Family Lomariopsidaceae, genus Dryopteris;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Dryopteris, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

    Dryopteris species P to Z and hybrids.
    Dryopteris species A to G. Here.   |   Dryopteris species H to O. Here.
005.00 Class Filicopsida
		Trivial names: Ferns
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
047.000 Order Dryopteridales
		Statistics: 3 families, 30 genera, 1421-1752 spp. 
		[1989 species and 158 hybrids listed]
049.0000 Family Dryopteridaceae Ching; Acta Phytotax. Sin. 10: 1 (1965) s. str.
		Statistics: 21 genera, 672-939 spp. 
		[1162 species and 147 hybrids listed]
		Revision date: 20.07.2001
		= Aspidiaceae Frank; Leunis, Syn. Pflanzenfam. ed. 2, 3: 1469 [nom. illeg.
		= Dryopteridaceae tribus Stenolepieae H.S. Kung; Proc. Int. Symp. Syst. Pteridol. 
		(eds. K.H. Shing & K.U. Kramer): 172 (1989)
049.0200 Tribus Dryopterideae

049.0201 Dryopteris Adanson; Fam. Pl. 2: 20 (1763) [nom. cons. Trivial names: Wurmfarn Statistics: 250-350 spp., northern temperate zone, tropical mountains, America (22, of them 9 in the American tropics). [476 species and 77 hybrids listed]. Most of the Chinese species are probably nomina nuda or synonyms. Many listed taxa may belong to other genera (Dryopteris used to be an "omnibus genus") Revision date: 20.07.2001 = Acrorumohra (H. Ito) H. Ito; Nakai & Honda, Nov. Fl. Japan 4: 101 (1939) = Anopodium J. Sm.; Cat. cult. ferns 16 (1857) = Arsenopteris Webb & Berthelot; Hist. Nat. Canar. 82. pars 3. 437 (1847) = Arthrobotrys Wallich; List n. 395 (1828) = Catapodium J. Smith; Cat. cult. ferns 16 (1857) = Desmopodium J. Smith ms.; ex Moore, Ind. 308 (1861) = Dichasium (A. Br.) Fée; Mem. Fam. Foug. 5 (Gen. Fil.) 302 (1852) = Diclisodon Moore; Ind. Filicum XCV (1857) = Filix Hill; Family Herbal 171 (1755) = Filix-Mas Hill; British Herbal 527 (1756) = Gleichenia Necker; Element. Bot. 3. 314 (1790) = Gymnothalamium Zenker ms. ex Kunze; Linn. 24. 287 (1851) = Lophodium Newman; Phytologist 4. 371, app. XVI (1851) = Megalastrum J. Smith; Hist. Fil. 216 (1875) = Microchlaena Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 322 (1938) = Microdictyon Fée; 11 mém. 66 (1866) = Monogonia Presl; Tent. Pterid. 146 (1836) = Monophlebia Moore; Index LXXXIX (1857) = Nephrodium Richard; Marthe, Cat. Jard. Méd. Paris 120 (1801) = Oligocampia Trevisan; Atti Istituto Veneto II. 2. 165 (1851) = Pachyderris J. Sm. ms.; ex Moore, Ind. CXLVI (1857) = Parapolystichum Keyserling; Pol. Cyath. Hb. Bung. 11 (1873) = Psidopodium Necker; Element. Bot. 3: 315 (1790) = Pteris Gleditsch; Syst. Pl. 289 (1764) = Pycnopteris Moore; Gard. Chr. 468 (1855) = Syneuron J. Sm. ms. ex Hook.; Ic. plant. t. 950 (1854) Dryopteris pachyphylla Hayata; Ic. Fl. Formosa 4. 168 f. 108 (1914) Dryopteris pacifica (Nakai) Tagawa; Col. Ill. Jap. Pterid. 211 (1959) [non Christ (1912) Dryopteris paleifera Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 2 (1984) Dryopteris pallida (Dory) Maire & Petitmengin; Bull. Séances Soc. Sci Nancy ser. 3, 9: 480 (1908) ssp. pallida (Dory) Maire & Petitmengin; Bull. Séances Soc. Sci Nancy ser. 3, 9: 480 (1908) ssp. balearica (Litard.) Fraser-Jenkins; Candollea 32: 314 (1977) ssp. libanotica (Rosenstock) Nardi; Webbia 32: 97 (1900) ssp. raddeana (Fomin) Fraser-Jenkins; Candollea 32(2): 315 (1977) Dryopteris paludicola Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 21 (1984) Dryopteris panda (Clarke) C. Chr.; Ind. Suppl. III. 94 (1934) Dryopteris paomowanensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 10 (1984) Dryopteris papuana C. Chr.; Ind. 282 (1905) Dryopteris parachinensis Ching & F.Z.Li; Bull. Bot. Res., Harbin 5(1): 157 (1985) Dryopteris parafuscipes Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 23 (1984) Dryopteris paralunanensis S.G.Lu ex W.M.Chu; Guihaia 11(3): 225 (1991) Dryopteris paravaria Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 22 (1987) Dryopteris parrisiae Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 14(3): 199 (1986) Dryopteris patentissima (Franchet) Nair; Indian Forester 94: 169 (1968) Dryopteris patula (Sw.) Underw.; Native Ferns ed. 4, 177 (1893) Dryopteris pauliae Fraser-Jenkins; New Species Syndrome in Indian Pteridology and the Ferns of Nepal 140 (1997) Dryopteris peninsulae Kitag.; Rep. First Sc. Exped. Manchoukuo 4(2). 54 (1935) Dryopteris pentheri (Krasser) C. Chr.; Ind. 284 (1905) Dryopteris permagna M.G.Price; Gard. Bull. Singapore 30: 249 (1977) Dryopteris perrieriana C. Chr.; Bonap., N. Pt. 16. 176 t, 4 (1925) Dryopteris persimilis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 16 (1983) [non (Bak.) C. Chr. Dryopteris phaeolepis Hayata; Ic. Fl. Formosa 4. 169 t 109 (1914) Dryopteris podophylla (Hook.) O. Ktze.; Rev. Gen. Pl. 2. 813 (1891) Dryopteris poilanei Tardieu; Bull. Soc. Bot. France 87: 368 (1941) Dryopteris polita Rosenst.; Fedde Repert. 18. 218 (1914) Dryopteris polylepis (Franch. & Sav.) C. Chr.; Ind. 285 (1905) Dryopteris porosa Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 460 (1938) Dryopteris pseudatrata Ching; Fl. Fujianica 1: 601 (1982) Dryopteris pseudobissetiana Ching ex K.H. Shing & J.F.Cheng; Jiangxi Sci. 8(3): 49 (1990) Dryopteris pseudodontoloma Ching; Fl. Xizangica 1: 265 (1983) Dryopteris pseudoerythrosora Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 27 (1983) [non Kodama 1913 Dryopteris pseudofilix-mas (Fée) Rothm.; Candollea 10: 96 (1945) Dryopteris pseudofuscipes Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 3 (1987) Dryopteris pseudolunanensis Tagawa; Acta Phytotax. Geobot. 9: 145 (1940) Dryopteris pseudoparasitica Alderw.; Nova Guinea 14. 19 (1924) Dryopteris pseudo-sabaei Hayata; Ic. Fl. Formosa b. 283 f. 110 (1915) Dryopteris pseudosieboldii Hayata; Ic. Fl. Formosa 4. 171 f. 111 (1914) Dryopteris pseudosikkimensis Ching & S. K. Wu; Fl. Xizangica 1: 267 (1983) Dryopteris pseudosparsa Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 489 (1938) Dryopteris pseudouniformis Ching; Fl. Fujianica 1: 601 (1982) Dryopteris pudouensis Ching; Bull. Bot. Res., Harbin 3(3): 11 (1983) Dryopteris pulvinulifera (Bedd.) O. Ktze.; Rev. Gen, Pl. 2: 213 (1891) Dryopteris purpurascens (Blume) Christ; Phil. Journ. Sci. Bot. 2. 213 (1907) Dryopteris purpurella Tagawa; Acta Phytotax. 1. 307 (1932) Dryopteris pycnopteroides (Christ) C. Chr.; Ind. Suppl. 38 (1913) Dryopteris qinyuanensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 7 (1987) Dryopteris quadrifida Ching ex K. H. Shing & J.F.Cheng; Jiangxi Sci. 8(3), 49 (1990) Dryopteris quatanensis Ching; Wuyi Sci. J. 1: 6 (1981) Dryopteris ramosa (Hope) C. Chr.; Ind. Fil. 266 (1905) Dryopteris rarissima Kurata; J. Geobot. 13: 43 (1964) Dryopteris raynalii Tard.; Adansonia 5: 200, t. 2, f. 1-4 (1965) Dryopteris redactopinnata Basu & Panigr.; Indian J. For. 3 (3): 270. t. 2. f. 39 6 (1980) Dryopteris reflexa Ching; Bull. Fan Mem. Inst. 12. 193. t. 8 (1931) Dryopteris reflexipinna Hayata; Ic. Fl. Formosa 4. 174 f. 113 (1914) Dryopteris reflexosquamata Hayata; Icon. pl. Formos. 4 176 f. 114 (1914) Dryopteris remota (Döll) Druce; List Brit. Pl.: 87 (1908) Trivial names: Entferntfiedriger Dornfarn Dryopteris remotipinnula Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 15 (1983) [nom. illeg., non Bonap. 1925 Dryopteris remotipinnula Bonap.; N. Pt. 16. 178 (1925) Dryopteris remotissima (Christ) Koidzumi; Bot. Mag. Tokyo 43. 386 (1929) Dryopteris retrorsopaleacea Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 20 (1983) Dryopteris rheophila Mitsuta ex Darnaedi; M.Kato & K.Iwats., J. Jap. Bot. 64(10): 302 (1989) Dryopteris rhomboideo-ovata H. Itô; Journ. Jap. Bot. 9. 55 (1933) Dryopteris rigidiuscula Ching ex K. H. Shing & J.F.Cheng; Jiangxi Sci. 8(3): 47 (1990) Dryopteris rosea (E.Fourn.) Mickel & Beitel; Pterid. Fl. Oaxaca (Mem. New York Bot. Gard. 46): 165 (1988) Dryopteris rossii C. Chr. in H. Ross; Mem. Acad. Nac. Ci. "Antonio Alzate" 32: 178 (1912) Trivial names: Ross' Wood Fern Dryopteris rosthornii (Diels) C. Chr.; Ind. 289 (1905) Dryopteris rubrobrunnea W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 46 (1992) Dryopteris rufosquamosa Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 6 (1987) Dryopteris ruwenzoriensis C. Chr. ex Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 14(3): 204 (1986) Dryopteris ryo-itoana Kurata; J. Geobot. 15: 84 (1967) Dryopteris sabaei (Franch. & Sav.) C. Chr.; Ind. 290 (1905) Dryopteris sacrosancta Koidzumi; Bot. Mag. Tokyo 88. 108 (1924) Dryopteris saffordii C. Chr.; Amer. Fern Journ. 1. 94 (1911) Dryopteris sandwicensis (Hook. & Arn.) C. Chr.; Ind. 290 (1905) Dryopteris sanmingensis Ching; Fl. Fujianica 1: 600 (1982) Dryopteris saxifraga H.Itô; Bot. Mag. Tokyo 60: 125 (1936) Dryopteris scabripes Ching; Bot. Res. Academia Sinica 1987(2): 9 (1987) Dryopteris schimperiana (Hochst.) C. Chr.; Ind. 291 (1905) Dryopteris schnellii Tardieu; Notul. Syst. [Paris] 13: 370 (1949) Dryopteris scottii (Bedd.) Ching; Bull. Dept. Biol. Col. Sci. Sun. Yatsen Univ. 6: 3 (1933) Dryopteris semipinnata Ching; Fl. Tsinlingensis 2: 226 (1974) Dryopteris sericea C. Chr.; Bot. Gazette 64. 336 (1913) Dryopteris serratodentata (Bedd.) Hayata; Ic. Pl. Taiwan 4. 179 f. 116 (1914) Dryopteris sessilipinna Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 29 (1987) [nom. illeg., non Copel. Dryopteris shandongensis J.X.Li & F. Li; Acta Phytotax. Sin. 26(5): 406 (1988) Dryopteris shanghaiensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 24 (1987) Dryopteris shangqianensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 20 (1984) Dryopteris shanmenensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 2 (1987) Dryopteris shensicola Ching & Y.T.Hsieh; Fl. Tsinlingensis 2: 229 (1974) Dryopteris shibipedis Kurata; J. Geobot. 14: 84 (1966) Dryopteris shiroumensis Kurata & Nakam.; Journ. Geobot. 9: 54 (1960) Dryopteris shuichangensis P.S.Chiu & Yao ex Ching; Bull. Bot. Res., Harbin 2(2): 63 (1982) Dryopteris sichotensis Komarov; Bull. Jard. Bot. Pierre le Grand Petrop. 16. 146 (1916) Dryopteris sieboldii (Moore) O. Ktze.; Rev. Gen. Pl. 2. 813 (1891) Dryopteris sikkimensis (Bedd.) O. Ktze.; Rev. Gen. Pl. 2 813 (1891) Dryopteris simasakii (H. Itô) Kurata; J. Geobot. 18: 5 (1970) var. paleacea (H. Ito) Sa. Kurata (1970) Dryopteris sinica Christ; Not. syst. 1. 38 (1909) Dryopteris sinobissetiana Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 8 (1984) Dryopteris sinodickinsii Ching ex K. H. Shing & J.F.Cheng; Jiangxi Sci. 8(3): 46 (1990) Dryopteris sinoerythrosora Ching & K.H. Shing; Fl. Fujianica 1: 600 (1982) Dryopteris sinosparsa Ching & K. H. Shing; Fl. Fujianica 1: 601 (1982) Dryopteris sinovaria Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 6(1): 179 (1986) [nom. nov. Dryopteris siranensis Nakai; Bull. Nation. Sc. Mus. Tokyo 33: 3 (1953) Dryopteris sledgei Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 361 (1989) Dryopteris sordidipes Tagawa; Acta Phytotax. Geobot. 3: 29 (1934) Dryopteris sparsa (D. Don) O. Ktze.; Rev. Gen. Pl. 2. 613 (1891) var. viridescens (Baker in Hook. & Baker) Ching (1938) Dryopteris splendens (Hook.) O. Ktze.; Rev. Gen. Pl. 2. 813 (1891) Dryopteris squamistipes Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 7 (1984) Dryopteris stenochlamys Ching ex K. H. Shing & J.F.Cheng; Jiangxi Sci. 8(3): 49 (1990) Dryopteris stenolepis (Bak.) C. Chr.; Ind. 294 (1905) Dryopteris stewartii Fraser-Jenk.; Kalikasan 7(3): 272 (1978 publ. 1979) Dryopteris subarborea (Bak.) C. Chr.; Ind. 295 (1905) Dryopteris subassamensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 451 (1938) Dryopteris subatrata Tagawa; Acta Phytotax. Geobot. 9: 207 (1940) Dryopteris subbipinnata W.H.Wagner & Hobdy; Contr. Univ. Michigan Herb. 19: 69 (1993) Dryopteris subchampionii Ching; Bot. Res. Academia Sinica 1987(2): 14 (1987) Dryopteris subconformis C. Chr.; Engl. Jahrb. 66. 47 (1933) Dryopteris subconjugata S.G.Lu; Guihaia 11(3): 224 (1991) Dryopteris subconjuncta Christ; Not. syst. 1. 185 c. fig (1910) Dryopteris subcrenulata (Bak.) C. Chr.; Ind. 295 (1905) Dryopteris subcuspidata Rosenst.; Med. Rijks Herb. no. 31. 7 (1917) Dryopteris subexaltata (Christ) C. Chr.; Ind. 295 (1905) Dryopteris subfuscipes Ching ex K. H. Shing & J.F.Cheng; Jiangxi Sci. 8(3): 49 (1990) Dryopteris subimpressa Loyal; Nova Hedwigia 16: 467, t. 177, 178, f. 9-16 (1969) Dryopteris subintegriloba Serizawa; Journ. Jap. Bot. 46: 17f.1a. (1971) Dryopteris sublacera Christ; Lecomte, Not. Syst. 1: 43 (1909) Dryopteris sublaeta Ching & Hsu; Fl. Tsinlingensis 2: 227 (1974) Dryopteris submarginalis Ching, Boufford & K. H. Shing; J. Arnold Arbor. 64(1): 30 (1983) [nom. illeg., non (Langsd. & Fisch.) C. Chr. 1905 Dryopteris submonticola Nakai; Bot. Mag. Tokyo 46. 98 (1931) Dryopteris subpycnopteroides Ching ex Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 14(3): 199 (1986) Dryopteris subtenuicula Ching & P.S.Chiu; Bot. Res. Academica Sinica 1987(2): 29 (1987) Dryopteris subtriangularis (Hope) C. Chr.; Ind. Fil. 296 (1905) Dryopteris sunii Ching; Bull. Fan Mem. Inst. Biol. ser. 2. 1: 295 (1949) Dryopteris supraimpressa Ching; Boufford & K. H. Shing, J. Arnold Arbor. 64(1): 30 (1983) Dryopteris tahmingensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 480 (1938) Dryopteris taishanensis Li; Acta Phytotax. Sin. 20(3): 344 (1982) Dryopteris takeuchiana Koidzumi; Acta Phytotax. 1. 30 (1932) Dryopteris tamiensis Brause; Engl. Jahrb. 49. 23 (1912) Dryopteris taquetii Christ; Fedde Repert. 5. 284 (1908) Dryopteris tarningensis Ching; Wuyi Sci. J. 1: 8 (1981) Dryopteris tenuicula Matthew & Christ; Not. syst. 1. 56 (1909) Dryopteris tenuipes (Rosenstock) Seriz.; J. Jap. Bot. 46: 19 (1971) Dryopteris tetrapinnata W. H. Wagner & Hobdy Dryopteris tetsu-yamanakae Kurata; J. Geobot. 10: 100 (1962) Dryopteris tieanzuensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 24 (1987) Dryopteris tieluensis Ching & Hsu; Fl. Tsinlingensis 2: 228 (1974) Dryopteris tingiensis Ching & S. K. Wu ex Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 14(3): 202 (1986) Dryopteris tokyoensis (Matsum.) C. Chr.; Ind. 298 (1905) Dryopteris tonompokensis C. Chr.; Gardens Bull. S. S. 7. 248 (1934) Dryopteris triangularifrons Ching; Bull. Bot. Res., Harbin 2(2): 64 (1982) Dryopteris tsoongii Ching; Bot. Res. Academia Sinica 1987(2): 14 (1987) Dryopteris tsugiwoi Kurata; J. Geobot. 17: 34 (1969) Dryopteris tsutsuiana Kurata; J. Geobot. 17: 35 (1969) Dryopteris tyrrhena Fraser-Jenkins & Reichst.; Fern Gaz. 11: 180 (1975) Dryopteris undulata O. Ktze.; Rev. Gen. Pl. 2. 814 (1891) Dryopteris unidentata (Hook. & Arn.) C. Chr.; Ind. 299 (1905) var. paleacea (Hillebr.) Fraser-Jenk.; Thaiszia 4(1): 31 (1994) Dryopteris uniformis (Mak.) Mak.; Bot. Mag. Tokyo 23. 145 (1909) var. rufomarginata K. H. Shing; Jiangxi Sci. 8(3): 47 (1990) Dryopteris ursipes Hayata; Ic. Fl. Formosa 5. 291 f. 116 (1915) Dryopteris varia (L.) O. Ktze.; Rev. Gen, Pl. 2: 814 (1891) Dryopteris venosa Ching & S. K. Wu; Fl. Xizangica 1: 264 (1983) Dryopteris vescoi (Drake) C. Chr.; Ind. 300 (1905) Dryopteris villarii (Bell) Woynar ex Schinz & Thell.; Vierteljahrsschr. Naturf. Ges. Zürich 60: 339 (1915) Trivial names: Starrer Wurmfarn var. australe (Ten.) Diklic & V.Nikolic; Fl. SR Srbije 10: 10 (1986) f. fallax (Milde) Niketic; Fl. Serbia 1: 155 (1992) Dryopteris viridis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 474 (1938) Dryopteris wallichiana (Spreng.) Hyl.; Bot. Notis. 1963: 352 (1953) ssp. wallichiana (Spreng.) Hyl.; Bot. Notis. 1963: 352 (1953) ssp. coriacea (Fraser-Jenk.) Fraser-Jenk.; Thaiszia 4(1): 23 (1994) ssp. madrasensis (Fraser-Jenk.) Fraser-Jenk.; Thaiszia 4(1): 22 (1994) ssp. himalayensis Fraser-Jenkins; New Species Syndrome in Indian Pteridology and the Ferns of Nepal 134 (1997) ssp. nepalensis Fraser-Jenkins; New Species Syndrome in Indian Pteridology and the Ferns of Nepal 134 (1997) Dryopteris wallichii Rosenst.; Med. Rijks Herb. no. 31. 6 (1917) Dryopteris wangii Ching; Bot. Res. Academia Sinica 1987(2): 13 (1987) Dryopteris wardii (Bak.) O. Ktze.; Rev. Gen. Pl. 2. 814 (1891) Dryopteris wenchuanensis H.S.Kung; Acta Bot. Yunnan. 4(4): 342 (1982) Dryopteris wherryi F.W. Crane; Amer. Fern Journ. 50: 271. t.23(6) (1960) Dryopteris willsii (Bak.) C. Chr.; Ind. 301 (1905) Dryopteris wilsoni (Bak.) C. Chr.; Ind. 301 (1905) Dryopteris woodsiisora Hayata; Ic. Pl. Taiwan 6, 158 (1916) Dryopteris wuliangshanicola S.G.LU ex W.M.Chu; Guihaia 11(3): 221 (1991) Dryopteris wuyiganensis Ching; Bot. Res. Academia Sinica 1987(2): 15 (1987) Dryopteris wuyishanensis Ching; Fl. Fujianica 1: 600 (1982) Dryopteris wuyishanica Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 34 (1987) Dryopteris wuyishanicola Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 31 (1987) Dryopteris x algonquinensis D.M.Britton; Canad. Field-Naturalist 89(2): 165 (1975) Remarks: Dryopteris fragrans x marginalis Dryopteris x ambroseae Fraser-Jenk. & Jermy; Fern Gaz. 11(5): 338 (1977) Remarks: Dryopteris dilatata (Hoffm.) A. Gray x Dryopteris expansa (C. Presl) Fraser-Jenkins Dryopteris x arecesiae Perez Carro & T.E.Diaz; Anales Jard. Bot. Madrid 47(1). 239 (1989 publ. 1990) Remarks: Dryopteris aemula x corleyi Dryopteris x asturiensis Fraser-Jenk. & Gibby; Fern Gaz. 13(2): 113 (1986) Remarks: Dryopteris affinis ssp. affinis x corleyi Dryopteris x atropalustris Small; Ferns SE. States 274 cum tab (1938) Remarks: Dryopteris celsa x Dryopteris cristata Dryopteris x australis (Wherry) Small; Ferns SE. States 279 cum tab., 477 (1938) Remarks: Dryopteris celsa x Dryopteris ludoviciana Dryopteris x benedictii (Farw.) Wherry; Amer. Fern J. 51: 34 (1961) Remarks: Dryopteris clintoniana x Dryopteris spinulosa Dryopteris x bootii (Tuckerman) Underwood; Our nat. Ferns ed. IV. 11 7 (1893) Remarks: Dryopteris cristata x Dryopteris intermedia Dryopteris x borbasii Litard; Bull. Soc. bot. Deux-Sevres 1909-10. 85 (1910) Dryopteris x brathaica Fraser-Jenk. & Reichest.; Fern Gaz. 11(5): 337 (1977) Remarks: Dryopteris carthusiana (Vill.) H. P. Fuchs x Dryopteris filix-mas (L.) Schott Dryopteris x burgessii Boivin; Nat. Canad. 93: 272 (1966) Remarks: ? Dryopteris cristata var. clintoniana x marginalis Dryopteris x burnatii Christ & Wilcz.; Ann. Cons. Jard. bot. Genève 16-16. 346 t.2, 3 (1913) = Dryopteris dilatata x rigida Christ & Wilczek; Ann. Cons. Jard. bot. Geneve 16. 345 t. 2, 3 (1913) Dryopteris x cebennae Fraser-Jenk.; Candollea 32(2): 318 (1977) Remarks: Dryopteris pseudomas x submontana Dryopteris x cedroensis Gibby & Widén; Fern Gaz. 12(5): 267 (1983) Remarks: Dryopteris guanchica x oligodonta Dryopteris x complexa Fraser-Jenk.; Sommerfeltia 6: xi (1987) nssp. complexa Fraser-Jenk.; Sommerfeltia 6: xi (1987) Remarks: Dryopteris affinis x filix-mas nssp. contorta Fraser-Jenk.; Sommerfeltia 6: xii (1987) Remarks: Dryopteris affinis ssp. cambrensis x filix-mas nssp. critica Fraser-Jenk.; Sommerfeltia 6 (1987) Remarks: Dryopteris affinis ssp. borreri x filix-mas Dryopteris x deweveri (Jansen) Jansen & Wachter; Heukels & Wachter, Geillustr. Schoolfl. Nederl. ed. 11. 93 (1934) Remarks: Dryopteris carthusiana (Vill.) H. P. Fuchs x Dryopteris dilatata (Hoffm.) A. Gray Dryopteris x doeppii Rothm.; Candollea 10: 93 (1945) Remarks: Dryopteris paleacea x spinulosa Dryopteris x doluchanovii Askerov; Izv. Akad. Nauk Azerbaidzhnsk. S.S.R., So. Biol. Med. Nauk. 1978(4): 4 (1978 publ. 1979) Remarks: Dryopteris dilatata (Hoffm.) A. Gray x Dryopteris pseudomas (Wollaston) Holub & Pouzar Dryopteris x dowellii (Farw.) Wherry; Amer. Fern. J. 51: 34 (1961) Remarks: Dryopteris clintoniana x Dryopteris intermedia Dryopteris x euxinensis Fraser-Jenkins & Corley; Brit. Fern Gaz. 10: 225 (1972) Remarks: Dryopteris caucasica (A. Br.) Fraser-Jenkins & Corley x Dryopteris filix-mas (L.) Schott Dryopteris x fraser-jenkinsii Gibby & Widén; Fern Gaz. 12(5): 268 (1983) Remarks: Dryopteris affinis x Parentage uncertain Dryopteris x fujipedis Kurata; J. Geobot. 14: 35 (1965) Remarks: Dryopteris crassirhizoma x lacera Dryopteris x furadensis Bennert, Rasbach, K.Rasbach & Viane; Fern Gazette 14(1): 10 (1991) Remarks: Dryopteris aitoniana x maderensis Dryopteris x ghatakii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 460 (1989) Remarks: Dryopteris austroindica x cochleata Dryopteris x gomerica Gibby & Widén; Fern Gaz. 12(5): 267 (1983) Remarks: Dryopteris aemula x guanchica Dryopteris x gotenbaensis Nakaike; J. Geobot. 17: 36, fig (1969) Remarks: Dryopteris hondoensis x uniformis Dryopteris x graeca Fraser-Jenk. & Gibby; Candollea 35(1): 309 (1980) Remarks: Dryopteris pallida ssp. pallida x submontana Dryopteris x haganecola Kurata; J. Geobot. 18: 4 (1970) Remarks: Dryopteris atrata x uniformis Dryopteris x hakonecola Kurata; Journ. Geobot. 7: 13 (1958) Remarks: Dryopteris dickinsii x uniformis Dryopteris x hisatsuana Kurata; J. Geobot. 13: 42 (1964) Remarks: Dryopteris handeliana x lacera Dryopteris x initialis Fraser-Jenkins & Corley; Brit. Fern Gaz. 10: 230 (1972) Remarks: Dryopteris caucasica (A. Br.) Fraser-Jenkins & Corley x Dryopteris oreades Fomin Dryopteris x laschii E. Walter in Walter & Callé; Pterid. Exsic. 1(2): p. XXXVI (1939) Remarks: Aspidium spinulosum x cristatum Lasch ex Milde Dryopteris x lawalreei Janchen; Cat. Fl. Austr., Pteridophyte & Anthrophyt., Ergänz., 121 (1963) Remarks: Dryopteris carthusiana x tavelii Dryopteris x leedsii Wherry; Bartonia 21: 2. t.1(1) (1942) Remarks: Dryopteris celsa x D. marginalis, Dryopteris goldiana x D. marginalis Dowell Dryopteris x liddarensis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 460 (1989) Remarks: Dryopteris barbigera ssp. barbigera x serratodentata Dryopteris x litardierei Rothm.; Candollea 10: 94 (1945) Remarks: Dryopteris paleacea x pallida Dryopteris x loyalii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 462 (1989) Remarks: Dryopteris caroli-hopei x marginata Dryopteris x macdonellii Fraser-Jenk.; Bull. Brit. Mus. (NaL Hist.) Bot. 18(5): 464 (1989) Remarks: Dryopteris filix-mas x ramosa Dryopteris x madalenae Fraser-Jenk.; Arquipelago, Ser. G. Nat. 2: 96 (1981) Remarks: Dryopteris azorica x crispifolia Dryopteris x mantoniae Fraser-Jenkins & Corley; Brit. Fern Gaz. 10: 230 (1972) Remarks: Dryopteris abbreviata x filix-mas, Dryopteris filix-mas (L.) Schott x D. oreades Fomin Dryopteris x martinsiae Fraser-Jenk.; Arquipelago, Ser. G. Nat. 2: 95 (1981) Remarks: Dryopteris aemula x crispifolia Dryopteris x mituii Seriz.; J. Jap. Bot. 46:19 (1971) Remarks: Dryopteris lacera x uniformis Dryopteris x miyazakiensis Miyam.; J. Jap. Bot. 59(8): 243 (1984) Remarks: Dryopteris commixta x hangchowensis Dryopteris x neowherryi W. Wagner in W. & F. Wagner; Castanea 31: 136, f. 4E-H, 5, 6, 8G, H (1966) Remarks: Dryopteris goldiana x Dryopteris marginalis Dryopteris x picoensis Fraser-Jenk. & Gibby; Arquipelago, Ser. G. Nat. 2: 94 (1981) Remarks: Dryopteris affinis ssp. affinis x azorica Dryopteris x pittsfordensis Slosson; Rhodora 6. 75 (1904) Remarks: Dryopteris marginalis x Dryopteris spinulosa Dryopteris x poyseri Wherry; Amer. Fern Journ. 50: 90 (1960) Remarks: Dryopteris goldiana x Dryopteris spinulosa Dryopteris x pseudoabbreviata Jermy; Brit. Fern Gaz. 10: 12 (1968) Remarks: Dryopteris abbreviata x aemula Dryopteris x pseudocommixta Kurata; J. Geobot. 13: 43 (1964) Remarks: Dryopteris commixta x pycnopteroides Dryopteris x pseudohangchowensis Miyam.; J. Jap. Bot. 59(8): 242 (1984) Remarks: Dryopteris hangchowensis x pycnopteroides Dryopteris x pteridiiformis Christ; Buit. Geogr, Bot. Mans 137 (1907) [pro spec. Remarks: ?Dryopteris caroli-hopei x Dryopteris kunmingensis Dryopteris x remota Hayek; Fl. Steiermark 56 (1908) Dryopteris x sardoa Fraser-Jenkins & Reichst. in Fraser-Jenkins; Reichst. & Vida, Fern Gaz. 11: 193 (1975) Remarks: Dryopteris abbreviata x tyrrhena Dryopteris x sarvelii Fraser-Jenk. & Jermy; Fern Gaz. 11(5): 339 (1977) Remarks: Dryopteris carthusiana (Vill.) H. P. Fuchs x Dryopteris assimilis S. Walker Dryopteris x satsumana Kurata; J. Geobot. 17: 89 (1969) Remarks: Dryopteris commixta x dickinsii Dryopteris x saxifragi-varia Nakai; Journ. Jap. Bot. 18: 286 (1942) Remarks: Dryopteris saxifraga x varia Dryopteris x separabilis Small; Ferns SE. States 284 cum tab (1938) Remarks: Dryopteris celsa x Dryopteris intermedia Dryopteris x shibisanensis Kurata; J. Geobot. 14: 37 (1965) Remarks: Dryopteris atrata x commixta Dryopteris x shorapanensis Askerov; Izv. Akad. Nauk. Azerbaidzhnsk. S.S.R. So. Biol. Med. Nauk. 1978(4): 4 (1978 publ. 1979) Remarks: Dryopteris caucasica (A. Br.) Fraser-Jenkins & Corley x Dryopteris pseudomas (Wollaston) Holub & Pouzar Dryopteris x sjoegrenii Fraser-Jenk.; Arquipelago, Ser. G. Nat. 2: 96 (1981) Remarks: Dryopteris azorica x dilatata Dryopteris x slossonae (Hahne) Wherry; Bartonia 21: 15 (1942) Remarks: Dryopteris cristata x Dryopteris marginalis Dryopteris x subaustriaca Rothm.; Candollea 10: 100 (1945) Remarks: Dryopteris austriaca x filix-mas Dryopteris x sugino-takaoi Kurata; J. Geobot. 14: 36 (1965) Remarks: Diyopteris lacera x polylepis Dryopteris x takachihoensis Miyam.; J. Jap. Bot. 59(8): 240 (1984) Remarks: Dryopteris hangchowensis x uniformis Dryopteris x tauschii (Celak.) Domin; Preslia 13-15: 8 (1936) Remarks: Dryopteris cristata x spinulosa Dryopteris x tavelii Rothm.; Candollea 10: 92 (1945) , Dryopteris affinis (Lowe) Fraser-Jenkins x D. filix-mas (L.) Schott Dryopteris x telesii Fraser-Jenk.; Arquipelago, Ser. G. Nat. 2: 97 (1981) Remarks: Dryopteris crispifolia x dilatata Dryopteris x tokudai Sugim.; Journ. Geobot. 7: 97 (1958) Remarks: Dryopteris polylepis x crassirhizoma Dryopteris x triploidea Wherry; Amer. Fern Journ. 50: 90 (1960) Remarks: Dryopteris intermedia (Willd.) Gray x Dryopteris spinulosa var. fructuosa (Gilbert) Wherry Dryopteris x uliginosa (A.Br. ex Döll) O. Ktze. ex Druce; List Brit. Pl. 87 (1908), et in Hayward's Bot. Pocket-book ed. 13. 259. (1909), Comital Fl. Brit. Isles 383 (1932) Remarks: Dryopteris carthusiana (Vill.) H. P. Fuchs x Dryopteris cristata (L.) A. Gray Dryopteris x vidae Fraser-Jenk. & Gibby; Candollea 35(1): 306 (1980) Remarks: Dryopteris pallida ssp. pallida x villarii ssp. villarii Dryopteris x vidyae Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 466 (1989) Remarks: Dryopteris sparsa x splendens Dryopteris x watanabei Kurata; J. Geobot. 14: 36 (1965) Remarks: Dryopteris crassirhizoma x uniformis Dryopteris x wechteriana Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 466 (1989) Remarks: Dryopteris chrysocoma x nigropaleacea Dryopteris x woynarii Rothm.; Candollea 10: 94 (1945) Remarks: Dryopteris austriaca x paleacea Dryopteris x yamashitae Kurata; J. Geobot. 17: 4 (1969) Remarks: Dryopteris hayatae x sparsa Dryopteris x yuyamae Kurata; J. Geobot. 15: 8 (1966) Remarks: Dryopteris pycnopteroides x uniformis Dryopteris x zygoparentalis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 467 (1989) Remarks: Dryopteris darjeelingensis x scottii Dryopteris xanthomelas (Christ) C. Chr.; Ind. Suppl. 41 (1913) Dryopteris xiangxinensis S.F.Wu & C.M.Zhang; Keys Vasc. Pl. Wuling Mount. (ed. W.T.Wang & al.): 570 (1995) Dryopteris xunwuensis Ching & K. H. Shing; Jiangxi Sci. 8(3): 48 (1990) Dryopteris yakusilvicola Kurata; J. Geobot. 15: 85 (1967) Dryopteris yandongensis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 13 (1983) Dryopteris yaoi Ching; Bull. Bot. Res., Harbin 2(2): 64 (1982) Dryopteris yigongensis Ching; Fl. Xizangica 1: 253 (1983) Dryopteris yongdeensis W.M.Chu ex S.G.Lu; Acta Phytotax. Sin. 28(6): 473 (1990) Dryopteris yonpingensis C. Chr. & Ching ex Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 450 (1938) Dryopteris yoroii Seriz.; J. Jap. Bot. 46: 20 (1971) Dryopteris yushanensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 28 (1987) Dryopteris zayuensis Ching & S. K. Wu; Fl. Xizangica 1: 255 (1983) Dryopteris zhangii Ching; Bull. Bot. Res., Harbin 3(3): 26 (1983) Dryopteris zhenangensis Ching & P.S.Chiu; Bot. Res. Academia Sinica 1987(2): 11 (1987) Dryopteris zinongii Z.R.Wang & Fraser-Jenk.; Acta Phytotax. Sin. 23(5). 345 (1985) Dryopteris zunyiensis Ching; Bot. Res. Academia Sinica 1987(2): 18 (1987)
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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