Family Athyriaceae, genus Diplazium;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Diplazium, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

             Diplazium species A to G.
             For Diplazium species G to O, Here   |  For Diplazium species P to Z and hybrids, Here


 005.00 Class Filicopsida
		Trivial names: Ferns
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
041.000 Order Athyriales
		Statistics: 6 families, 54 genera, 1782-2215 spp. [2362 species and 88 hybrids listed]
045.0000 Family Athyriaceae Alston; Taxon 5: 25 (1956) s. str.
		Statistics: 10 genera, 546-698 spp. [772 species and 56 hybrid listed]
		Revision date: 20.07.2001
		Remarks: according to PCR results, the family has probably to be divided. 
		There are two groups: Athyrium, Adenoderris and Cornopteris, which are monophyletic, 
		and the group Diplazium, Dictyodroma, Diplaziopsis, Hemidictyum and Deparia, which 
		is monophyletic with Thelypteridaceae and therefore has to be included there as 
		subfamily. At this time we have retained the traditional scheme to avoid confusion.
		= Dryopteridaceae subfam. Athyrioideae
		= Dryopteridaceae Tribus Physematieae J. Sm.; 1875

045.0006 Diplazium Swartz; J. Bot. (Schrader) 1800(2): 61 (1801) Statistics: 300-400 spp. cosmopolitan, 100 spp. in tropical America [474 species and 8 hybrids listed]. Borders between this genus, Athyrium and Allantodia very unclear. Many probably invalid microspecies in China Revision date: 20.07.2001 Trivial names: Twin-Sorus Fern = Anisogonium Presl; Tent. Pterid. 115 (1836) = Asplenium grex Triblemma J.Sm. = Brachysorus Presl; Epim. Bot. 70 (1851) = Callipteridastrum Fée; Gen. Fil. 213 (1850-52) = Callipteris Bory; Voyage 1. 282 (1804) = Didymochlamys Moore; Index LV (1857) = Digrammaria Presl; Tent. Pterid. 116 (1836) = Homalosorus Small ex Pichi-Serm.; Webbia 31(1): 246 (1977) = Lotzea Klotzsch & Karsten; Linnaea 20. 358 (1847) = Microstegia Presl; Epim. Bot. 90 (1851) = Monomelangium Hayata; Bot. Mag. Tokyo 42. 343 (1928) = Ochlogramma Presl; Epim. Bot. 93 (1849-51) = Oxygonium Presl; Tent. Pterid. 117 (1836) = Pteriglyphis Fée; Xe. Congr. scient. de France 1: 178 (1843) = Pteriglyphis Fée; Mem. Fam. Foug. 5 (Gen. Fil.) 219 (1852) [nom. superfl. = Triblemma (J.Sm.) Ching; Acta Phytotax. Sin. 16(4): 23 (1978) = Dictyodroma Ching; Acta Phytotax. Sin. 9: 57 (1964) = Diplaziopsis C. Chr.; Ind. Fil. 227 (1905) = Allantodia Wallich; Pl. asiat. rar. 1. 44 t. 22 (1830) [non R. Brown (1810) = Allantodia R. Brown; Prod. Fl. N. Holl. 149 (1810) Diplazium aberrans Maxon & Morton; Amer. Fern Journ. 36: 93 (1946) Diplazium acanthopus C. Chr.; Svensk Bot. Tids. 16. 94 (1922) Diplazium andapense (Tardieu) comb. ined. Diplazium acrocarpum Rosenst.; Fedde Repert. 10. 328 (1912) Diplazium acrotis Christ; Bull. Boiss. II. 6. 1000 (1906) Diplazium aculeatum Alderw.; Bull. Buit. II no. 28. 20 (1918) Diplazium aemulum Underw. & Maxon; Bull. Torr. Cl. 29. 583 (1902) Diplazium aequibasale (Bak.) C. Chr.; Ind. 227 (1905) Diplazium agyokuense Tagawa; Acta Phytotax. Geobot. 7: 80 (1938) Diplazium albido-squamatum Alderw.; Bull. Buit. II no. 23. 9 (1916) Diplazium alienum (Mett.) Hier.; Hedwigia 69. 336 (1917) Diplazium alsophilum Maxon; Amer. Fern Journ. 14. 75 (1924) Diplazium altissimum (Jenm.) C. Chr.; Ind. 227 (1905) Diplazium altum Copel.; C. Chr., Ind. Suppl. III. 72 (1934) Diplazium amamianum Tagawa; Acta Phytotax. Geobot. 3: 35 (1934) Diplazium ambiguum Raddi; Opusc. sci. Bol. 3. 292 (1819) var. dissectum Stolze; Fl. Ecuador 49(14:5B): 20 (1994) Diplazium amplifrons Alderw.; Bull. Jard. bot. Buit. II nr. XI. 8 (1913) Diplazium amplissimum (Bak.) Diels; Nat. Pfl. 1(4). 227 (1899) Diplazium angulosum C. Chr.; Ind. 227 (1905) Diplazium angustatum Amann; Nat. Tijdschr. Ned. Ind. 23. 406 (1861) Diplazium angustipinna (Holtt.) Holtt.; Fl. Malaysia ed. 2, 2: 637 (1968) Diplazium angustisquamatum (Holtt.) Parris; Fern Gaz. 12(2): 118-119 (1980) Diplazium apatelium Alderw.; Bull. Buit. III. 6. 195 (1922) Diplazium apollinaris L'Herm. ex Fée; 11 mem. 40 t. 10 f. 1 (1866) Diplazium arborescens Bory ex Sw.; Syn. 9 (1806) Diplazium armatum (Copel.) Holtt.; Blumea 13: 135 (1965) Diplazium asperulum Alderw.; Bull. Jard. bot. Buit. II nr. XVI. 8 (1914) Diplazium asperum Presl; Epim. 195 (1828) Diplazium assimile (Endl.) Bedd.; Ferns br. Ind. t. 294 (1808) Diplazium asterothrix C. Chr.; Brittonia 2: 292 (1937) Diplazium atirrense (Donn. Sm.) Lellinger; Amer. Fern J. 67: 58 (1977) var. lobulatum C.D. Adams. Novon 2: 292 (1992) Diplazium atratum Christ; Phil. Journ. Sci. Bot. 2. 163 (1907) Diplazium atropurpureum Rosenst.; Fedde Repert. 12. 528 (1913) [non Athyrium atropurpureum Copel. Diplazium atrosquamosum (Copel.) C. Chr.; Ind. Suppl. III. 72 (1931) Diplazium australe (R. Br.) Wakef.; Vict. Nat. 58: 142. f.3 (1942) Trivial names: Austral Lady fern Diplazium avitaguense Hieron.; Hedwigia 47. 216 t. 2 f. 7 (1908) Diplazium baishanzuensis (Ching & Chiu) comb. ined. Diplazium balliviani Rosenst.; Fedde Repert. 6. 311 (1909) Diplazium banahaoense (Copel.) C. Chr.; Ind. Suppl. III. 72 (1934) Diplazium bantamense Bl.; Enum. 191 (1828) Diplazium barbatum C. Chr.; Gardens Bull. 7. 272 t. 59 (1934) Diplazium barisanicum (Bak.) C. Chr.; Ind. 228 (1905) Diplazium basahense Ching; Ching & Wang, Acta Phytotax. Sin. 9: 351 (1964) Diplazium beamanii M.G.Price; Contr. Univ. Michigan Herb. 16: 193 (1987) Diplazium beddomei C. Chr.; Ind. 228 (1905) Diplazium bellum (C. B. Clarke) Bir in Mehra & Bir; Res. Bull. Panjab Univ., Sci. 15: 148 (1964) Diplazium betimusense Alderw.; Bull. Buit. III. 2. 142 (1920) Diplazium bicolor Stolze; Fieldiana, Bot. 27: 74 (1991) Diplazium biolleyi H. Christ; Bull. Herb. Boissier sér. 2, 7: 269 (1907) Diplazium birgeri C. Chr.; Dansk Bot. Ark. 9(3): 58 (1937) Diplazium bittyuense Tagawa; Acta Phytotax. 2. 196 (1933) Diplazium blanchetii Mett.; Linn. 36. 104 (1869) (syn.) Diplazium bogotense (Karst.); Hedwigia 47. 218 (1908) Diplazium bolsteri Copel.; Phil. Journ. Sci. 1. Suppl. IV. 254 t. 2 (1906) Diplazium bombonasae Rosenst.; Fedde Repert. 7. 294 (1909) var. petiolulatum Stolze; Fl. Ecuador 49(14:5B): 53 (1994) Diplazium bostockii D. L. Jones; Fl. Australia 48: 712 (1998) Diplazium brachylobum (Sledge) Manickam & V.Irudayaraj; Indian Fern J. 7(1-2): 119 (1990) Diplazium brachysoroides Copel.; Phil. Journ. Sci. Bot. 2. 127 t. 1 f. A (1907) Diplazium brasilianum Rosenst.; Hedwigia 46. 107 (1906) Diplazium brausei Rosenst.; Repert. Spec. Nov. Regni Veg. 9: 68 (1910) Diplazium brevipes (Bak.) C. Chr.; Ind. 229 (1905) Diplazium brunonianum (Wall.) comb. ined. Diplazium buchtienii Rosenst.; Fedde Repert. 6. 312 (1909) Diplazium burmanicum Ching apud W. M. Chu & Z. R. He; Acta Phytotax. Sin. 36(4): 381, t. 3 (1998) Diplazium calliphyllum (Copel.) M.G.Price; Contr. Univ. Michigan Herb. 17: 270 (1990) Diplazium calogramma Christ; Not. syst. 1. 45 (1909) Diplazium calogrammoides (Ching ex W. M. Chu & Z. R. He) comb. ined. Diplazium camarinum (Bak.) C. Chr.; Ind. 229 (1905) Diplazium campbellii (Jenm.) C. Chr.; Ind. 229 (1905) Diplazium caracasanum (Willd.) Kze. ex Moore; Ind. 324 (1861) Diplazium cariomorphum Alderw.; Bull. Buit. III. 6. 196 (1922) Diplazium carnosum H. Christ; Bull. Herb. Boissier sér. 2, 6: 170 (1906) Diplazium caudatum (Cav.) Jermy; Brit. Fern Gaz. 9: 161 (1964) Diplazium cavalerianum (Christ) M.Kato; Bot. Mag. (Tokyo) 90(1017): 31 (1977) Diplazium celtidifolium Kze.; Bot. Zeit. 285 (1845) var. puberulum Stolze; Fieldiana, Bot. 27: 84 (1991) Diplazium centripetale (Bak.) Maxon; Pter. Puerto Rico 441 (1926) Diplazium ceramicum (Miq.) C. Chr.; Ind. 229 (1905) Diplazium ceratolepis (H. Christ) H. Christ; Farnkr. Erde 221 (1897) Diplazium chattagramicum (Clarke) Ching; Lingnan Sc. Journ. 15: 277 (1936) Diplazium chimboanum (Sod.) C. Chr.; Ind. Suppl. 26 (1913) Diplazium chimborazense (Spruce) Christ; Farnkr. 221 (1867) Diplazium chimuense C.D. Adarns; Novon 2: 292 (1992) Diplazium chinense (Bak.) C. Chr.; Ind. 229 (1906) Diplazium chiriquense C.D. Adams; Novon 2: 292 (1992) Diplazium chocoense (Triana) Hier.; Engl. Jahrb. 34. 459 (1904) Diplazium christii C. Chr.; Ind. 229 (1905) Diplazium ciliatum M.G.Price; Contr. Univ. Michigan Herb. 17: 270 (1990) Diplazium cognatum (Hieron.) Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 2: 308, t. 31, E 18 (1962) Diplazium cominsii (Bak.) C. Chr.; Ind. 230 (1905) Diplazium consacense Hieron.; Hedwigia 47. 213 t. 1 f. 3 (1908) Diplazium corderoi (Sod.) Diels; Nat. Pfl. 1(4). 228 (1899) Diplazium cordifolium Bl.; Enum. Pl. Jav. 190 (1828) var. integrifolium (Blume) Mitsuta; Acta Phytotax. Geobot. 36(1-3): 78 (1985) var. pariens (Copel.) C. Chr. (1934) Diplazium costale (Sw.) Presl; Tent. 114 (1836) var. robustum (Sodiro) Stolze; Fl. Ecuador 49(14:5B): 38 (1994) Diplazium costulisorum (Copel.) C. Chr.; Ind. Suppl. 26 (1913) Diplazium crassiusculum Ching; Lingnan Sc. Journ. 15: 279 (1936) Diplazium crenatoserratum (BI.) Moore; Ind. 121, 325 (1859) Diplazium crinitum (Bak.) C. Chr.; Ind. 230 (1905) Diplazium cristatum (Desr.) Alston; J. Bot. 74: 173 (1936) Diplazium croatianum C.D. Adams; Novon 2: 293 (1992) Diplazium cultratum Presl; Epim. 84 (1849-51) Diplazium cultrifolium (L.) Kze.; Bot. zeit. 286 (1845) Diplazium cumingii (Presl) C. Chr.; Ind. 230 (1905) Diplazium cuneifolium Rosenst.; Fedde Repert. 12. 470 (1913) Diplazium curtisii (Holtt.) Holtt.; Fl. Malaysia ed. 2, 2: 637 (1968) Diplazium cyatheifolium (Rich.) Presl; Epim. 84 (1849-51) Diplazium dameriae Pichi-Serm.; Webbia 45(2): 357 (1991) Diplazium davaoense Copel.; Phil. Journ. Sci. 1. Suppl. II. 151 (1906) Diplazium decompositum (Copel.) Parris; Kew Bull. 41(1): 69 (1986) Diplazium decrescens C. Chr.; Kungl. Svenska Vetens.-akad. Handl. ser. 3. 16(2). 47. t.I3(9-13) (1937) Diplazium deltoideum Presl; Tent. 114 (1836) Diplazium dielsii (Brause) Holtt.; Blumea 13: 136 (1965) Diplazium dietrichianum (Luerss.) C. Chr.; Ind. 231 (1905) Diplazium dinghushanicum (Ching & S.H.Wu) comb. ined. Diplazium diplazioides (Klotzsch & H. Karst.) Alston; J. Bot. 74: 174 (1936) Diplazium divaricatum Ching; Bull. Fan Mem. Inst. 2. 208 t. 25 (1931) Diplazium divergens Rosenst.; Fedde Repert. 12. 471 (1913) Diplazium divisissimum (Bak.) Christ; Farnkr. 220 (1897) Diplazium doederleinii (Luerss.) Mak.; Bot. Mag. Tokyo 13. 14 (1899) Diplazium dolichosorum Copel.; Phil. Journ. Sci. 1. Suppl. II. 151 (1906) Diplazium donianum (Mett.) Tard.; Aspl. du. Vietnam 58. t. 5 (1932) var. aphanoneuron (Ohwi) Tagawa (1962) var. lobatum Tag.; Acta Phytotax. Geobot. 10: 290 (1941) Diplazium donnell-smithii H. Christ; Bull. Herb. Boissier sér, 2, 7: 270 (1907) Diplazium drepanolobium A.R. Sm.; Amer. Fern J. 70: 21 (1980) Diplazium dulongjiangense (W.M.Chu) comb. ined. Diplazium dushanense (Ching ex W.M.Chu & Z. R. He) comb. ined. Diplazium ebenirachis (Copel.) Price Diplazium echinatum C. Chr.; Bull. Bishop Mus. 177: 75. t.2(B) (1943) Diplazium egenolfioides M. Price; Brit. Fern Gaz. 10: 258 (1972) Diplazium elatum Fée; Gen. 214 (1850-52) Diplazium ellipticum (Copel.) C. Chr.; Ind. Suppl. III. 73 (1934) Diplazium elmeri (Copel.) C. Chr.; Ind. Suppl. III. 73 (1934) Diplazium entecnum Mickel & Beitel; Pterid. Fl. Oaxaca [Mem. New York Bot. Gard. 46]: 154 (1988) Diplazium esculentoides M.Kato; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 15(4): 326 (1994) Diplazium esculentum (Retz.) Sw.; J. Bot. (Schrader) 1801(2): 312 (1803) Trivial names: Vegetable Fern Diplazium expansum Willd.; Sp. 5. 354 (1810) Diplazium fadyenii (Hook.) Proctor; Bull. Inst. Jamaica Sc. Ser. 5: 24 (1953) Diplazium falcinellum C. Chr.; Gardens Bull. 7. 270 (1934) Diplazium feei Schaffn. ex Fée; 8 mem. 85 (1857) Diplazium fieldingianum (Kunze) Panigr.; Phytologia 31 (3): 251 (1975) Diplazium fimbriichlamys Nakai; Bull. Nation. Sc. Mus. Tokyo 27: 17 (1949) Diplazium flavoviride Alston; Journ. of Bot. 78: 226 (1940) (Oct.), Diplazium flexuosum Presl; Tent. 114 (1836) Diplazium forbesii (Bak.) C. Chr.; Ind. 232 (1905) Diplazium formosanum Rosenst.; Hedwigia 66. 337 (1915) Diplazium forrestii (Ching ex Z.R.Wang) comb. ined. Diplazium fosbergii (Copel.) C. Morton; Contrib. U.S. Nation. Herb. 38: 229 (1973) Diplazium franconis Liebm.; Kongel. Danske Vidensk. Seisk. Skr., Naturvidensk. Math. Afd. ser. 5, 1: 256 (1849) Diplazium fraxinifolium Presl; Rel. Haenk. 1. 49 (1825) Diplazium frondosum (Clarke) Christ.; Not. Syst. Lacombe 1: 46 (1909) Diplazium fructuosum Copel.; Phil. Journ. Sci. 1. Suppl. II. 150 t. 8 (1906) Diplazium fuenzalidae Espin.; Bol. Mus. Nacion. Hist. Nat. [Chile] 20: 28. t.1-7 (1943) Diplazium fuertesii Brause; Urban, Symb. Ant. 7. 486 (1913) Diplazium fuliginosum (Hook.) M. Price; Brit. Fern Gaz. 10: 260 (1972) Diplazium furculicolum Alderw.; Bull. Buit. II. no. 28. 19 (1918) Diplazium fuscum (Bak.) C. Chr.; Ind. 232 (1905) Diplazium geophilum (Copel.) Alderw.; Bull. Jard. bot. Buit. II. nr. VII. 12 (1912) Diplazium gillespiei (Copel.) M.Kato; J. Fac. Sci. Univ. Tokyo, Seet. 3, Bot. 13(4): 389 (1984) Diplazium glingense (Ching & Y.X.Lin) comb. ined. Diplazium godmanii (Bak.) C. Chr.; Ind. 233 (1905) Diplazium gomezianum C.D. Adams; Novon 2: 294 (1992) Diplazium gracilescens (Mett.) C. Chr. Diplazium grandifolium (Sw.) Sw.; J. Bot. (Schrader) 1800 (2): 62 (1801) var. andicola Stolze; Fieldiana, Bot. 27: 82 (1991) Diplazium grashoffii Rosenst.; Fedde Repert. 18. 215 (1914) Diplazium griffithii (T. Moore) Diels; Nat. Pfl. 1(4). 227 (1899)
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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