Family Thelypteridaceae, genus Christella;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Christella, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

005.00 Class Filicopsida
		Trivial names: Ferns
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
041.000 Order Athyriales
		Statistics: 6 families, 54 genera, 1782-2215 spp. [2362 species and 88 hybrids listed]
046.0000 Family Thelypteridaceae Pichi-Sermolli; Webbia 24: 711 (1970) 
		Statistics: 1-30 genera (depending on genus concept), 975-1217 spp. 
		[1271 species and 14 hybrids listed]
		Remarks: This Checklist follows closely the Index of Thelypteridaceae (Kew), 
		even if the necessary combination for the genus are not available.
		Revision date: 20.07.2001 
		= Aspidiaceae subfam. Thelypteridoideae (C. Chr.) C. Reed; Phytologia 17: 254 (1968)
		= Polypodiaceae tribe Thelypterideae C. Chr.

046.0019 Christella Lév.; Fl. de Kouy-tchéou (1915) 472 [emend. Holtt. 1971 Statistics: 30-140 spp. India, SE-Asia, China (127 Cyclosorus described), 2 Hawaii, several Africa/Madagascar, 2 neotropical. [176 species and 8 hybrids listed]. The concept of the Chinese authors (in Fl. Reipubl. Pop. Sin.) is largely incompatible with other systems. Their "Cyclosorus" includes species of no less than 6 other subgenera. The Cyclosorus-microspecies of China are listed here in Christella, but they may belong to other subgenera, like Amphineuron or Sphaerostephanos. Revision date: 20.07.2001 = Nephrodium Schott; Gen. Fil. (1834) t. 10 = Christella sect. Pelazoneuron Holtt.; J. S. Afr. Bot. 40: 144 (1974) Christella abbreviata (Ching & K.H. Shing) comb. ined. Christella acuminata (Houtt.) Lév.; Fl. Kouy-tchéou (1915) 476 Christella acutiloba (Ching ex Shing) comb. ined. Christella acutissima (Ching ex K.H. Shing & J.F. Cheng) comb. ined. Christella adenopelta Holtt.; Kew Bull. 31 (1976) 322 Christella afzelii (C. Chr.) Holtt.; J. S. Afr. Bot. 40: 146 (1974) Christella albicaulis (Fée) comb. ined. Christella albidipilosa (Bonap.) comb. ined. Christella angusta (Ching) comb. ined. Christella angustipinna (Ching) comb. ined. Christella angustipinnata (C. Chr. & Tardieu ex Tardieu) comb. ined. Christella appendiculata (C.Presl) Holtt.; Kew Bull. 31(2): 311 (1976) Christella arida (D. Don) Holtt.; Nayar & Kaur, Comp. to Bedd. (1974) 206 Christella attenuata (Ching ex Shing) comb. ined. Christella augescens (Link) Pichi-Serm.; Webbia 31(1): 352 (1977) Trivial names: Abrupt-tip Maiden Fern Christella aureoglandulifera (Ching ex Shing) comb. ined. Christella aureoglandulosa (Ching & Shing) comb. ined. Christella baiseensis (Ching ex Shing) comb. ined. Christella balansae (Ching) Holtt.; Kew Bull. 31(2): 321 (1976) Christella bangii (C. Chr.) comb. ined. Christella berroi (C. Chr.) comb. ined. Christella blepharis (A.R. Sm.) comb. ined. Christella boninensis (Kodama ex Koidz.) Holtt.; Kew Bull. 31(2): 316 (1976) Christella boydiae (A.A.Eaton) Holtt.; Kew Bull. 31(2): 325 (1976) Christella brevipes (Ching) comb. ined. Christella burmanica (Ching) Holtt.; Kew Bull. 31(2): 317 (1976) Christella burundensis Pichi-Serm.; Bull. Jard. Bot. Belg. 53(1-2): 277 (1983) Christella buwaldae (Holtt.) Holtt.; Fl. Males. Ser. 2 l(5): 554 (1982) Christella caii (Ching ex Shing) comb. ined. Christella callensii (Alston) Holtt.; J. S. Afr. Bot. 40: 146 (1974) Christella calvescens (Ching) Holtt.; Kew Bull. 31(2): 328 (1976) Christella cangnanensis (Shing & C. F. Zhang) comb. ined. Christella carolinensis (Hosok.) Holtt.; Kew Bull. 31(2): 307 (1976) Christella chaseana (Schelpe) Holtt.; J. S. Afr. Bot. 40: 148 (1974) Christella chengii (Ching ex K.H. Shing & J.F. Cheng) comb. ined. Christella ciliensis (Shing) comb. ined. Christella clarkei (Beddome) Holtt., B. Nayar & Kaur; Companion Beddome's Handb. Ferns Brit. India, 203 (1974) Christella clavata (Shing) comb. ined. Christella conspersa (Schrad.) A.Löve & D.Löve; Taxon 26(2-3): 326 (1977) Christella contracta (Ching) comb. ined. Christella cretacea (A. R. Sm.) A.Löve & D.Löve; Taxon 26(2-3): 326 (1977) Christella crinipes (Hook.) Holtt., B. Nayar & Kaur; Companion Beddome's Handb. Ferns Brit. India, 208 (1974) Christella cuneata (Ching ex Shing) comb. ined. Christella cyatheoides (Kaulf.) Holtt.; Kew Bull. 31(2): 323 (1976) Christella damingshanensis (Ching ex Shing) comb. ined. Christella decipiens (Ching) comb. ined. Christella dehuaensis (Ching & K.H. Shing) comb. ined. Christella densissima (Ching ex Shing) comb. ined. Christella dentata (Forsskal) Brownsey & Jermy; Brit. Fern Gaz. 10: 338 (1973) Trivial names: Downy Maiden Fern, Downy woodfern var. buchananii Schelpe (1965) var. caespitosa Holtt.; Kew Bull. 41(3): 518 (1986) Christella depilata (A. R. Sm.) comb. ined. Christella dissita (Ching ex Shing) comb. ined. Christella distans (Hook.) Holtt.; J. S. Afr. Bot. 40: 140 (1974) Christella dulongjiangensis (W.M.Chu) comb. ined. Christella elata (Ching ex Shing) comb. ined. Christella ensifera (Tag.) Holtt. ex Kuo, H. L. Liu, T. C. Huang, T. Koyama & DeVol; Fl. Formosa 1: 406 (1975) Christella euphlebia (Ching) Holtt.; Kew Bull. 31(2): 328 (1976) Christella evoluta (Bedd.) Holtt.; Nayar & Kaur, Comp. to Bedd. Handb. 208 (1974) Christella excelsior (Ching & K. H. Shing) comb. ined. Christella fengii (Ching ex Shing) comb. ined. Christella fraxinifolia (Ching & K. H. Shing) comb. ined. Christella friesii (Brause) Holtt.; J. S. Afr. Bot. 40: 145 (1974) Christella fukienensis (Ching) Holtt.; Kew Bull. 31(2): 318 (1976) Christella gaoxiongensis (Ching ex Shing) comb. ined. Christella glabrescens (Ching ex Shing) comb. ined. Christella goedenii (Rosenstock) comb. ined. Christella grandissima (Ching & K. H. Shing) comb. ined. Christella gretheri (W.H.Wagner) Holtt.; Allertonia l(3): 219 (1977) Christella grossodentata (Ching ex Shing) comb. ined. Christella guamensis Holtt.; Allertonia l(3): 222 (1977) Christella gueintziana (Mett.) Holtt.; J. S. Afr. Bot. 40: 147 (1974) Christella guineensis (Christ) Holtt.; J. S. Afr. Bot. 40: 145 (1974) Christella gustavii (Bedd.) Holtt.; Nayar & Kaur, Comp. to Bedd. Handb. 208 (1974) Christella harveyi (Mett.) Holtt.; Kew Bull. 31 (1976) 306 var. connivens Holtt.; Kew Bull. 31(2): 307 (1976) Christella hirtipes (Shing & C. F. Zhang) comb. ined. Christella hispidula (Decne.) Holtt.; Kew Bull. 31 (1976) 312 Trivial names: Hairy Maiden Fern var. inconstans (C. Chr.) comb. ined. var. versicolor (R.P. St. John) comb. ined. Christella hokouensis (Ching) Holtt.; Kew Bull. 31 (2): 327 (1977) Christella houi (Ching) comb. ined. Christella jaculosa (Christ) Holtt.; Webbia 30(1): 193 (1976) Christella jinghongensis (Ching ex Shing) comb. ined. Christella jiulongshanensis (P.S.Chiu & Yao ex Ching) comb. ined. Christella kuizhouensis (Shing) comb. ined. Christella kweichowensis (Ching ex Shing) comb. ined. Christella lanosa (C. Chr.) A.Löve & D.Löve; Taxon 26(2-3): 326 (1977) Christella lebeufii (Baker) Holtt.; B. Nayar & Kaur, Companion Beddome's Handb. Ferns Brit. India, 206 (1974) Christella leipoensis (Ching & H. S. Kung) comb. ined. Christella longqishanensis (Shing) comb. ined. Christella macrophylla (Ching & Z.Y.Liu) comb. ined. Christella meeboldii (Rosenst.) Holtt.; Nayar & Kaur, Comp. to Bedd. Handb. 208 (1974) Christella medogensis (Ching & S. K. Wu) comb. ined. Christella megongensis (Ching ex Shing) comb. ined. Christella mianningensis (Shing) comb. ined. Christella microbasis (Baker) Holtt.; J. S. Afr. Bot. 40: 146 (1974) Christella minima Holtt.; Kew Bull. 31(2): 304 (1976) Christella modesta Holtt.; J. S. Afr. Bot. 40: 144 (1974) Christella mollissima (Ching ex Shing) comb. ined. Christella multiauriculata Punetha; J. Ind. Bot. Soc. 69(1-2): 177 (1990) Christella multifrons (C. Chr.) Holtt.; J. S. Afr. Bot. 40: 145 (1974) Christella multisora (Ching ex Shing) comb. ined. Christella namburensis (Bedd.) Holtt.; Nayar & Kaur, Comp. to Bedd. Handb. 206 (1974) Christella nana Holtt.; Kew Bull. 31(2): 304 (1976) Christella nanchuanensis (Ching & Liu) comb. ined. Christella nanlingensis (Ching ex K. H. Shing & J. F. Cheng) comb. ined. Christella nanpingensis (Ching) comb. ined. Christella nanxiensis (Ching ex Shing) comb. ined. Christella nigrescens (Ching ex Shing) comb. ined. Christella normalis (C. Chr.) Holtt.; Webbia 30(1): 193 (1976) Trivial names: Widespread Maiden Fern Christella oblanceolata (Ching) comb. ined. Christella omeigensis (Ching) Holtt.; Kew Bull. 31(2): 318 (1976) Christella opposita (Ching ex Shing) comb. ined. Christella orientalis (Ching ex Shing) comb. ined. Christella orizabae (Fée) comb. ined. Christella ovata (R.P.St.John) A.Löve & D.Löve; Taxon 26(2-3): 326 (1977) Trivial names: Ovate Maiden Fern var. lindheimeri (C. Chr.) A.R. Sm., Amer. Fern J. 61: 30 (1971) Trivial names: Lindheimers Maiden Fern Christella pacifica Holtt.; Kew Bull. 31(2): 320 (1976) Christella papilio (C. Hope) Holtt.; B. Nayar & Kaur, Companion Beddome's Handb. Ferns Brit. India, 208 (1974) Christella papilionacea (Shing & C. F. Zhang) comb. ined. Christella papyracea (Bedd.) Holtt.; Nayar & Kaur, Comp. to Bedd. Handb. 206 (1974) Christella paracuminata (Ching ex K. H. Shing & J. F. Cheng) comb. ined. Christella paradentata (Ching ex Shing) comb. ined. Christella paralatipinna (Ching ex Shing) comb. ined. Christella pararida (Ching ex Shing) comb. ined. Christella parasitica (L.) Lév.; Fl. Kouytchéou (1915) 475 var. pitcairnensis (E. Br.) comb. ined. var. rapensis (E. Br.) comb. ined. var. rurutuensis (E. Br.) comb. ined. var. mangarevensis (E. Br.) comb. ined. Christella parvifolia (Ching) comb. ined. Christella parviloba (Ching & Shing ex Shing) comb. ined. Christella patens (Sw.) Holtt.; Webbia 30(1): 193 (1976) var. scabriuscula (Smith) comb. ined. var. dissimilis (Schrad.) comb. ined. Christella paucipinna (Ching & C. F. Zhang ex Shing) comb. ined. Christella pauciserrata (Ching & C.F.Zhang) comb. ined. Christella peekelii (Alderw.) Holtt.; Kew Bull. 31 (1976) 306 Christella perpubescens (Alst.) Holtt.; Kew Bull. 31 (1976) 304 Christella pingshanensis (Ching & H.S. Kung ex Shing) comb. ined. Christella procurrens (Rosenst) comb. ined. Christella prolixa (Willd.) Holtt.; Amer. Fern J. 63: 83 (1973) Christella proxima (Ching) comb. ined. Christella pseudoarida (Ching ex Shing) comb. ined. Christella pseudocuneata (Ching ex Shing) comb. ined. Christella puberula A.Löve & D.Löve; Taxon 26(2-3): 326 (1977) Trivial names: Sonoran Maiden Fern var. sonorensis (A.R. Smith) comb. ined. Christella pumila (Ching ex Shing) comb. ined. Christella pustulifera (Ching ex Shing) comb. ined. Christella pygmaea (Ching & C.F.Zhang) comb. ined. Christella repens (Sledge) comb. ined. Christella rupicola (Ching) comb. ined., non (Hosok.) Holtt. Christella rupicola (Hosok.) Holtt.; Allertonia l(3): 217 (1977) Christella sanduensis (Shing & P. S. Wang) comb. ined. Christella scaberula (Ching) Holtt.; Kew Bull. 31(2): 329 (1976) Christella semisagittata (Roxb. ex Griff.) Holtt.; Kew Bull. 31 (2): 334 (1977) Christella serra (Sw.) Holtt.; Webbia 30(1): 194 (1976) Christella serrifera (Ching ex Shing) comb. ined. Christella shapingbaensis (Ching ex Shing) comb. ined. Christella shimenensis (Shing & C. M. Zhang) comb. ined. Christella simillima (Ching ex Shing) comb. ined. Christella sinodentata (Ching & Z.Y.Liu) comb. ined. Christella sparsisora (Ching ex Shing) comb. ined. Christella stenopes (Ching & Shing ex Shing) comb. ined. Christella subacuminata (Ching ex K. H. Shing & J. F. Cheng) comb. ined. Christella subacuta (Ching) comb. ined. Christella subcoriacea (Ching ex Shing) comb. ined. Christella subdentata Holtt.; Kew Bull. 31 (1976) 335 Christella subelata (Bak.) Holtt.; Kew Bull. 31 (2): 331 (1977) Christella subjuncta (Baker) Holltum; Allertonia l(3): 220 (1977) Christella subnamburensis (Ching ex Shing) comb. ined. Christella subpubescens (Bl.) Holtt.; Webbia 30 (1976) 193 Christella taprobanica (Panigrahi) Holtt.; Kew Bull. 31(2): 335 (1976) Christella tarningensis (Ching) comb. ined. Christella timorensis Holtt.; Kew Bull. 31 (1976) 316 Christella transitoria (Ching ex Shing) comb. ined. Christella tuerckheimii (Donn. Sm.) comb. ined. Christella wailele T.Flynn & W.H. Wagner Christella wangii (Ching) comb. ined. Christella wangmoensis (Shing ex P. S. Wang) comb. ined. Christella wenzhouensis (Shing & C. F. Zhang) comb. ined. Christella wulingshanensis (C.M.Zhang) comb. ined. Christella x altissima Holtt.; J. S. Afr. Bot. 40: 141 (1974) Christella x incesta (W.H.Wagner) Nakaike & Kawabata; J. Nat. Hist. Mus. Inst. Christella x intermedia (W.C.Shieh & J.L.Tsai) comb. ined. Christella x kumaunica Holtt.; Kew Bull. 31 (2): 318 (1977) Christella x linii (Fraser-Jenkins) comb. ined. Christella x nareshii (Fraser-Jenkins) comb. ined. Christella x palmeri W.H.Wagner Christella x parahispidula (Fraser-Jenkins) comb. ined. Christella xunwuensis (Ching ex K. H. Shing & J. F. Cheng) comb. ined. Christella yandangensis (Ching & K. H. Shing) comb. ined. Christella yuanjiangensis (Ching ex Shing) comb. ined. Christella yunnanensis (Ching ex Shing) comb. ined. Christella zeylanica (Fée) Holtt.; Nayar & Kaur, Comp. to Bedd. Handb. 201 (1974) Christella zhangii (Shing) comb. ined.
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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