Family Athyriaceae, genus Athyrium;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Athyrium, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

            Lists Athyrium species A to J inclusive. Click here for species K to Z and hybrids.


005.00 Class Filicopsida
		Trivial names: Ferns
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
041.000 Order Athyriales
		Statistics: 6 families, 54 genera, 1782-2215 spp. [2362 species and 88 hybrids listed]
045.0000 Family Athyriaceae Alston; Taxon 5: 25 (1956) s. str.
		Statistics: 10 genera, 546-698 spp. [772 species and 56 hybrid listed]
		Revision date: 20.07.2001
		Remarks: according to PCR results, the family has probably to be divided. 
		There are two groups: Athyrium, Adenoderris and Cornopteris, which are monophyletic, 
		and the group Diplazium, Dictyodroma, Diplaziopsis, Hemidictyum and Deparia, which 
		is monophyletic with Thelypteridaceae and therefore has to be included there as 
		subfamily. At this time we have retained the traditional scheme to avoid confusion.
		= Dryopteridaceae subfam. Athyrioideae
		= Dryopteridaceae Tribus Physematieae J. Sm.; 1875

045.0002 Athyrium Roth; Tent. Fl. Germ. 3: 58 (1790) Statistics: 170-200 spp. worldwide, mostly northern hemisphere, many in China (127, but this number is most certainly be too high), several in trop. Africa and Madagascar, some in North America south to Mexico and Greater Antilles, only 3 in temperate South America [200 species and 42 hybrids listed]

Revision date: 20.07.2001 Trivial names: Lady Fern = Anisocampium Presl; Epim. Bot. 58 (1849-51) = Cystoathyrium Ching; Acta Phytotax. Sin. 11: 22 (1966) = Hypochlamys Fée; Gen. Fil. 200 (1850-52) = Kuniwatsukia Pichi-Serm.; Webbia 28: 455 (1975) = Pseudathyrium Newm.; Phytol. 4: 370 (1851) = Pseudocystopteris Ching; Acta Phytotax. Sin. 9: 76 (1964) = Solenopteris Zenker ms. ex Kze.; Linn. 24. 267 (1851) Athyrium adpressum Ching & W.M.Chu; Acta Bot. Boreal-Occid. Sin. 6(1): 13 (1986) Athyrium alpestre (Hoppe) Clairv.; Man. Herbor. Suisse 301 (1811) ssp. alpestre (Hoppe) Clairv.; Man. Herbor. Suisse 301 (1811) Trivial names: Alpen-Frauenfarn, Berg-Frauenfarn ssp. americanum (Butters) Lellinger; Amer. Fern J. 71(3): 91 (1981) Athyrium ammifolium (Mett.) C. Chr.; Ind. 139 (1905) Athyrium amoenum C. Chr.; Gardens Bull. S. S. 7. 267 t. 56 (1934) Athyrium anisopterum Christ; Bull. Herb. Boiss. 6: 962 (1898) Athyrium annae Komas; Kew Bull. 33(1): 99 (1978) Athyrium appendiculiferum Alderw.; Bull. Jard. bot. Buit. II nr. XVI. 3 (1914) Athyrium araiostegioides Ching; Acta Bot. Boreal-Occid. Sin. 6(2): 103 (1986) Athyrium archboldii Copel.; Univ. Calif. Publ. Bot. 18: 222 (1942) Athyrium arcuatum Liebm.; Vid. Selsk. Skr. V. 1. 278 (seors. 126) (1849) Athyrium arisanense (Hayata) Tagawa; Acta Phytotax. 2. 195 (1933) Athyrium atkinsonii Bedd.; Ferns br. Ind. Suppl. 11 t. 369 (1876) Athyrium atratum Bedd.; Handb. Ferns Brit. lndia Suppl. 33 (1892) Athyrium attenuatum (Clarke) Tagawa; Acta Phytotax. et Geobot. 16: 177 (1956) Athyrium atuntzeensis (Ching) comb. ined. Athyrium austro-orientale Ching; Fl. Xizangica 1: 141 (1983) Athyrium baishanzuense Ching & Y.T.Hsieh; Acta Bot. Boreal-Occid. Sin. 6(3): 157 (1986) Athyrium baoxingense Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 14 (1986) Athyrium barnebyanum Mickel & Beitel; Pterid. Fl. Oaxaca (Mem. New York Bot. Gard. 46): 77 (1988) Athyrium bomicola Ching; Fl. Xizangica 1: 136 (1983) Athyrium brevifrons Nakai ex Kitag.; Rep. First Sci. Exp. Manchoukuo 4(2): 75 (1935) Athyrium bourgaei Fourn.; Mex. pl. 1. 102 (1872) Athyrium brevipinnulum Copel.; Phil. Journ. Sci. Bot. 8. 291 (1908) Athyrium brevisorum (Wall. ex Hook.) Moore; Ind. Fil. 117 (1859) f. cristato-flabellatum (Makino) comb. ined. Athyrium brevistipes Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 12 (1986) Athyrium brooksii Copel.; Phil. Journ. Sci. Bot. 9. 229 (1914) Athyrium caespitosum Nakai; Bull. Nation. Sc. Mus. Tokyo 33: 3 (1953) [nom. nov. Athyrium caudatum Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 21 (1986) Athyrium chinense (Ching) comb. ined. Athyrium chingianum X.C.Zhang ex Z.R.Wang & X.C.Zhang; Bull. Bot. Res., Harbin 11(3): 12 (1991) Athyrium christensenii Tardieu-Blot; Aspl. du Tonkin 80 t. 12 (1932) Athyrium chungtienense Ching; Bull. Fan Mem. Inst. Biol. ser. 2. 1: 277 (1949) Athyrium clemensiae Copel.; Phil. Journ. Sci. 12 C. 58 (1917) Athyrium clivicola Tagawa; Acta Phytotax. Geobot. 3: 32 (1934) var. rotundum (Ching) Z. R. Wang (1999) Athyrium concinnum Nakai; Bot. Mag. Tokyo 45. 92 (1931) Athyrium costulalisorum Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 15 (1986) Athyrium crassipes Ching; Acta Bot. Boreal-Occid. Sin. 6(2): 103 (1986) Athyrium criticum Ching; Fl. Tsinlingensis 2: 216 (1974) Athyrium cryptogrammoides Hayata; Ic. Pl. Taiwan 6. 156 (1916) Athyrium cumingianum (Presl) Milde; Bot. Zeit. 353 (1870) Athyrium cuspidatum (Bedd.) M.Kato; Bot. Mag. (Tokyo) 90(1017): 27 (1977) Athyrium daxianglingense Ching & H.S.Kung; Acta Bot. Boreal-Occid. Sin. 6(1): 16 (1986) Athyrium delavayi Christ; Bull. Soc. Fr. 52, Mem. 1. 47 (1905) Athyrium delicatulum Ching & S. K. Wu; Fl. Xizangica 1: 139 (1983) Athyrium dentilobum Ching & S. K. Wu; Fl. Xizangica 1: 139 (1983) Athyrium devolii Ching; Sunyatsenia 3: 1. t.1 (1935) Athyrium dissitifolium (Bak.) C. Chr.; Contr. U. S. Nat. Herb. 26: 296, t. 18 (1931) var. funebre (Christ) Ching & Z. R. Wang (1999) Athyrium distans (Don) Moore; Ind. 125, 181 (1860) Athyrium dombeyi Desv.; Prod. 266 (1827) (dombei auctt.) Athyrium drepanopterum (Kunz.) A. Br. ex Milde; Fil. Fur. 49 (1867) Athyrium duthiei (Bedd.) Bedd.; Handb. Ferns. Brit. India Suppl. 34 (1892) Athyrium elongatum Ching; Acta Bot. Boreal-Occid. Sin. 6(2), 101 (1986) Athyrium emeicola Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 13 (1986) Athyrium epirachis (Christ) Ching; C. Chr., Ind. Suppl. III. 41 (1934) Athyrium erythropodum Hayata; Ic. Fl. Formosa 4. 233 t 163 (1914) Athyrium excelsius Nakai; Fedde Repert. 18. 243 (1914) Athyrium falcatum Bedd.; Ferns South. India t. 151 (1865) Athyrium fallaciosum Milde; Fil. Europ. et Atlant. 6: 54 (1867) Athyrium ferulaceum (T. Moore ex Hook.) H. Christ; Bull. Boiss. II. 4. 968 (1904) Athyrium filix-femina (L.) Roth; Tent. Fl. Germ. 3: 65 (1799) Trivial names: Wald-Frauenfarn, Lady Fern var. angustum (Willd.) Lawson; Edinburgh New Philos. J., n.s. 19: 114 (1864) var. asplenioides (Michx.) Farwell; Pap. Michigan Acad. Sci. 2: 13 (1923) Trivial names: Southern Lady Fern var. californicum Butters; Rhodora 19: 201 (1918) var. cyclosorum Rupr.; Distr. Crypt. Vasc. Ross. 41 (1845) Trivial names: Western Lady Fern var. retusa (Decne.) Bedd.; Handb. Ferns. Brit. India 170 (1883) Athyrium fimbriatum (Wall.) Moore; Ind. Fil. 185 (1860) Athyrium flabellulatum (Clarke) Tardieu-Blot; Aspl. du Tonkin 81 t. 12 (1932) Athyrium foliolosum (Wall.) Moore ex Sm.; Cat. Hort. no. 6: 22 (1859) Athyrium gedeanum (Rac.) Christ; Journ. de Bot. 19, 68 (1905) Athyrium grantii Copel.; Bishop Mus. Bull. 59. 12 (1932) Athyrium guangnanense Ching; Acta Bot. Boreal-Occid. Sin. 6(2): 103 (1986) [nom. nov. Athyrium gymnocarpum Copel.; Phil. Journ. Sci. 40. 301 t. 4 (1929) Athyrium hahasimense Nakai; Bot. Mag. Tokyo 48. 4 (1933) Athyrium hainanense Ching; Acta Bot. Boreal-Occid. Sin. 6(2): 101 (1986) Athyrium hakonense (Mak.) Ohwi; Bull. Nation. Sc. Mus. Tokyo. ser. 2. 3: 100 (1956) Athyrium heterocarpum Nakai; Bot. Mag. Tokyo 33. 131 (1921) Athyrium heterophyllum Nakai; Bot. Mag. Tokyo 45. 92 (1931) Athyrium himalaicum Ching ex Mehra & Bir; Res. Bull. (N. S.) Punjab. Univ. 15: 137. f. 26-29 (1964) Athyrium hirtirachis Ching & Hsu; Fl. Tsinlingensis 2: 215 (1974) Athyrium hochreutineri Christ; Ann. Conserv. Jard. bot. Geneve 15-16. 197 (1912) Athyrium hohenackerianum (Kunze) Moore; Ind. Fil. XLIX (1857) Athyrium infrapuberulum Ching; Acta Bot. Boreal-Occid. Sin. 6(2): 100 (1986) Athyrium interjectum Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 15 (1986) Athyrium intermixtum Ching & P. S. Chui; Acta Bot. Boreal-Occid. Sin. 6(1): 21 (1986) Athyrium irayense Copel.; Philip. Journ. Sc. 81: 39 (1952) Athyrium iseanum Rosenst.; Fedde Repert. 13. 124 (1913) var. chuanqianense Z. R. Wang (1999) Athyrium jinshajiangense Ching & Shing; Acta Bot. Boreal-Occid. Sin. 6(1): 14 (1986)


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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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