Family Athyriaceae, genus Athyrium;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Athyrium, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

            Lists Athyrium species K to Z inclusive, including hybrids. Click here for species A to J.


005.00 Class Filicopsida
		Trivial names: Ferns
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
041.000 Order Athyriales
		Statistics: 6 families, 54 genera, 1782-2215 spp. [2362 species and 88 hybrids listed]
045.0000 Family Athyriaceae Alston; Taxon 5: 25 (1956) s. str.
		Statistics: 10 genera, 546-698 spp. [772 species and 56 hybrid listed]
		Revision date: 20.07.2001
		Remarks: according to PCR results, the family has probably to be divided. 
		There are two groups: Athyrium, Adenoderris and Cornopteris, which are monophyletic, 
		and the group Diplazium, Dictyodroma, Diplaziopsis, Hemidictyum and Deparia, which 
		is monophyletic with Thelypteridaceae and therefore has to be included there as 
		subfamily. At this time we have retained the traditional scheme to avoid confusion.
		= Dryopteridaceae subfam. Athyrioideae
		= Dryopteridaceae Tribus Physematieae J. Sm.; 1875

Athyrium kawabatae Kurata; Sci. Rep. Yokosuka City Mus. 6: 22 (1961) Athyrium kenzo-satakei Kurata; Journ. Geobot. 7: 75 (1958) var. jieguishanense (Ching) Z. R. Wang (1999) Athyrium kidoanum Kurata; J. Geobot. 10: 10 (1961) Athyrium kirishimaense Tagawa; Acta Phytotax. 2. 22 (1933) Athyrium kumaonicum Punetha; Indian Fern J. 2(1-2): 29 (1985) Athyrium kuratae Serizawa; J. Jap. Bot. 45: 117 (1970) Athyrium kutaiense C. Chr.; Dansk Bot. Ark. 9(3): 55 (1937) Athyrium latifolium Presl; Tent. 98 t. 3 f. 4 (1836) Athyrium lewalleanum Pichi-Serm.; Webbia 27: 440 (1972) publ. (1973) Athyrium lineare Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 11 (1986) Athyrium longius Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 178 (1940) Athyrium ludingense Z.R. Wang & L.B. Zhang in L.B. Zhang, Z.R. Wang & X.S. Guo (1995) Athyrium mackinnoniorum (Hope) C. Chr.; Ind. Fil. 143 (1905) ('mackinnoni', emend. Fraser-Jenkins 1997) var. glabratum Y.T. Hsieh & Z.R. Wang in C.Y. Wu (1983) var. yigongense Ching & S.K. Wu in C.Y. Wu (1983) Athyrium macraei (Hook. & Grev.) Copel.; Gen. Fil. 149 (1947) Athyrium macrocarpum (BI.) Bedd.; Ferns South. India 51. t. 153 (1865) Athyrium majus Mak.; Bot. Mag, Tokyo 28. 178 (1914) Athyrium maoshanense Ching & P.S.Chiu; Acta Bot. Boreal-Occid. Sin. 6(3): 157 (1986) Athyrium masachikanum (Sa.Kurata) Seriz.; J. Jap. Bot. 56(7): 196 (1981) Athyrium masamunei Seriz.; Acta Phytotax. Geobot. 32(5-6): 176 (1981) Athyrium mearnsianum (Copel.) Alderw.; Mal. Ferns Suppl. 279 (1917) Athyrium medium (Carm.) Moore; Ind. 186 (1860) Athyrium medogense X.C.Zhang; Bull. Bot. Res., Harbin 11(3): 11 (1991) Athyrium melanolepis (Franch. & Sav.) Christ; Bull. Boiss 4. 668 (1896) Athyrium mengtzeense Hier.; Hedwigia 59. 319 (1918) Athyrium microphyllum (Sm.) Alston; Phil. Journ. Sci. 50. 178 (descr. orig.) (1933) Athyrium micropterum Fraser-Jenkins; New Species Syndrome in Indian Pteridology and the Ferns of Nepal 58 (1997) Athyrium minimum Ching; Acta Bot. Boreal-Occid. Sin. 6(3): 151 (1986) Athyrium minutum Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942) Athyrium mohillense (Fée) Tardieu; Bull. Mus. Hist. Nat. Paris Ber.2. 29: 290 (1957) Athyrium monomachii (Kom.) Kom.; Bull. Jard. bot. Kieff no. 13. 145 (1931) Athyrium morobense Copel.; Philip. Journ. Sc. 78: 464. t.40 (1951) Athyrium multifidum Rosenst.; Fedde Repert. 18. 125 (1913) var. gracillimum (Ching) Z. R. Wang (1999) Athyrium multijugum Nakai; Bot. Mag. Tokyo 45. 93 (1931) Athyrium multipinnum Y.T.Hsieh & Z.B. Wang; Acta Bot. Boreal-Occid. Sin. 7(1): 55 (1987) Athyrium myer-dreesii Copel.; Univ. Calif. Publ. Bot. 18: 222 (1942) Athyrium nakanoi Makino; Bot. Mag. Tokyo 23: 247 (1909) Athyrium neglectum Seriz.; J. Jap. Bot. 55(8): 250 (1980) ssp. neglectum Seriz.; J. Jap. Bot. 55(8): 250 (1980) ssp. australe Seriz.; J. Phytogeogr. Taxon 33: 76, f. 2 (1985) Athyrium nephrodioides (Bak.) Christ; Bull. Soc. Fr. 62 Mém. 1. 47 (1905) Athyrium newtonii (Bak) Diels; Nat. Pfl. 1(4). 224 (1899) Athyrium nikkoense Mak.; Bot. Mag. Tokyo 17. 43 (1903) Athyrium nitidulum (Kze.) Milde; Bot. Zeit. 370 (1870) Athyrium nudum Copel.; Fern Fl. Philip. 3: 391 (1960) Athyrium oblitescens Kurata; J. Geobot. 17: 3 (1969) Athyrium obtusilimbum Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 11 (1986) Athyrium omeiense Ching; Bull. Fan Mem. Inst. Biol. ser. 2. 1: 282 (1949) Athyrium oosorum (Bak.) Christ; Engl. Jahrb. 28. 349 (1896) Athyrium oppositipinnum Hayata; Journ. Coll. Sci. Univ. Tokyo 80. 441 (1911) Athyrium otophorum (Miq.) Koidzumi; Fl. Synth. Orient. As. 40 (1930) var. okanum Sa. Kurata; J. Geobot. 17: 3 (1969) Athyrium pachyphyllum Ching; Acta Bot. Boreal-Occid. Sin. 6(2): 102 (1986) Athyrium palmense (H. Christ) Lellinger; Proc. Biol. Sec. Wash. 89: 708 (1977) Athyrium palustre Seriz.; J. Jap. Bot. 45: 264 (1970) Athyrium parasnathense (C. B. Clarke) Ching ex Mehra & Bir; Res. Bull. Panjab Univ., Sci. 15: 141 (1964) Athyrium paucijugum (Alderw.) comb. ined. Athyrium pectinatum (Wall. ex Hope) Presl; Tent. Pterid. 98 (1836) Athyrium philippinense Christ; Copel., Phil. Journ. Sci. 1. Suppl. IV. 254 (1906) Athyrium pinetorum Tagawa; Acta Phytotax. 2. 16 (1933) Athyrium pitcairnense Copel.; Occ. Pap. Bishop Mus. 14: 60. t. 11 (1938) Athyrium praetermissum Sledge; Am & Mag. Nat. Hist. ser. 12. 9: 457. t. 16 (1956) Athyrium pubicostatum Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 7 (1984) Athyrium pulcherrimum Copel.; Phil. Journ. Sci. Bot. 8. 141 t. 3 (1913) Athyrium puncticaule (BI.) Moore; Ind. Fil. 186 (1860) Athyrium pygmaei-silvae Kurata; Sci. Rep. Yokosuka City Mus. 6: 20 (1961) Athyrium reflexipinnum Hayata; Ic. Fl. Formosa 4. 234 t 164 (1914) Athyrium regulare Koidzumi; Bot. Mag. Tokyo 88. 111 (1924) Athyrium repens (Ching) comb. ined. Athyrium rhachidosorum (Hand.-Mazz.) Ching; C. Chr., Ind. Suppl. III. 43 (1934) Athyrium rigidulum Nakai; Journ. Jap Bot. 16: 61 (1940) Athyrium roseum Christ; Bull. Boiss. 6. 961 (1898) var. fugongense Z.R.Wang; Vasc. Pl. Hengduan Mount. 1: 85 (1993), without latin descr. or type, et Z.R. Wang, Fl. Reipubl. Pop. Sin. 3(2): 233 (1999) Athyrium rubricaule (Edgew. ex C. B. Clarke) Bir; Nova Hedwigia 4: 169, t. 54 (1962) Athyrium rubripes (Kom.) Kom.; Bull. Jard. Bot. Kieff no. 13. 145 (1931) Athyrium ruilicolum W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 41 (1992) Athyrium rupestre Kodama; Matsum. Ic. Pl. Koisik. 4. 63 t. 244 (1919) Athyrium rupicola (Hope) C. Chr.; Ind. Fil. 145 (1905) Athyrium sarasinorum Christ; Verh. Nat. Ges. Basel 11. 434 (1896) Athyrium satowii H.Itô; Journ. Jap. Bot. 27: 337. f.1-2 (1952) Athyrium scandicinum (Willd.) Presl; Tent. 98 (1836) var. rhodesianum Schelpe (1967) f. bipinnata Rakotoudr.; Bull. Mus. Nat. Hist. Nat., B, Adansonia Ser. 4 10(4): 375 (1988 publ. 1989) Athyrium schimperi Moug. ex Fée; Mém. Gen. Fil. 187 (1850-52) ssp. schimperi Moug. ex Fée; Mém. Gen. Fil. 187 (1850-52) ssp. biserrulatum (Christ) Fraser-Jenkins; New Species Syndrome in Indian Pteridology and the Ferns of Nepal 60 (1997) Athyrium schizochlamys (Ching) K.Iwats., H.Ohba & S.B.Malla; Himalayan Pl. 1 (Univ. Mus. Univ. Tokyo Bull. 31): 313 (1988) Athyrium sericellum Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 17 (1986) Athyrium setuligerum Kurata; J. Geobot. 15: 6 (1966) Athyrium sheareri (Bak.) Ching; C. Chr., Ind. Suppl. III. 44 (1934) Athyrium silvicola Tagawa; Acta phytotax. 12. 17 (1933) Athyrium sinense Rupr.; Distr. Cr. vasc. Ross. 41 (1845) Athyrium sitchense (Rupr.) N.N. Tsvelev; Novosti Sist. Vyssh. Rast. 26: 8 (1989) Athyrium skinneri (Baker) C. Chr.; Index Filic. 146 (1905) Athyrium sohayakiense Nakaike; New Fl. Japan, Pterid.: 839 (1992) [nom. nov. Athyrium solenopteris (Kunze) Moore; Ind. Fil. 43. 1857 , 187 (1860) var. pusillum (Kunze) Moore; Ind. Fil. 187 (1860) Athyrium spinescens Kurata; J. Geobot. 15: 7 (1966) Athyrium spinulosum (Maxim.) Milde; Bot. Zeit. 376 (1866) Athyrium setiferum C. Chr.; Ind. 146 (1905) Athyrium nigripes (BI.) Moore; Ind. Fil. XLIX. 98 (1858) Athyrium strigillosum (T. Moore ex E.J. Lowe) Salomon (1883) Athyrium subimbricatum Nakai; Bot. Mag. Tokyo 35. 1: 11 (1921) Athyrium subrigescens Hayata; Ic. Fl. Formosa 4. 219 (syn.) (1914) ex H. Ito Athyrium subtriangulare (Hook.) Bedd.; Ferns Brit. India t. 242 (1868) Athyrium suprapuberulum Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 14 (1986) Athyrium suprapubescens Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 10 (1986) Athyrium supraspinescens C. Chr.; Contr. U. S. Nat. Herb. 297 t. 19 (1931) Athyrium tarulakaense Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 12 (1986) Athyrium tashiroi Tagawa; Acta Phytotax. 2. 21 (1932) Athyrium tenuipaleatum Copel.; Occ. Pap. Bishop Mus. 14: 60. t. 12 (1938) Athyrium triangulare Alderw.; Bull. Jard. bot. Buit. II nr. XX. 8 (1915) Athyrium uniforme Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 11 (1986) Athyrium venulosum Ching; Acta Bot. Boreal-Occid. Sin. 6(1): 13 (1986) Athyrium vidalii (Fr. & Sav.) Nakai; Bot. Mag. Tokyo 39. 110 (1925) var. amabile (Ching) Z. R. Wang (1999) Athyrium viridescentipes Kurata; J. Geobot. 13: 75 (1965) Athyrium viviparum Christ; Bull. Geogr. Bot. Mans 13 (1910) Athyrium wallichianum Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 497 (1938) Athyrium wangii Ching; Bull. Fan Mem. Inst. Biol. ser. 2. 1: 279 (1949) Athyrium wardii (Hook.) Mak.; Bot. Mag. Tokyo 18. 15 (1899) var. densipinnum Z.R. Wang & L.B. Zhang in L.B. Zhang, Z.R. Wang & X.S. Guo (1995) var. glabratum Y. T. Hsieh & Z. R. Wang (1999) Athyrium x akiense Kurata; J. Geobot. 15: 5 (1966) Athyrium x anceps Kurata; J. Geobot. 13: 76 (1965) Athyrium x arnagi-pedis Kurata; Sci. Rep. Yokosuka City Mus. 6: 23 (1961) Athyrium x austrojaponense Kurata; J. Geobot., 16: 99 (1968) Athyrium x awatae Seriz.; J. Phytogeogr. Taxon. 28(1): 34 (1980) Athyrium x bicolor Seriz.; J. Jap. Bot. 55(8): 252 (1980) Athyrium x calophyllum Sa.Kurata ex Seriz.; J. Jap. Bot. 55(8): 254 (1980) Athyrium x flavosorum Seriz.; J. Phytogeogr. Taxon. 33(2): 77 (1985) Athyrium x fuscopaleaceum Kurata; Sci. Rep. Yokosuka City Mus. 6: 24 (1961) Athyrium x heterosporum Hsieh & Z. R. Wang (1989) Athyrium x hisatsuanum Kurata; J. Geobot. 15: 6 (1966) Athyrium x hohuanshanense Yoshik.; J. Phytogeogr. Taxon. 33(2): 80 (1985) Athyrium x ikutae Kurata; J. Geobot. 16: 33 (1968) Athyrium x inabaense Seriz.; J. Phytogeogr. Taxon. 28(1): 34 (1980) Athyrium x inouei Kurata; J. Geobot. 17: 89 (1969) Athyrium x kawabataeoides Kurata; J. Geobot. 16: 33 (1968) Athyrium x masayukianum Kurata; J. Geobot. 13: 76 (1965) Athyrium x megayakusimense Kurata; J. Geobot. 14: 5 (1965) Athyrium x mentiens Kurata; J. Geobot. 16: 34 (1968) Athyrium x minakuchii Kurata; J. Geobot. 14: 85 (1966) Athyrium x neoelegans Kurata; J. Geobot. 14: 4 (1965) Athyrium x paludicola Sa.Kurata ex Seriz.; J. Jap. Bot. 55: 254 (1980) Athyrium x pseudocryptogrammoides Yoshik.; J. Phytogeogr. Taxon. 29(1): 27 (1981) Athyrium x pseudoiseanum Kurata; Journ. Geobot. 8: 84 (1960) Athyrium x pseudopinetorum Seriz.; J. Jap. Bot. 55(8): 253 (1980) Athyrium x pseudospinescens Seriz.; J. Phytogeogr. Taxon. 28(1): 35 (1980) Athyrium x pseudowardii Seriz.; J. Phytogeogr. Taxon. 28(1): 35 (1980) Athyrium x purpurascens (Tag.) Kurata; J. Geobot. 14: 4 (1965) Athyrium x purpureipes Kurata; Sci. Rep. Yokosuka City Mus. 6: 23 (1961) Athyrium x quaesitum Kurata; J. Geobot. 15: 5 (1966) Athyrium x reichsteinii Schneller & Rasbach. nssp. reichsteinii Schneller & Rasbach nssp. microderris Rasbach, Reichst. & J.Schneller; Bot. Helv. 101(2): 217 (1991) nssp. praetermissum Rasbach, Reichst. & J.Schneller; Bot. Helv. 101(2): 220 (1991) Athyrium x saitoanum (Sugim.) Seriz.; J. Phytogeogr. Taxon. 34(2): 68 (1986) Athyrium x satsumense Seriz.; J. Phytogeogr. Taxon. 28(1): 34 (1980) Athyrium x sefuricola Kurata; J. Geobot. 15: 6 (1966) Athyrium x shiibaense (Seriz.) Nakaike; New Fl. Japan, Pterid.: 839 (1992) Athyrium x shikoku-montanum Seriz.; J. Jap. Bot. 55(8): 255 (1980) Athyrium x sitchense (C.Presl) Tzvelev; Novosti Sist. Vyssh. Rast. 26: 8 (1989) Athyrium x subcrassipes Seriz.; J. Phytogeogr. Taxon. 33(2): 77 (1985) Athyrium x tokashikii Kurata; J. Geobot. 12: 15 (1963) Athyrium x tsurutanum Kurata; J. Geobot. 13: 77 (1965) Athyrium x watanabei Seriz.; J. Jap. Bot. 55(8): 252 (1980) Athyrium x yakuinsulare Kurata; J. Geobot. 13: 77 (1965) Athyrium xichouense Y.T.Hsieh & Z.R.Wang; Acta Phytotax. Sin. 27(2): 155 (1989) Athyrium yakumonticola Kurata; J. Geobot. 12: 15 (1963) Athyrium yakusimense Tagawa; Acta Phytotax. Geobot. 6: 94 (1937) Athyrium yokoscense (Franch. & Sav.) Christ; Bull. Boiss. 4. 668 (1896) Athyrium yuanyangense Y.T.Hsieh & W.M.Chu; Acta Bot. Boreal-Occid. Sin. 7(1): 54 (1987) Athyrium yui Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 6 (1940) Athyrium zayuense Z.R.Wang; Acta Phytotax. Sin. 32(1): 84 (1994) Athyrium zhenfengense Ching; Acta Bot. Boreal-Occid. Sin. 6(3): 151 (1986)
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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