Family Ophioglossaceae, genus Botrychium;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Botrychium, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.


005.00 Class Filicopsida
		Trivial names: Ferns
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.

 Family Ophioglossaceae


007.0003 Botrychium Swartz; J. Bot. (Schrader) 1800(2): 8 (1802) Statistics: ca. 60 spp. cosmopolitan, mostly North America and E-Asia [61 species and 5 hybrids listed] Revision date: 20.07.2001 Trivial names: Mondraute, Rautenfarn = Botrypus Michaux; Flora Boreali-Americana 2: 274 (1803) = Botrypus Richard; Cat. Hort. Med. Paris 120 (1801) = Holubiella Skoda; Preslia 68(4): 345 (1996) [1997] = Japanobotrychium Masamune; Jour. Soc. Trop. Agric. Formosa 3: 246 (1931) = Osmundopteris (Milde) Small; Ferns SE. States, 377, 482 (1938) = Osmundopteris Milde; Fil. Europae 209 (1867) = Phyllotrichium Prantl; Ber. deut. bot. Ges. 1. 349 (1883) = Sceptridium Lyon; Bot. Gazette 40: 457 (1905) Botrychium acuminatum Wagner; Contr. Univ. Michigan Herb. 17: 321 (1990) Trivial names: Pointed Moonwort Remarks: Subgen. Botrychium Botrychium arisanense (Masam.) Masam.; Short Fl. Form 1 (1936) Botrychium ascendens Wagner; Amer. Fern J. 76: 36, figs. 1, 2 (1986) Trivial names: Upswept Moonwort Remarks: Subgen. Botrychium Botrychium australe R. Br.; Prodr., 164 (1810) ssp. australe R. Br.; Prodr., 164 (1810) Trivial names: Parsley fern, Patotara ssp. negeri (H. Christ) R.T. Clausen; Memoirs of the Torrey Botanical Club 19(2): 40 (1938) Botrychium biforme Col.; Tr. New Zeal. Inst. 18. 223 (1886) Trivial names: Fine-leaved parsley fern Botrychium biternatum (Sav.) Underw.; Bot. Gaz. 22. 407 t. 21 (1896) Trivial names: Sparse-lobed Grape Fern Remarks: Subgen. Sceptridium Botrychium boreale (Fr.) Milde; Bot. Zeit. 15(51): 880 (1857) Remarks: Subgen. Botrychium Botrychium campestre Wagner & Farrar; Amer. Fern J. 76: 39, figs. 2, 4, 5 (1986) Trivial names: Prairie Moonwort Remarks: Subgen. Botrychium Botrychium chamaeconium Bitter & Hier.; Nat. Pfl. 1(4). 471 (1900) Botrychium crenulatum Wagner; Amer. Fern J. 71: 21 (1981) Trivial names: Dainty Moonwort Remarks: Subgen. Botrychium Botrychium daucifolium (Wall.) Hook. & Grev.; Icones Filicum 2(9): t. 161 (1830) Remarks: Subgen. Sceptridium var. parvum v. Ald. v. Ros.; Bull. Jard. Bot. Buit. 1: 3 (1911) var. intermedium (Sahashi) comb. ined. var. kanaii (Sahashi) comb. ined. Botrychium decompositum Mart. & Gal.; Mém. Ac. Brux. 15. 15 t. 1 (1842) Remarks: Subgen. Sceptridium Botrychium decurrens Ching; Chien & Chun, Fl. Reipubl. Pop. Sin. 2: 329 (1959) Botrychium dissectum Spreng.; Anleit. Kenntn. Gew. 3: 172 (1804) Trivial names: Cut-Leaf Grape Fern, lacy leaf grape fern. Remarks: Subgen. Sceptridium Botrychium dusenii (Christ) Alston; Lilloa 30: 107 (1960) Botrychium echo Wagner; Amer. Fern J. 73: 57 (1983) Trivial names: Echo Moonwort Remarks: Subgen. Botrychium Botrychium formosanum Tagawa; Acta Phytotax. Geobot. 9: 87 (1940) Botrychium gallicomontanum Farrar & Johnson-Groh; Amer. Fern J. 81: 1 (1991) Trivial names: Frenchman's Bluff Moonwort Remarks: Subgen. Botrychium Botrychium hesperium (Maxon & Clausen) Wagner & Lellinger; Amer. Fern J. 71: 92 (1981) Trivial names: Western Moonwort Remarks: Subgen. Botrychium Botrychium japonicum (Prantl) Underw.; Bull. Torr. Cl. 26. 538 (1898) var. silvicola (Sahashi) M.Kato; Fl. Japan 1: 26 (1995) Botrychium javanicum (Sahashi) comb. ined. Botrychium jenmanii Underw.; Fern Bull. 8: 59 (1900) Trivial names: Alabama Grape Fern Remarks: Subgen. Sceptridium Botrychium lanceolatum (S. G. Gmelin) Angström; Bot. Not. 14: 68 (1854) Trivial names: Triangle Moonwort ssp. lanceolatum (S. G. Gmelin) Angström; Bot. Not. 14: 68 (1854) Remarks: Subgen. Botrychium ssp. angustisegmentum (Paese & Moore) Clausen; Bull. Torrey Club 64: 280 (1937) Trivial names: Narrow Triangle Moonwort Botrychium lanuginosum (Wall.) Hook. & Grev.; Icones Filicum 1(4): t. 79 (1828) Remarks: Subgen. Japanobotrychium var. nepalense Nishida; Journ. Jap. Bot. 41: 319 (1966) [fide Fraser-Jenkins 1997 Botrychium lineare W.H.Wagner; Amer. Fern J. 84(1): 5 (1994) Botrychium longipedunculatum Ching; Chien & Chun, Fl. Reipubl. Pop. Sin. 2: 330 (1959) Botrychium lunaria (L.) Swartz; J. Bot. (Schrader) 1800(2): 110 (1802) Trivial names: Gemeine Mondraute, Moonwort Remarks: Subgen. Botrychium f. gracile (Schur) Asch. & Graebn. var. onondagense (Underwood) House; New York State Museum Bulletin 243-244: 47 (1923) f. subincisum (Roep.) Milde (1859) f. tripartitum (Moore) Weath. Botrychium lunarioides (Michx.) Swartz; Syn. Fil. 172 (1806) Trivial names: Winter Grape Fern Remarks: Subgen. Sceptridium Botrychium manshuricum Ching; Chien & Chun, Fl. Reipubl. Pop. Sin. 2: 329 (1959) Botrychium matricariifolium (Döll) A. Braun ex Koch; Syn. Deut. Schweiz. Fl. ed. 2, 7: 1009 (1847) Trivial names: Ästige Mondraute, Ästiger Rautenfarn, Daisy-leaf Grape Fern Remarks: Subgen. Botrychium f. gracile (House) Weath. Botrychium microphyllum (Sahashi) M.Kato; Fl. Japan 1: 27 (1995) Botrychium minganense Vict.; Proc. & Trans. Roy. Soc. Canada ser. 3, 21: 331 (1927) Trivial names: Mingan Moonwort Remarks: Subgen. Botrychium Botrychium modestum Ching; Chien & Chun, Fl. Reipubl. Pop. Sin. 2: 329 (1959) Botrychium montanum Wagner; Amer. Fern J. 71: 29 (1981) Trivial names: Western Goblin Remarks: Subgen. Botrychium Botrychium mormo Wagner; Amer. Fern J. 71: 26 (1981) Trivial names: Little Goblin Remarks: Subgen. Botrychium Botrychium multifidum (S. G. Gmelin) Rupr.; Beitr. Pflanzenk. Russ. Reiches 11: 40 (1859) Trivial names: Vielteilige Mondraute (Rautenfarn), Leathery Grape Fern Remarks: Subgen. Sceptridium var. robustum (Rupr.) C. Chr.; Kongliga Svenska Vetenskapsakademiens Handlingar 5(1): 48, t. 3 (1927) Botrychium nipponicum Mak.; Journ. Jap. Bot. 1: 5 (1916) Remarks: Subgen. Sceptridium var. minus (H.Hara) M.Kato; Fl. Japan 1: 26 (1995) Botrychium officinale Ching; Chien & Chun, Fl. Reipubl. Pop. Sin. 2: 330 (1959) Botrychium oneidense (Gilb.) House; Amer. Midl. Naturalist 7: 126 (1905) Trivial names: Blunt-lobed Grape Fern Remarks: Subgen. Sceptridium Botrychium pallidum Wagner; Amer. Fern J. 80: 74 (1990) Trivial names: Pale Moonwort Remarks: Subgen. Botrychium Botrychium paradoxum Wagner; Amer. Fern J. 71: 24 (1981) Trivial names: Paradox Moonwort Remarks: Subgen. Botrychium Botrychium parvum Ching; Chien & Chun, Fl. Reipubl. Pop. Sin. 2: 330 (1959) Botrychium pedunculosum Wagner; Amer. Fern J. 76: 43 (1986) Trivial names: Stalked Moonwort Remarks: Subgen. Botrychium Botrychium pinnatum H. St. John; Amer. Fern J. 19: 11 (1929) Trivial names: Northern Moonwort Remarks: Subgen. Botrychium Botrychium pseudopinnatum Wagner; Contr. Univ. Michigan Herb. 17: 322 (1990) Trivial names: False Northwestern Moonwort Remarks: Subgen. Botrychium Botrychium pumicola Coville; Underwood, Native Ferns ed. 6, 69 (1900) Trivial names: Pumice Moonwort Remarks: Subgen. Botrychium Botrychium rugulosum Wagner; Contr. Univ. Michigan Herb. 15: 315 (1982) Trivial names: Ternate Grape Fern, St. Lawrence Grape Fern Remarks: Subgen. Sceptridium Botrychium schaffneri Underw.; Bulletin of the Torrey Botanical Club 30: 51 (1903) Remarks: Subgen. Sceptridium Botrychium simplex E. Hitchc.; Amer. J. Sci. Arts 6: 103 (1823) Trivial names: Einfache Mondraute, Least Moonwort var. compositum (Lasch) Milde; Verhandlungen der Zoologisch-botanischen Gesellschaft in Wien 19: 140 (1869) f. laxifolium (Clausen) Fern. f. simplicissimum (Lasch) Milde Botrychium socorrense W.H.Wagner; G.A.Levin & R.C.Moran, Vasc. Fl. Isla Socorro, Mexico (Mem. San Diego Soc. Nat. Hist. 16): 20 (1989) Botrychium spathulatum Wagner; Amer. Fern J. 80: 77 (1990) Trivial names: Spatulate Moonwort Remarks: Subgen. Botrychium Botrychium strictum Underw.; Bulletin of the Torrey Botanical Club 20: 52 (1902) Remarks: Subgen. Osmundopteris Botrychium subbifoliatum Brackenr.; Expl. Exp. 16. 317 t. 44 f. 2 (1854) Botrychium sutchuanense Ching; Chien & Chun, Fl. Reipubl. Pop. Sin. 2: 330 (1959) Botrychium ternatum (Thunb.) Sw.; Schrad. Journ. 1800(2). 111 (1801) var. pseudoternatum (Sahashi) M.Kato; Fl. Japan 1: 27 (1995) Botrychium tibeticum (Ching) comb. ined. Botrychium triangularifolium (Sahashi) M.Kato; Fl. Japan 1: 27 (1995) Botrychium underwoodianum Maxon; Bull. Torr. Cl. 32. 220 t. 6 (1905) Botrychium virginianum (L.) Sw.; Schrad. Journ. 1800(2). 111 (1801) Trivial names: Virginische Mondraute (Rautenfarn), Rattlesnake Fern Remarks: Subgen. Osmundopteris Botrychium x argutum (Sahashi) comb. ined. Remarks: Sceptridium japonicum var. japonicum x triangularifolium Botrychium x elegans (Sahashi) comb. ined. Remarks: Sceptridium japonicum var. japonicum x nipponicum var. nipponicum Botrychium x longistipitatum (Sahashi) comb. ined. Remarks: Sceptridium atrovirens x ternatum Botrychium x pulchrum (Sahashi) comb. ined. Remarks: Sceptridium atrovirens x nipponicum var. nipponicum Botrychium x watertonense W.H.Wagner; Canad. J. Bot. 62(4): 630 (1984) Remarks: Subgen. Botrychium, Botrychium hesperium x Botrychium paradoxum Botrychium yunnanense Ching; Chien & Chun, Fl. Reipubl. Pop. Sin. 2: 329 (1959)
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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