Family Grammitidaceae, genus Grammitis;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Grammitis, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

005.00 Class Filicopsida
		Trivial names: Ferns
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
050.000 Order Polypodiales
		Statistics: 9 families, 82 genera, 1996-2140 spp. 
		(2293 species and 31 hybrids listed]
		= Order Davalliales
055.0000 Family Grammitidaceae (Presl) Ching; Sunyatsenia 5: 264 (1940)
		Statistics: 18 genera, 561-591 spp. 
		[632 species and 2 hybrids listed]
		Revision date: 20.07.2001
		= Polypodiaceae Tribus Grammitideae Presl; Tent. Pterid. 205 (1836) 

055.0001 Grammitis Sw.; Schrad. J. Bot. 1800(2): 3, 17 (1802) Statistics: ca. 200 spp. centered in tropical Asia (many in New Guinea), several Pacific islands, Australia, Neotropics, Africa, Madagascar, Mascarenes [207 species listed] Revision date: 20.07.2001 = Amphoradenium Desv.; Prodr. 335 (1827) = Lomaphlebia J. Sm.; Hist. Fil. 182 (1875) = Mecosorus Kl.; Linnaea 20: 404 (1847) = Nanopteris Vareschi; Fl. Venez. 1: 881 (1969) = Oreogrammitis Copeland; Philipp. J. Sci. C 12: 64 (1917) Grammitis adspersa (Blume) Blume; Fl. Jav. Fil. 115: t. 48, f. 2 (1828) Grammitis ahenobarba Parris; Blumea 29(1): 142 (1983) Grammitis albosetosa (Bailey) Parris; Bot. J. Linn. Soc. 70: 36 (1975) Grammitis alepidota M. Price; Philippine Agriculturist 57: 34 (1973) Grammitis alta Parris; New Zealand J. Bot. 18(1): 146 (1980) [nom. nov. Grammitis andicola Stolze; Fieldiana, Bot. 32: 103 (1993) Grammitis angustifolia Gilli; Ann. Naturhist. Mus. Wien 81: 24 (1977 publ. 1978) Grammitis archboldii (C. Chr.) Copel.; Philip. Journ. Sc. 80: 186. f. 55 (1952) Grammitis argyrata (Bory) C. Morton; Contrib. U.S. Nation. Herb. 38: 255 (1973) Grammitis attenuata Kunze; Linnaea 24: 251 (1851) [non Polypodium attenuatum H. B. Willd. (1810) Grammitis baldwinii (Bak.) Copel.; Philip. Journ. Sc. 80: 158. f.27 (1952) Grammitis barbatula (Bak. in Hook. & Bak.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 240 (1941) Grammitis basalis (Maxon) Lellinger; Amer. Fern J. 74(2): 58 (1984) Grammitis beddomeana (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 240 (1941) Grammitis billardieri Willd.; Sp. 5: 139 (1810) Trivial names: Common strap fern Grammitis bongorensis (Copel.) Copel.; Philip. Journ. Sc. 80: 208, 232, 271. f.90 (1952) Grammitis brassii Copel.; Philip. Journ. Sc. 60: 112. t.21 (1936) Grammitis brevipila Copel.; Univ. Calif. Publ. Bot. 16: 106 (1929) Grammitis bryophila (Maxon) F. Seym.; Phytologia 31: 172 (1975) Grammitis bufonis L.D. Gómez; Phytologia 52: 154 (1982) Grammitis caespitosa Bl.; Enum. 115 (1828) Grammitis carnosula (Christ) F. Seymour; Phytologia 31: 177 (1975) Grammitis ceratocarpa Copel.; Univ. Calif. Publ. Bot. 18: 223 (1942) Grammitis cervicornis (Alderw.) Adelb.; v.Borss. Waalk., Ann. Bogor. 4: 25 (1960) Grammitis cheesemanii Parris; New Zealand J. Bot. 31(1): 15 (1993) Grammitis christophersenii Copel.; Occ. Pap. Bishop Mus. 15: 89. f.8 (1939) Grammitis ciliata Col.; Tasm. Journ. 2: 166 (1811) Grammitis cincta Copel. ex Parris; Allertonia 7(4): 297 (1997) Grammitis clavipila Parris; Blumea 29(1): 136 (1983) Grammitis clemensiae (Copel.) Parris; Fern Gaz. 12(2): 118 (1980) Grammitis collina Parris; Blumea 29(1): 174 (1983) Grammitis conformis J. Sm.; Hist. Fil. 181 (1875) Grammitis congener Bl.; Enum. 115 (1828) Grammitis conjunctisora (Baker) C. Morton; Contrib. U.S. Nation. Herb. 38: 256 (1973) Grammitis copelandii Tardieu; Notul. Syst. [Paris] 15: 421. t.1(5-8) (1959) Grammitis coredrosora (Alderw.) Copel.; Philip. Journ. Sc. 80: 179. f.49 (1952) Grammitis crenulata Parris; Blumea 29(1): 81 (1983) Grammitis crinifera Parris; Blumea 29(1): 180 (1983) Grammitis crispatula Holtt.; Gard. Bull. Singapore 11: 268 (1947) Grammitis cryptophlebia (Bak.) Copel.; Philip. Journ. Sc. 80: 133 (1952) Grammitis cuencana (Hieron.) C. Morton; Contrib. U.S. Nation. Herb. 38: 111 (1967) Grammitis curtipila Parris; Blumea 29(1): 112 (1983) Grammitis daguensis (Hieron.) C. Morton; Contrib. U.S. Nation. Herb. 38: 98 (1967) Grammitis debilifolia Copel.; Univ. Calif. Publ. Bot. 18: 223 (1942) Grammitis demissa Parris; Blumea 29(1): 39 (1983) Grammitis deplanchei (Bak. in Hook. & Bak.) Copel.; Gen. Fil. 211 (1947) Grammitis dictymioides Copel.; Univ. Calif. Publ. Bot. 18: 223 (1942) Grammitis diminuta (Bak. in Hook. & Bak.) Copel.; Philip. Journ. Sc. 80: 141. f.13 (1952) Grammitis dolichosora (Copel.) Copel.; Philip. Journ. Sc. 80: 188. f.57 (1952) Grammitis dorsipila (Christ. in Warb.) C. Chr. & Tardieu; Notul. Syst. [Paris] 8: 179 (1939) Grammitis duplopilosa Copel.; Univ. Calif. Publ. Bot. 16: 106 (1929) Grammitis ebenina (Maxon) Tardieu; Mém. Inst. Fr. Afr. Noire. 28: 211. t.42(11-12) 1953 Grammitis excelsa Parris; Blumea 29(1): 171 (1983) Grammitis fasciata Bl.; Enum. 116 (1828) Grammitis forbesiana W.H.Wagner; Contr. Univ. Michigan Herb. 19: 70 (1993) Grammitis friderici-et-pauli (Christ) Copel.; Philip. Journ. Sc. 80: 250. f.102 (1952) Grammitis frigida (Ridl.) Copel.; Univ. Calif. Publ. Bot. 18: 223 (1942) Grammitis gameriana (Vareschi) Duek & Lellinger; Amer. Fern J. 68(4): 120 (1978) Grammitis garrettii Parris; Fl. Australia 48: 713 (1998) Grammitis givenii Parris; New Zealand J. Bot. 14(1): 96 (1976) Grammitis glabrata Brownlie; Nova Hedwigia Beih. 55 (Pterid. Fl. Fiji): 348 (1977) Grammitis glossophylla Parris; Blumea 29(1): 47 (1983) Grammitis graminea (Swartz) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 240 (1941) Grammitis graminella (C. Chr.) Sledge; Brit. Fern Gaz. 9: 340 (1967) Grammitis graminifolia Copel.; Univ. Cald. Publ. Bot. 18: 223 (1942) Grammitis gunnii Parris; Fl. Australia 48: 713 (1998) Grammitis habbemensis Copel.; Univ. Calif. Publ. Bot. 18: 223 (1942) Grammitis havilandii (Bak.) Copel.; Gen. Fil. 211 (1947) Grammitis hirtella (Blume) Tuyama; Bot. Mag. Tokyo 61: 125 (1937) Grammitis hirtelloides (Copel.) Copel.; Philip. Journ. Sc. 80: 146. f.18 (1952) Grammitis hirtiformis (Rosenst.) Copel.; Philip. Journ. Sc. 80: 190. f. 59 (1952) Grammitis hispida Copel.; Univ. Calif. Publ. Bot. 18: 224 (1942) Grammitis holophlebia (Bak.) Copel.; Philip. Journ. Sc. 80: 136 (1952) Grammitis holttumii Copel.; Univ. Calif. Publ. Bot. 18: 224 (1942) Grammitis hookeri (Brackenr.) Copel.; Philip. Journ. Sc. 80: 157 (1952) Grammitis imberbis Parris; Blumea 29(1): 128 (1983) Grammitis impressa Copel.; Philip. Journ. Sc. 80: 242. f.95 (1952) Grammitis inconstans (Alderw.) Copel.; Philip. Journ. Sc. 80: 193 (1952) Grammitis insularis Copel.; Univ. Calif. Publ. Bot. 18: 223 (1942) Grammitis interrupta (Bak.) Copel.; Gen. Fil. 214 (1947) Grammitis intromissa (Christ) Parris; Fern Gaz. 12(3): 180 (1981) Grammitis jagoriana (Mett. ex Kuhn) Tagawa; Acta Phytotax. Geobot. 10: 284 (1941) Grammitis jeraiensis Parris; Hooker's Icon. Pl. 40(4): t. 3976 (1990) Grammitis kairatuensis M.Kato & Parris; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 15(2): 129 (1992) Grammitis kinabaluensis (Copel.) Copel.; Philippine J. Sci. 56: 479 (1935) Grammitis knutsfordiana (Bak.) Copel.; Univ. Calif. Publ. Bot. 18: 224 (1942) Grammitis kyimbilensis (Brause ex Brause & Hieron.) Copel.; Philip. Journ. Sc. 80: 257. f.104 (1952) Grammitis lasiosora (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 15 (1940) Grammitis latifolia DeVol in H. L. Li; Liu, T. C. Huang, T. Koyama & DeVol, Fl. Formosa 1: 223 (1975) Grammitis leptopoda (C.H. Wright) Copel.; Philipp. J. Sci. 80: 255 (1952) Grammitis ligulata (Bak. in Hook. & Bak.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 240 (1941) Grammitis limbata Fée; Mém. Foug. 6: 6 (1854) Grammitis locellata (Bak.) Copel.; Occ. Pap. Bishop Mus. 16: 86 (1939) Grammitis loculosa (Alderw.) Copel.; Univ. Calif. Publ. Bot. 18: 223 (1942) Grammitis loheriana (Christ) Copel.; Philip. Journ. Sc. 80: 213. f.77 (1952) Grammitis magellanica Desv.; Berl. Mag. 5. 313 (1811) ssp. magellanica Desv.; Berl. Mag. 5. 313 (1811) f. nana (Brackenr.) de la Sota ex T.R.Dudley; Rhodora 83(836): 483 (1981) ssp. nothofageti Parris; New Zealand J. Bot. 14(1): 100 (1976) Grammitis marginella (Sw.) Sw.; J. Bot. (Schrader) 1800(2): 17 (1801) Grammitis marginelloides (J.W.Moore) Copel.; Occ. Pap. Bishop Mus. 14: 72 (1938) Grammitis marivelesensis Copel.; Univ. Calif. Publ. Bot. 18: 224 (1942) Grammitis marquesensis Parris; Allertonia 7(4): 297 (1997) Grammitis maireaui Copel.; Occ. Pap. Bishop Mus. 14: 71. t.22 (1938) Grammitis maxwellii (Baker) Parris; Fern Gaz. 12(2): 118 (1980) Grammitis medialis (Bak. in Hook. & Bak.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 15 (1940) Grammitis meijer-dreesii Copel.; Univ. Calif. Publ. Bot. 18: 223 (1942) ('myer-dreesii') Grammitis melanoloma (Cordem.) Tardieu; Notul. Syst. [Paris] 15: 423. t.1(1-4) 1959 Grammitis merrillii (Copel. in Perkins) Copel.; Philip. Journ. Sc. 80: 224 (1952) Grammitis mesocarpa (Alderw.) Copel.; Philip. Journ. Sc. 80: 181. f.51 (1952) Grammitis microglossa (C. Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 240 (1941) Grammitis miltiblepharis Copel.; Occ. Pap. Bishop Mus. 14: 71. t.23 (1938) Grammitis moelleri (Baker ex Henriq.) Schelpe; Bol. Soc. Brot. Ser. 2, 40: 162 (1966) Grammitis mollipila (Bak.) Copel.; Univ. Calif. Publ. Bot. 18: 224 (1942) Grammitis montana Parris; Blumea 29(1): 167 (1983) Grammitis monticola Sledge; Brit. Fern Gaz. 9: 344, f. 2, t. 3 (1967) Grammitis multifolia Copel.; Phil. Journ. Sci. 46. 219 (1931) Grammitis murrayana (C. Chr.) Copel.; Philip. Journ. Sc. 80: 200. f.71 (1952) Grammitis nanodes (A. Peter) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 241 (1941) Grammitis neocaledonica Copel.; Univ. Calif. Publ. Bot. 18: 222 (1942) Grammitis nigrocincta Alston; Exell, Bull. Inst. Fr. Afr. Noire ser. A. 21: 444 (1959) Grammitis nigropaleata Copel.; Univ. Calif. Publ. Bot. 18: 224 (1942) Grammitis nipponica Tag. & Iwatsuki; Amer. Fern J. 51: 150, f A-C (1961) Grammitis nuda Tagawa; Acta Phytotax. Geobot. 10: 284 (1941) Grammitis nudicarpa Copel.; Univ. Calif. Publ. Bot. 18: 222 (1942) Grammitis oblanceolata (Bak.) Copel.; Philip. Journ. Sc. 80: 166. f.34 (1952) Grammitis obscura Bl.; Enum. 115 (1828) Grammitis obtusa Willd. ex Kaulf.; Enum. 94 (1824) Grammitis oncobasis M.G.Price; Contr. Univ. Michigan Herb. 17: 274 (1990) Grammitis ornatissima (Rosenst.) Copel.; Philip. Journ. Sc. 80: 195. f.65 (1952) Grammitis padangensis (Bak.) Copel.; Philip. Journ. Sc. 80: 212. f.75 (1952) Grammitis palauensis Hosok.; Trans. Nat. Hist. Soc. Taiwan 31: 475 (1941) Grammitis papuensis (Alderw.) Parris; Fern Gaz. 12(2): 118 (1980) Grammitis paramicola L.E. Bishop; Amer. Fern J. 67: 105 (1977) Grammitis parva (Brause) Copel.; Gen. Fil. 211 (1947) Grammitis parvula (Bory) C. Morton; Contrib. U.S. Nation. Herb. 38: 261 (1973) Grammitis patagonica (C. Chr.) Parris; New Zealand J. Bot. 14(1): 108 (1976) Grammitis pellucidovenosa (Bonap.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 241 (1941) Grammitis peninsularis Copel.; Univ. Calif. Publ. Bot. 18: 224 (1942) Grammitis peritimundi L.E.Bishop & A. R. Sm.; Ann. Missouri Bot. Gard. 77(2): 258 (1990) Grammitis pervillei (Mett. ex Kuhn) Tardieu; Notul. Syst. [Paris] 15: 424. t.1(12-14) 1959 Grammitis pilifera Ravi & J.Joseph; J. Bombay Nat. Hist. Soc. 76(2): 348 (1979 publ. 1980) Grammitis pilosiuscula Bl.; Enum. 115 (1828) Grammitis piruensis M.Kato & Parris; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 15(2): 127 (1992) Grammitis plana (Alderw.) Parris; Fern Gaz. 12(3): 180 (1981) Grammitis pleiosora (Mett. ex Kuhn) Copel.; Philip. Journ. Sc. 56: 105 (1935) Grammitis pleurogrammoides (Rosenst.) Copel.; Occ. Pap. Bishop Mus. 15: 86 (1939) Grammitis poeppigiana (Mett.) Pichi-Serm.; Webbia 32(2): 461 (July 1978) Grammitis ponapensis Copel.; Occ. Pap. Bishop Mus. 16: 88. f.7 (1939) Grammitis pseudoaustralis Fourn.; Bull. Soc. Fr. 16. 423 (1869) Grammitis pseudociliata Parris; New Zealand J. Bot. 14(1): 106 (1976) Grammitis pseudolocellata Parris; Blumea 29(1): 189 (1983) Grammitis pseudoxiphopteris Parris; Hooker's Icon. Pl. 40(4): t. 3975 (1990) Grammitis pubinervis Bl.; Enum. 116 (1828) Grammitis pygmaea (Mett. ex Kuhn) Copel.; Philip. Journ. Sc. 80: 138 (1952) Grammitis queenslandica Parris; Bot. J. Linn. Soc. 70: 39 (1975) Grammitis raiateensis (J.W.Moore) Copel.; Occ. Pap. Bishop Mus. 14: 72 (1938) Grammitis rawlingsii Parris; New Zealand J. Bot. 14(1): 105 (1976) Grammitis recondita C. Morton; Phytologia 22: 78 (1971) Grammitis reducta (Alderw.) Copel.; Philip. Journ. Sc. 80: 167. f.35 (1952) Grammitis reinwardtii Bl.; Enum. addend (1828) Grammitis reinwardtioides Copel.; Philip. Journ. Sc. 56: 478. t.10 (1935) Grammitis reptans Parris; Blumea 29(1): 156 (1983) Grammitis rigida Hombr. & Jacq.; Voy. Pole sud t. 2 f. F (1852) Grammitis rivularis Parris; Kew Bull. 41(3): 514 (1986) Grammitis rupestris Parris; Blumea 29(1): 172 (1983) Grammitis salticola Parris; Blumea 29(1): 92 (1983) Grammitis samoensis (Bak. in Hook. & Bak.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 241 (1941) Grammitis scabristipes (Bak.) Copel.; Univ. Calif. Publ. Bot. 18: 223 (1942) Grammitis scleroglossoides Copel.; Occ. Pap. Bishop Mus. 16: 86. f.6 (1939) Grammitis sechellarum (Baker) C. Morton; Amer. Fern J. 60: 124, 1970 Grammitis setosa (Kaulf.) Presl; Tent. 208 (1836) [non Bl. (1828) Grammitis silvicola Parris; Blumea 29(1): 141 (1983) Grammitis solida (Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 241 (1941) Grammitis stenophylla Parris; Bot. J. Linn. Soc. 70: 35 (1975) Grammitis stipitata Proctor; Rhodora 68: 467 (1966) Grammitis subdichotoma (Racib. ex Alderw.) Parris; Fern Gaz. 12(3): 180 (1981) Grammitis subfasciata (Rosenst.) Copel.; Philip. Journ. Sc. 80: 170. f.39 (1952) Grammitis subreticulata (Copel.) Copel.; Philip. Journ. Sc. 80: 185. f.54 (1952) Grammitis subscabra (Klotzsch) C. Morton; Phytologia 22: 80 (1971) Grammitis subspathulata (Brackenr.) Farwell; Amer. Midl. Nat. 12. 294 (1931) Grammitis sumatrana (Bak.) Copel.; Philip. Journ. Sc. 66: 105 (1935) Grammitis synsora (Bak.) Copel.; Philip. Journ. Sc. 80: 137 (1952) Grammitis taeniophylla Parris; Blumea 29(1): 67 (1983) Grammitis tahitensis (C. Chr.) Copel.; Philip. Journ. Sc. 80: 153. f.24 (1952) Grammitis tegetiformis L.E.Bishop; Amer. Fern J. 67(4): 105 (1977) Grammitis tehoruensis M.Kato & Parris; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 15(2): 131 (1992) Grammitis tenella Kaulf.; Enum. 84 (1824) [non Polypodium tenellum Forst (1786) Grammitis tenuis Parris; Blumea 29(1): 130 (1983) Grammitis tomaculosa Parris; Blumea 29(1): 152 (1983) Grammitis tomensis Schelpe; Contrib. Bolus Herb. 1: 6 (1969) Grammitis torricelliana (Brause) Parris; Fern Gaz. 12(3): 180 (1981) Grammitis trachycarpa (Mett. ex Kuhn) Copel.; Occ. Pap. Bishop Mus. 14: 72 (1938) Grammitis trichopoda (F.v.Muell.) Copel.; Univ. Calif. Publ. Bot. 18: 224 (1942) Grammitis trogophylla Copel.; Univ. Calif. Publ. Bot. 18: 224 (1942) Grammitis tuberculata Parris; Blumea 29(1): 99 (1983) Grammitis turquina (Maxon) Copel.; Gen. Fil. 211 (1947) Grammitis tuyamae H. Ohba; Sci. Rep. Tohoku Univ. 4, 36: 121. 1971 Grammitis uapensis (E. D. Br.) Parris; Allertonia 7(4): 297 (1997) Grammitis universa (Bak.) Copel.; Philip. Journ. Sc. 80: 218 (1952) Grammitis vaupelii (Brause) Copel.; Philip. Journ. Sc. 80: 265 (1952) Grammitis viridula (Alderw.) Parris; Fern Gaz. 12(2): 118 (1980) Grammitis vitiensis Brownlie; Nova Hedwigia Beih. 55 (Pterid. Fl. Fiji): 346 (1977) Grammitis vittariifolia (C. Chr.) Parris; Fern Gaz. 12(2): 118 (1980) Grammitis wallii (Bedd.) Copel.; Philip. Journ. Sc. 80: 240 (1952) Grammitis wattsii Copel.; Univ. Calif. Publ. Bot. 18: 222 (1942) Grammitis whitmeei (Bak.) Copel.; Philip. Journ. Sc. 80: 150. f.22 (1952) Grammitis wurunuran Parris; Bot. J. Linn. Soc. 70: 40 (1975) Grammitis youngii Stolze; Fieldiana. Bot. 32: 97 (1993) Grammitis zeylanica Fée; Gen. 234 (1850-52)
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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