Family Lomariopsidaceae, genus Dryopteris;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Dryopteris, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

    Dryopteris species H to O.
    Dryopteris species A to G. Here.   |   Dryopteris species P to Z and hybrids. Here.
005.00 Class Filicopsida
		Trivial names: Ferns
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
047.000 Order Dryopteridales
		Statistics: 3 families, 30 genera, 1421-1752 spp. 
		[1989 species and 158 hybrids listed]
049.0000 Family Dryopteridaceae Ching; Acta Phytotax. Sin. 10: 1 (1965) s. str.
		Statistics: 21 genera, 672-939 spp. 
		[1162 species and 147 hybrids listed]
		Revision date: 20.07.2001
		= Aspidiaceae Frank; Leunis, Syn. Pflanzenfam. ed. 2, 3: 1469 [nom. illeg.
		= Dryopteridaceae tribus Stenolepieae H.S. Kung; Proc. Int. Symp. Syst. Pteridol. 
		(eds. K.H. Shing & K.U. Kramer): 172 (1989)
049.0200 Tribus Dryopterideae

049.0201 Dryopteris Adanson; Fam. Pl. 2: 20 (1763) [nom. cons. Trivial names: Wurmfarn Statistics: 250-350 spp., northern temperate zone, tropical mountains, America (22, of them 9 in the American tropics). [476 species and 77 hybrids listed]. Most of the Chinese species are probably nomina nuda or synonyms. Many listed taxa may belong to other genera (Dryopteris used to be an "omnibus genus") Revision date: 20.07.2001 = Acrorumohra (H. Ito) H. Ito; Nakai & Honda, Nov. Fl. Japan 4: 101 (1939) = Anopodium J. Sm.; Cat. cult. ferns 16 (1857) = Arsenopteris Webb & Berthelot; Hist. Nat. Canar. 82. pars 3. 437 (1847) = Arthrobotrys Wallich; List n. 395 (1828) = Catapodium J. Smith; Cat. cult. ferns 16 (1857) = Desmopodium J. Smith ms.; ex Moore, Ind. 308 (1861) = Dichasium (A. Br.) Fée; Mem. Fam. Foug. 5 (Gen. Fil.) 302 (1852) = Diclisodon Moore; Ind. Filicum XCV (1857) = Filix Hill; Family Herbal 171 (1755) = Filix-Mas Hill; British Herbal 527 (1756) = Gleichenia Necker; Element. Bot. 3. 314 (1790) = Gymnothalamium Zenker ms. ex Kunze; Linn. 24. 287 (1851) = Lophodium Newman; Phytologist 4. 371, app. XVI (1851) = Megalastrum J. Smith; Hist. Fil. 216 (1875) = Microchlaena Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 322 (1938) = Microdictyon Fée; 11 mém. 66 (1866) = Monogonia Presl; Tent. Pterid. 146 (1836) = Monophlebia Moore; Index LXXXIX (1857) = Nephrodium Richard; Marthe, Cat. Jard. Méd. Paris 120 (1801) = Oligocampia Trevisan; Atti Istituto Veneto II. 2. 165 (1851) = Pachyderris J. Sm. ms.; ex Moore, Ind. CXLVI (1857) = Parapolystichum Keyserling; Pol. Cyath. Hb. Bung. 11 (1873) = Psidopodium Necker; Element. Bot. 3: 315 (1790) = Pteris Gleditsch; Syst. Pl. 289 (1764) = Pycnopteris Moore; Gard. Chr. 468 (1855) = Syneuron J. Sm. ms. ex Hook.; Ic. plant. t. 950 (1854) Dryopteris habaensis Ching; Sunyatsenia 6: 2 (1941) Dryopteris hadanoi Kurata; J. Geobot. 14: 83 (1966) Dryopteris handeliana C. Chr. Dansk Bot. Ark. 9(3): 62. t.6(5-6) (1937) Dryopteris hangchowensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8 s 414 (1938) Dryopteris hasseltii (BI.) C. Chr.; Ind. 269 (1905) Dryopteris hawaiiensis (Hillebr.) Robinson; Bull. Torr. Cl. 39. 594 (1912) Dryopteris hayatai Tagawa; Acta Phytotax. 1. 156 (1932) Dryopteris herbacea Alderw.; Bull. Jard. bot. Buit. II nr. XX. 13 (1915) Dryopteris heterolepia Alderw.; Bull. Jard. bot. Buit. II nr. XVI. 23 (syn.) (1914) Dryopteris heteroneura (Tagawa) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 398 (1938) Dryopteris himachalensis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 367 (1989) Dryopteris hirticarpa Ching; Bull. Fan Mem. Inst. 2. 197 t. 11 (1931) Dryopteris hirtipes (BI.) O. Ktze.; Rev. Gen. Pl. 2. 813 (1891) Dryopteris hololepis (Hayata) Tagawa; Acta Phytotax. Geobot. 4: 133 (1935) Dryopteris hondoensis Koidzumi; Acta Phytotax. 1. 31 (1932) Dryopteris huangangshanensis Ching; Fl. Fujianica 1: 602 (1982) Dryopteris huanglungensis Ching; Bot. Res. Academia Sinica 1987(2): 10 (1987) Dryopteris huberi (Christ) C. Chr.; Ind. Suppl. 34 (1913) Dryopteris hunanica Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 179 (1940) Dryopteris hupehensis Ching; Fl. Tsinlingensis 2: 226 (1974) Dryopteris hwangii Ching; Fl. Fujianica 1: 600 (1982) Dryopteris hypophlebia Hayata; Icon. Pl. Formosa 4: 154. f. 96 (1914) Dryopteris immixta Ching; Fl. Tsinlingensis 2: 225 (1974) Dryopteris inaequalis (Schlecht.) O. Ktze.; Rev. Gen. Pl. 2. 813 (1891) var. atropaleacea Schelpe (1967) Dryopteris indonesiana Darnaedi; M.Kato & K.Iwats., J. Jap. Bot. 64(I0): 304 (1989) Dryopteris indusiata Mak. & Yamamoto; Suppl. Ic. Pl. Formos. V. 2 (1932) Dryopteris infrahirtella Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 22 (1983) Dryopteris infrapuberula Ching; Boufford & K. H. Shing, J. Arnold Arbor. 64(1): 28 (1983) Dryopteris insularis Kodama; Matsum., lc. Pl. Koisik. 2. 49 t. 109 (1913) var. chichishimensis (Nakai ex Ito) Ito (1939) Dryopteris integriloba C. Chr.; Buit. Dépt. Biol. Coll. Sci. Sun Yatsen Univ. no. 6. 5 (1933) Dryopteris integripinnula Ching; Wuyi Sci. J. 1: 7 (1981) Dryopteris intermedia (Muhl. ex Willd.) A. Gray; Manual 630 (1848) Trivial names: Fancy Fern ssp. intermedia (Muhl. ex Willd.) A. Gray; Manual 630 (1848) ssp. azorica (Christ) Jermy; Webbia 38: 408 (1984) ssp. maderensis (Milde ex Alston) Fraser-Jenk.; Arquipelago, Ser. G. Nat. 2: 85 (1981) Dryopteris inuyamensis H. Itô; Journ. Jap. Bot. 9. 54 (1933) Dryopteris jiangshanensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 34 (1987) Dryopteris jinfoshanensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 6(1): 179 (1986) [nom. nov. Dryopteris jiulungshanensis P.S.Chiu & Yao ex Ching; Bull. Bot. Res., Harbin 2(2): 62 (1982) Dryopteris juxtaposita Christ; Bull. Géogr. Bot. Mans 138 (1907) Dryopteris kaihuaensis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 24 (1983) Dryopteris karwinskyana (Mett.) Kuntze; Revis. Gen. Pl. 2: 813 (1891) Dryopteris khullarii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 362 (1989) Dryopteris kilemensis (Kuhn) O. Ktze.; Rev. Gen. Pl. 2. 813 (1891) (kilmensis) Dryopteris kinkiensis Koidzumi; Tagawa, Acta Phytotax. 2. 200 (1933) Dryopteris kinokuniensis Kurata; J. Geobot. 18: 6 (1970) Dryopteris kobayashii Kitag.; Rep. First. Sc. Exped. Manchoukuo 4(2): 56. f.11 (1935) Dryopteris kodamai Hayata; Ic. Fl. Formosa 4. 156 f. 97 (1914) Dryopteris koidzumiana Tagawa; Acta Phytotax. 2. 190 (1933) Dryopteris komarovii Kossinsky; Not. Syst. Hort. Petr. 2. 1 (1921) Dryopteris kominatoensis Tagawa; Acta Phytotax. Geobot. 6: 154 (1937) Dryopteris koraiensis Tagawa; Acta Phytotax. Geobot. 6: 254 (1936) Dryopteris kwanzanensis Tagawa; Acta Phytotax. Geobot. 7: 186 (1938) Dryopteris kweichowicola Ching ex P.S.Wang; J. Arnold Arbor. 71(1): 119 (1990) Dryopteris labordei (Christ) C. Chr.; Ind. 273 (1905) Dryopteris lacera (Thunb.) Ktze.; Reg. Gen. Pl. 2: 813 (1891) var. chinensis Ching (1938) Dryopteris laeta (Komarov) C. Chr; Ind. 267 (1905) Distribution: E-Siberia / Amur / Ussuri, China (Hebei, Beijing, Shanxi, Shaanxi, Gansu, var. giraldii (H. Christ in Baroni & H. Christ) Ching; Fl. Tsinlingensis 2: 170 (1974) f. triangulare (Nakai) Kitag.; Neo-Lineam. Fl. Manshur.: 35 (1979) Dryopteris laetevirens Rosenst.; Hedwigia 66: 368 (1915) Dryopteris laodianensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 16 (1987) Dryopteris laoshanensis J.X.Li & S.T. Ma; Bull. Bot. Res., Harbin 3(4): 139 (1983) Dryopteris laurentiorum (Christ) C. Chr.; lnd. 664 (1906) Dryopteris laurisilvicola T. Suzuki; Journ. Jap. Bot. 11: 646. f.2 (1935) Dryopteris layardii (Bak.) C. Chr.; Ind. 274 (1905) Dryopteris lepidocaulon Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 19 (1987) Dryopteris lepidopoda Hayata; Icon. Pl. Formos. 4: 161. f 101 (1914) Distribution [complete]: China (Sichuan, Yunnan, Hubei), Bhutan, Nepal, India (Kashmir, Dryopteris lepinei (Kuhn) O. Ktze.; Rev. Gen. Pl. 2. 813 (1891) Dryopteris lewalleana Pichi-Serm.; Bull. Jard. Bot. Belg. 55(1-2): 158 (1985) Dryopteris liankwangensis Ching; lcon. Fil. Sin. 4: t.180 (1937) Dryopteris linganensis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 23 (1983) Dryopteris lingii Ching; Bot. Res. Academia Sinica 1987(2): 23 (1987) Dryopteris linyingensis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 30 (1983) Dryopteris liui Ching; Bull. Bot. Res., Harbin 4(4): 12 (1984) Dryopteris lividis Ching & Wang; Acta Phytotax. Sin. 8: 168. t.22(26) (1959) Dryopteris liyangensis Ching & Y.C.Lan; Fl. Kiang Su (Rec. Fl. Jiangsu (Kiangsu) Prov.) 1: 466 (1977) Dryopteris longirostrata Ching ex K. H. Shing & J.F.Cheng; Jiangxi Sci. 8(3): 47 (1990) Dryopteris ludens (Bak.) C. Chr.; Ind. 276 (1906) Dryopteris ludoviciana (Kunze) Small; Ferns S.E. States 281 (1938) Trivial names: Southern Wood Fern Dryopteris lunanensis (Christ) C. Chr.; Ind. 276 (1905) Distribution: Tibet, China (Yunnan, Sichuan, Guizhou, Guangxi, Hunan, Gansu), Dryopteris lungjingensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 27 (1987) Dryopteris lungnanensis Ching ex K. H. Shing & J.F.Cheng; Jiangxi Sci. 8(3): 48 (1990) Dryopteris lungquanensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 1 (1987) Dryopteris lungshanensis Ching; Bot. Res. Academia Sinica 1987(2): 5 (1987) Dryopteris lushanensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 20 (1987) Dryopteris macrochlamys (Fée) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 453 (1989) Dryopteris macrolepidota Copel.; Bishop Mus. Bull. 98. 8 t. 7 B (1932) Dryopteris makinoi Koidzumi; Acta Phytotax. 1. 26 (1932) Dryopteris mangindranensis Tardieu; Notul. Syst. [Paris] 15: 161 (1956) Dryopteris manniana (Hook.) C. Chr.; Ind. 276 (1905) Dryopteris manshurica Ching ex Wang & al. in Liou; Herb. Fl. N.-E. China 1: 69 (1958) Dryopteris marginalis (L.) Gray; Man. ed. 1. 632 (1848) Trivial names: Marginal Wood Fern Dryopteris marginata (Wall ex Christ) Christ; Philip. Journ. Sci. 2: 212 (1907) Dryopteris matsuzoana Koidzumi; Bot. Mag. Tokyo 39. 15 (1925) Dryopteris mauiensis C. Chr.; Ind. 277 (1905) Dryopteris maximowiczii (Bak.) O. Ktze.; Rev. Gen. Pl. 2. 813 (1891) Dryopteris maximum (Haines) Raizada & N.P.Chowdhury; Pteridophyte Fl. Dryopteris maxonii Underw. & C. Chr.; Amer. Fern J. 1: 96 (1911) Dryopteris mayebarae Tagawa; Acta Phytotax. Geobot. 7: 185 (1938) Dryopteris medialisora Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 6 (1987) Dryopteris medioxima Koidzumi; Acta Phytotax. 1. 32 (1932) Dryopteris megacarpa Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 3 (1984) Dryopteris melanolepia Alderw.; Bull. Jard. bot. Buit. II nr. XVI. 26. (syn.) (1914) Dryopteris metafuscipes Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 19 (1983) Dryopteris metcalfii Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 463 (1938) Dryopteris microlepioides C. Chr.; Bull. Mus. Paris II. 6. 102 (1934) Dryopteris mimetica Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 21 (1983) Dryopteris mindshelkensis N.Pavl.; Vestn. Akad. Nauk Kaz. SSR. 8: 129. f.31 (1954) Dryopteris mingetsuensis Hayata; Ic. Fl. Formosa 6. 281 f. 109 (1916) Dryopteris minimisora Nakai; Bull. Nation. Sc. Mus. Tokyo 27: 19 (1949) Dryopteris montigena Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 64 (1941) Dryopteris multijugata Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3):12 (1983) Dryopteris munchii A.R. Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 218 (1975) Dryopteris nakanensis Ching; Bull. Fan-Mem. Inst. Biol. Bot. 8: 413 (1938) Dryopteris namegatae (Kurata) Kurata; J. Geobot. 17: 89 (1969) Dryopteris nanchuanensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 20 (1983) Dryopteris napoleonis (Bory) O. Ktze.; Rev. Gen. Pl. 2. 813 (1891) Dryopteris neglecta Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 4 (1984) Dryopteris neoassamensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 480 (1938) Dryopteris neofuscipes Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 22 (1984) Dryopteris neolacera Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 440 (1938) Dryopteris neorosthornii Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 62 (1941) Dryopteris neosordidipes Ching ex K. H. Shing & J.F.Cheng; Jiangxi Sci. 8(3): 48 (1990) Dryopteris nigrisquama Hayata; Ic. Fl. Formosa 4. 167 f. 106 (1914) Dryopteris nigropaleacea (Fraser-Jenk.) Fraser-Jenk.; Bol. Soc. Brot. 55: 238 (1982) Dryopteris nipponensis Koidz.; Mayebara, Fl. Austro-Higoensis 5 (1931), Acta Phytotax. 1. 28 (1932) Dryopteris nobilis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 65 (1941) Dryopteris nubigena Maxon & C.V. Morton; Proc. Biol. Soc. Wash. 50: 179 (1937) Dryopteris nudistipes Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 30 (1987) Dryopteris oblancifolia Tagawa; Acta Phytotax. Geobot. 5: 190 (1936) Dryopteris oblongipinnula Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 5 (1987) Dryopteris obovata (Bak.) C. Chr.; Ind. 280 (1905) Dryopteris occidentalizhejiangensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 17 (1987) Dryopteris odontoloma (Moore) C. Chr.; Act. Hort. Gothob. 1: 59 (1924) Dryopteris odontophora Copel.; Philip. Journ. Sc. 60: 106 (1936) Dryopteris ogawai H.Itô in Nakai; Iconogr. Pl. As. Or. 1: 18. t.9 (1935) Dryopteris okinawensis H.Itô; Bot. Mag. Tokyo 49: 360. f.3,4(3) (1935) Dryopteris oligodonta (Desv.) Pic.Ser.; Webbia 8: 150. f.2 (1951) Dryopteris oligophylloides Sehnem; Fl. Ilustr. Catar. 1 (Aspidiaceas): 124 (1979) [nom. nov. Dryopteris omeicola Ching; Acta Phytotax. Sin.10: 303 (1965) Dryopteris oppositipinna Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 25 (1984) Dryopteris oreades Fomin; Vestn. Tiflissk. Bot. Sada 18: 20 (1911) Trivial names: Kleiner Wurmfarn Dryopteris otomasui Kurata; J. Geobot. 15: 83 (1967) Dryopteris oxyodon (Franch.) Kitag. Rep. First Sc. Exped. Manchoukuo 4(2): 58 (1935)
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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