): 46 (1986) Arachniodes sporadosora (Kunze) Nakaike; Enum. Pteridophyt. Japon. Filicales, 192 (1975) Arachniodes standishii (T. Moore) Ohwi; J. Jap. Bot. 37: 76 (1962) Trivial names: Upside-down fern Arachniodes subamabilis Kurata; Sci. Rep. Yokosuka City Mus. 7: 37 (1962) Arachniodes subamoena Ching; Bull. Bot. Res., Harbin 6(3): 51 (1986) Arachniodes subaristata Ching & Y.T.Hsieh; Bull. Bot. Res., Harbin 4(2): 103 (1984) Arachniodes subreflexipinna (Ogata) Ching; Acta Bot. Sin. 10: 260 (1962) Arachniodes suijiangensis Ching & Y.T.Hsieh; Bull. Bot. Res., Harbin 6(4): 1 (1986) Arachniodes superba Fraser-Jenkins; New Species Syndrome in Indian Pteridology and the Ferns of Nepal 35 (1997) [nom. nov. Arachniodes tibetana Ching & S. K. Wu; Fl. Xizangica 1: 243 (1983) Arachniodes tiendongensis Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 9 (1983) Arachniodes tomitae Sa.Kurata; J. Geobot. 25(4): 98 (1978) Arachniodes tonkinensis (Ching) Ching; Acta Bot. Sin. 10: 260 (1962) Arachniodes triangularis Ching; Bull. Bot. Res., Harbin 6(3): 26 (1986) Arachniodes tripinnata (Goldm.) Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 5: 41 (1973) Arachniodes valida Y.T.Hsieh; Acta Bot. Yunnan. 5(1): 56 (1983) Arachniodes walkerae (Hook.) Ching; Acta Bot. Sin. 10: 260 (1962) Arachniodes webbiana (A. Braun) Schelpe; Bol. Soc. Brot. Ser. 2, 41: 203 (1967) ssp. webbiana (A. Braun) Schelpe; Bol. Soc. Brot. Ser. 2, 41: 203 (1967) ssp. foliosa (C. Chr.) Gibby, Rasbach, Reichst., Widen & Viane; Bot. Helv. 102(2): 243 (1992) Arachniodes wulingshanensis S.F.Wu; Keys Vasc. Pl. Wuling Mount. (ed. W.T.Wang & al.): 573 (1995) Arachniodes x clivorum Kurata; J. Geobot, 16: 97 (1968) Remarks: Arachniodes exilis x pseudoaristata Arachniodes x ikeminensis Seriz.; J. Jap. Bot. 61(2): 55 (1986) Remarks: Arachniodes miqueliana x quadripinnata ssp. fimbriata Arachniodes x kenzo-satakei (Kurata) Kurata; Sci. Rep. Yokosuka City Mus. 7: 39 (1962) Remarks: Byrsopteris pseudoaristata x standishii Arachniodes x kurosawae Shimura & Kurata in Kurata; J. Geobot. 16: 5 (1968) Remarks: Arachniodes amabilis x pseudoaristata Arachniodes x masakii Kurata; J. Geobot. 16: 5 (1968) Remarks: Arachniodes nipponica x pseudoaristata Arachniodes x minamitanii Sa.Kurata; J. Geobot. 25(4): 97 (1978) Remarks: Arachniodes hiugana x nipponica Arachniodes x mirabilis Sa.Kurata; J. Geobot. 25(4): 97 (1978) Remarks: Arachniodes assamica x rhomboidea Arachniodes x pseudohekiana Kurata; J. Geobot. 16: 5 (1968) Remarks: Arachniodes amabilis x hekiana Arachniodes x respiciens Sa.Kurata; J. Geobot. 25(4): 98 (1978) Remarks: Arachniodes exilis x simplicior Arachniodes x sasamotoi Kurata; J. Geobot. 16: 98 (1968) Remarks: Arachniodes exilis x simplicior var. major Arachniodes x takayamensis Seriz.; J. Jap. Bot. 61(2): 54 (1986) Remarks: Arachniodes borealis x miqueliana (Maxim.) Ohwi Arachniodes xinpingensis Ching; Bull. Bot. Res., Harbin 6(3): 23 (1986) Arachniodes yaomashanensis Ching; Bull. Bot. Res., Harbin 6(3): 32 (1986) Arachniodes yaoshanensis (Y. C. Wu) Seriz.; J. Jap. Bot. 48: 219 (1973) Arachniodes yasu-inouei Kurata; J. Geobot. 16: 6 (1968) Arachniodes yinjiangensis Ching; Bull. Bot. Res., Harbin 6(3): 44 (1986) Arachniodes yoshinagae (Makino) Ching; Acta Phytotax. Sin. 9: 383 (1964) Arachniodes yunnanensis Ching; Bull. Bot. Res., Harbin 6(3): 24 (1986) Arachniodes yunqiensis Y.T.Hsieh; Bull. Bot. Res., Harbin 6(4): 3 (1986) Arachniodes ziyunshanensis Y.T.Xie; Acta Phytotax. Sin. 22(2): 162 (1984)
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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