Family Lycopodiaceae, genus Huperzia;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Huperzia, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.
          Huperzia species A to K.
          For Huperzia species L to Z including hybrids go here


001.00 Class Lycopsida Trivial names: Lycophytes Statistics: 3 orders, 3 families, 6 genera, 1313 spp. [1321 species and 28 hybrids listed] 001.000 Order Lycopodiales Statistics: 1 family, 483 spp. [487 species and 12 hybrids listed] 001.0000 Family Lycopodiaceae Beauv. ex Mirb.; Hist. Nat. Veg. (Lam. & Mirb.) 4: 293 (1802) Trivial names: Bärlappgewächse Statistics: 4 genera, 483 spp. [487 species and 12 hybrids listed] = Huperziaceae Rothm.; Fedde Repert. 66: 236 (1962) [nom. nov. = Urostachyaceae Rothm.; Fedde Repert. 54: 58 (1944) [nom. illeg. = Phylloglossaceae Kze.; Bot. Zeit. 1: 722 (1843) 001.0002 Huperzia Bernhardi; Schrad. J. Bot. 1800(2): 126 (1801)
Statistics: ca. 390 spp. nearly worldwide, Europe (1) [394 species and 6 hybrids listed] Revision date: 20.07.2001; 19.09.2006 Trivial names: Teufelsklaue = Mirmau Adans.; Fam. Pl. 2: 491. Jul-Aug (1763) [p. p., nom. illegit. = Phlegmariurus (Herter) Holub; Preslia 36 (1): 179 21 (1964) = Plananthus P. Beauv. ex Mirbel; Lam. & Mirbel, Hist. nat. vég. 3: 476, 4: 310 (1802) [nom. superfl., by lectotypification of Huperzia Bernh. = Selago Boehmer; Ludwig, Defin. Gen. Pl. ed. Boehmer 484. 1760 [non L. 1753, nec Browne 1756, nec Hill (1757) nom. illeg. = Selago J. Hill; Brit. Herb. 533. 1757 (28 jan.) [non L. 1753, nec Browne 1756] nom. inval. (Pichi Sermolli (1971: 140)) = Selago Schur; Enum. pl. Transsilv. 825. 1866 [non L. 1753, nec Browne 1756, nec Hill (1757]) = Urostachys Herter; Beih. Bot. Centralbl. 39(2): 249 (1922) [nom. superfl. Huperzia acerosa (Sw.) Holub; Folia Geobot. Phytotax. 20: 70 (1985) Distribution [complete]: Costa Rica, Guatemala, Panama, Colombia, Venezuela, Ecuador, Argentina (Misiones), Brazil Remarks: verticillata agg. = Lycopodium acerosum Sw.; Fl. Ind. Occid. 3: 1575 (1806) = Urostachys acerosus (Sw.) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 399, t. 18 (1927) Huperzia acifolia (Rolleri) Rolleri & Deferrari; Notas del Museo de la Plata, Botánica 21(100): 155 (1988) Distribution [complete]: Peru, Ecuador, Venezuela, Colombia, Bolivia Remarks: reflexa agg. = Lycopodium acifolium Rolleri; Notas Mus. La Plata, Bot. 14(87): 2, t. 1-2 (1985) Huperzia acuta (Rolleri) Rolleri & Deferrari; Notas del Museo de la Plata, Botánica 21(100): 155 (1988) Distribution: Bolivia Remarks: saururus agg. = Lycopodium acutum Rolleri; Notas Mus. La Plata, Bot. 13(71): 61-114 (1981) Huperzia acutifolia (Desv ex Poir.) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution [complete]: Mauritius Remarks: hippuris agg. = Lycopodium acutifolium Desv. ex Poiret; Encycl. suppl. 3: 559. 1813 [1814] = Lycopodium acutifolium Desv.; Encyclopédie Méthodique, Botanique , Suppl., 3: 559 (1813) Huperzia affinis Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) [nom. nov. Distribution [complete]: Ecuador, Peru, Colombia Remarks: reflexa agg. = Huperzia blepharodes (Maxon) Holub; Folia Geobot. Phytotax. 20: 71 (1985) [nom. superfl. for H. affinis Trevisan = Lycopodium affine Grev. & Hook.; Botanical Miscellany 2: 364 (1831) [nom. illeg., non Lycopodium affine Bory (1804) = Lycopodium blepharodes Maxon; Contributions from the U. S. National Herbarium 17(4): 423 (1914) = Urostachys affinis (Grev. & Hook.) Nessel; Repert. Spec. Nov. Regn. Veg. 39: 69 (1935) = Urostachys blepharodes (Maxon) Herter; Estudios Botánicos en la Región Uruguaya 20: 53 (1949) = Urostachys involutus Herter ex Nessel; Revista Sudamericana de Botánica 6(5-6): 162, t. 10, f. 39 (1940) Huperzia afromontana Pichi-Serm.; Webbia 27: 394, fig. 2 (1972 [1973]) Distribution [complete]: Burundi, Uganda, Tanzania (Mt. Kilimanjaro), E-Zaire, Rwanda, Malawi, Kenya Remarks: ophioglossoides agg. Huperzia albescens (Bailey) Holub; Folia Geobot. Phytotax. 20: 70 (1985) Distribution: New Guinea Remarks: carinata agg. = Lycopodium albescens Bailey; Queensl. Dept. Agric. Brisbane Bot. Bull. 14: 16 (1896) = Urostachys albescens (Bailey) Herter; Revista Sudamer. Bot. 8: 86 (1949) Huperzia aloifolia (Hook. & Grev.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution: E-India Remarks: hamiltonii agg. = Lycopodium aloifolium Wallich ex Hook. & Grev.; Icon. filic. 2: t. 233. 1831 [jun (1832]) = Lycopodium aloifolium Zenker (Herter 1949a: 3) = Urostachys aloifolius (Hook. & Grev.) Herter ex Nessel; Bärlappgewächse 70 (1939) Huperzia amentacea (B. Øllgaard) Holub; Folia Geobot. Phytotax. 20: 70 (1985) Distribution [complete]: Guatemala, Colombia, Venezuela Remarks: myrsinites agg. = Lycopodium amentaceum B. Øllgaard; Amer. Fern Journ. 72: 53 (1982) Huperzia andina (Rosenst.) Holub; Folia Geobot. Phytotax. 20: 70 (1985) Distribution [complete]: Peru, Bolivia = Lycopodium andinum Rosenst.; Repert. Spec. Nov. Regn. Veg. 5: 239 (1908) = Urostachys andinus (Rosenst.) Nessel; Die Bärlappgewächse 76 (1939) Huperzia apolinari-mariae (Nessel) Holub; Folia Geobot. Phytotax. 20: 70 (1985) Distribution [complete]: Colombia Remarks: selago agg. = Urostachys apolinari-mariae Nessel; Repert. Spec. Nov. Regni Veg. 36: 177, t. 170 (1934) Huperzia appalachiana Beitel & Mickel; Amer. Fern J. 82: 45 (1992) Distribution [complete]: Greenland, Canada (Quebec, Nova Scotia, Newfoundland, Labrador, Ontario), Saint Pierre & Miquelon, USA (Georgia, Maine, Massachusetts, Michigan, Minnesota, New Hampshire, New York, North Carolina, South Carolina, Tennessee, Vermont, Virginia) Huperzia aqualupiana (Spring) Rothm.; Feddes Repert. 54: 62 (1944) Distribution [complete]: Colombia, Venezuela, Trinidad, Puerto Rico, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent Remarks: aqualupiana agg. = Lycopodium aqualupianum Spring; Bulletin de l'Academie Royale des Sciences & Belles-lettres de Bruxelles 8: 518 (1841) = Urostachys aqualupianus (Spring) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 166 (1923) = Lycopodium guadalupianum Spring ex Fée; Hist. foug. Ant. 1310 t. 33p f. 1 (1866) [nom. superfl. Huperzia archboldiana (Nessel) Holub; Folia Geobot. Phytotax. 20: 70 (1985) Distribution: New Guinea Remarks: selago agg. = Urostachys archboldianus Nessel; Repert. Spec. Nov. Regni Veg. 39: 61, t. 189a (1935) = Lycopodium archboldianum (Nessel) Alston; J. Bot. 77: 290 (1939) Huperzia arcuata B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 15, f. 1D (1988) Distribution [complete]: Ecuador, Peru, S-Colombia Remarks: brongniartii agg. Huperzia aristei (Nessel) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 155 (1988) Distribution [complete]: Colombia Remarks: taxifolia agg. = Urostachys aristei Nessel; Repert. Spec. Nov. Regni Veg. 36: 181, t. 172 (1934) = Lycopodium aristei (Nessel) Rolleri; Revista Mus. La Plata (N. S.) Bot. 13: 94 (1981) Huperzia ascendens (Nessel) Holub; Folia Geobot. Phytotax. 20: 70 (1985) Distribution: Ecuador Remarks: saururus agg. = Urostachys ascendens Herter ex Nessel; [Bärlappgewächse 74. 1939 [nom. inval.] Revista Sudamer. Bot. 6: 161, t. 9, f. 30 (1940) Huperzia attenuata (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1874) Distribution [complete]: Costa Rica, Colombia, Ecuador, Peru, Bolivia Remarks: saururus agg. = Lycopodium attenuatum Spring; Mém. Acad. Roy. Sci. Belgique 24: 8 (1849) = Lycopodium tobarii Sodiro; Crypt. vasc. quit. 565. 1893 [Anales Univ. Quito 12: 320. 1895] = Urostachys attenuatus (Spring) Herter ex Nessel; Die Bärlappgewächse 101 (1939) = Urostachys tobarii (Sodiro) Herter ex Nessel; Die Bärlappgewächse 101 (1939) Huperzia australiana (Herter) Holub; Folia Geobot. Phytotax. 20: 70 (1985) Distribution: Borneo (Mt. Kinabalu, etc.), Macquarie Isl., New Zealand (North Isl., South Isl., Stewart Isl.), Campbell Isl., Auckland Isl., New South Wales, Victoria, Tasmania Remarks: selago agg. = Lycopodium australianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 42 (1909) = Urostachys australianus (Herter) Herter ex Nessel; Bärlappgewächse 49 (1939) = Urostachys australianus (Herter) Herter; Engl. Bot. Jb. 43, 42 (1909) Huperzia australis (Willd.) Holub; Folia Geobot. Phytotax. 20: 70 (1985) Distribution [complete]: Society Isl. Remarks: phlegmaria agg. = Lycopodium australe Willd.; Sp. pl. 5: 11 (1810) = Urostachys australis (Willd.) Herter ex Nessel; Bärlappgewächse 224 (1939) Huperzia austroecuadorica B. Øllg.; Flora of Ecuador 33: 31 (1988) Distribution [complete]: Ecuador Remarks: reflexa agg. Huperzia austrosinica Ching; Acta Botanica Yunnanica 3(3): 298-299 (1981) Distribution: Guangxi Remarks: selago agg. Huperzia badiniana B. Øllg. & Wind.; Bradea 5: 7, fig. 1A (1987) Distribution: Brazil (Minas Gerais) Remarks: saururus agg. Huperzia balansae (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution [complete]: New Caledonia Remarks: myrtifolia agg. = Lycopodium balansae Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 51 (1909) = Urostachys balansae (Herter) Herter ex Nessel; Bärlappgewächse 208 (1939) = Lycopodium balansae var. pomperyanum Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 52 (1909) = Urostachys pomperyanus (Herter) Herter; Index Lyc. 76 (1949) Huperzia bampsiana Pichi-Serm.; Bull. Jard. Bot. Nat. Belg. 55: 193 (1985) Distribution [complete]: Rwanda, Tanzania (Kilimanjaro) Remarks: ophioglossoides agg. Huperzia banayanica (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution: Philippines Remarks: phlegmaria agg. = Urostachys banayanicus Herter; Bot. Arch. 3: 17 (1923) = Lycopodium banayanicum (Herter) Copel.; Univ. Calif. Publ. Bot. 12: 384 (1931) Huperzia beccarii (Alderw.) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution: Sumatra Remarks: selago agg. = Lycopodium beccarii Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 7: 29 (1912) = Urostachys beccarii (Alderw.) Herter ex Nessel; Bärlappgewächse 44 (1939) Huperzia beiteliana Mickel; Brittonia 44(3): 312, f. 1A-E (1992) Distribution: Mexico Huperzia beitelii B. Øllg.; Amer. Fern Journal 79: 151 (1989) Distribution [complete]: Venezuela, British Guyana (Mt. Roraima) Huperzia biformis (Hook.) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution: Brazil (Organ Mts.) Remarks: myrsinites agg. = Lycopodium biforme Hook.; Icon. pl. 3: t. 228 (1839) = Urostachys biformis (Hook. ) Herter; Index Lyc. 52 (1949) Huperzia binervia (Herter) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution [complete]: Peru Remarks: brongniartii agg. = Lycopodium binervium Spring ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 48 (1909) = Urostachys binervius (Herter) Herter ex Nessel; Bärlappgewächse 110 (1939) Huperzia bolanica (Rosenstock) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution: New Guinea Remarks: phlegmaria agg. = Lycopodium bolanicum Rosenstock; Repert. Spec. Nov. Regni Veg. 12: 181 (1913) = Urostachys bolanicus (Rosenstock) Herter ex Nessel; Bärlappgewächse 234 (1939) Huperzia boliviana (Rosenst.) Rolleri & Deferrari; Notas del Museo de la Plata, Botánica 21 (100): 155 (1988) Distribution: Bolivia Remarks: brongniartii agg. = Lycopodium bolivianum Rosenst.; Repert. Spec. Nov. Regn. Veg. 21: 95 (1924) = Urostachys bolivianus (Rosenstock) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 396 (1927) Huperzia bonapartei (Nessel) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution: Pacific Remarks: unverified name, hippuris agg. = Urostachys bonapartei Nessel; Repert. Spec. Nov. Regni Veg. 36: 182, t. 172 (1934) Huperzia borneensis (Herter) comb. ined. Distribution: Borneo Remarks: unverified name, phlegmaria agg. = Urostachys borneensis Herter; Philipp. J. Sci. 22: 181 (1923) = Urostachys leonis Herter; Index Lyc. 67. 1949, nom. superfl. for Urostachys borneensis Herter Huperzia brachiata (Maxon) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution [complete]: Isla del Coco Remarks: verticillata agg. = Lycopodium brachiatum Maxon; Contr. U.S. Natl. Herb. 17: 176 (1913) = Urostachys brachiatus (Maxon) Herter ex Nessel; Die Bärlappgewächse 127 (1939) Huperzia brachystachys (Baker) Pichi-Serm.; Webbia 23: 162 (1968) Distribution [complete]: Cameroun, Bioko Isl. (Fernando Poo), Guinea, Sierra Leone, Ivory Coast, Nigeria, Equatorial Guinea Remarks: gnidioides agg. = Lycopodium brachystachys (Baker) Alston; Bol. Soc. Brot., ser. 2, 30: 19. 1956 [non brachystachyum Grev. & Hook. (1832) = Lycopodium dacrydioides Baker var. brachystachys Baker; Handb. fern-allies 18 (1887) = Urostachys brachystachys (Baker) Herter ex Nessel; Bärlappgewächse 189 (1939) Huperzia bradeorum (H. Christ) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution [complete]: Costa Rica, Guatemala, Panama, Isla Margarita, Venezuela, Ecuador, Nicaragua Remarks: wilsonii agg. = Huperzia brauseana (Herter) Rolleri & Deferrari; Notas del Museo de la Plata, Botánica 21 (100): 155 (1988) = Lycopodium bradeorum H. Christ; Repert. Spec. Nov. Regni Veg. 8: 20 (1910) = Lycopodium brauseanum Herter; Symbolae Antillarum 7(2): 165 (1912) = Urostachys bradeorus (H. Christ) Herter ex Nessel; Die Bärlappgewächse 154 (1939) = Urostachys brauseanus (Herter) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 162 (1923) Huperzia brasiliana (Herter) comb. ined. Distribution: Brazil Remarks: unverified name, brongniartii agg. = Lycopodium brasilianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 44 (1909) = Urostachys brasilianus (Herter) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 386 (1927) Huperzia brassii (Copel.) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution: Papua New Guinea Remarks: phlegmaria agg. = Lycopodium brassii (Nessel) Alston; J. Bot. 77: 290. 1939 [non Copel. (1929) = Lycopodium brassii Copel.; Journal of the Arnold Arboretum 10: 174 (1929) = Urostachys arietis Herter; Index Lyc. 51 (1949) [nom. superfl. = Urostachys brassii (Copel.) Herter; Index Lyc. 54. 1949 [non Nessel (1935]) = Urostachys brassii Nessel; Repert. Spec. Nov. Regni Veg. 39: 62, t. 190a (1935) Huperzia brevifolia (Grev. & Hook.) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution [complete]: Costa Rica, Colombia, Venezuela, Ecuador, Peru Remarks: brevifolia agg. = Lycopodium brevifolium Grev. & Hook.; Bot. Misc. 3: 104 (1832) = Urostachys brevifolius (Grev. & Hook.) Herter; Estudios Botánicos en la Región Uruguaya 20: 54 (1949) = Urostachys rufescens var. brevifolius (Grev. & Hook.) Nessel; Die Bärlappgewächse 103 (1939) Huperzia brongniartii (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution [complete]: Bolivia, Ecuador, Peru, Colombia Remarks: brongniartii agg. = Lycopodium brongniartii Spring; Bull. Acad. Roy. Sci. Bruxelles 8 (2): 515 (1841) = Lycopodium taxifolium var. brongniartii (Spring) Baker; Handbook of the Fern-Allies 16 (1887) = Urostachys brongniartii (Spring) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 388, t. 8 (1927) Huperzia bucahwangensis Ching; Acta Botanica Yunnanica 3(3): 301 (1981) Distribution: NW-Yunnan Remarks: selago agg. Huperzia buesii (Herter) B. Øllg.; Fieldiana, Bot., New Ser. 34: 42 (1994) Distribution [complete]: Peru Remarks: taxifolia agg. = Lycopodium buesii (Herter) C. Morton; Amer. Fern 1. 54: 72 (1964) = Urostachys buesii Herter; Revista Sudamer. Bot. 10: 126 (1953 [1954]) Huperzia callitrichifolia (Mett.) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution [complete]: Ecuador, Colombia, Venezuela Remarks: myrsinites agg. = Lycopodium callitrichifolium Mett.; Annales des Sciences Naturelles, Botanique, sér. 5, 3: 309 (1865) = Urostachys callitrichaefolius (Mett.) Nessel; Die Bärlappgewächse 253 (1939) Huperzia campestris (Alderw.) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution: New Guinea Remarks: selago agg. = Lycopodium campestre Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 24: 3 (1917) = Urostachys campestris (Alderw.) Herter ex Nessel; Bärlappgewächse 45 (1939) Huperzia campiana B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 109, f. 22B (1988) Distribution [complete]: Ecuador, Peru Remarks: myrsinites agg. Huperzia cancellata (Spring) Trev.; Atti. Soc. Ital. Sci. Nat. 17: 247 (1874) Distribution: India (Arunachal Pradesh), Bhutan, Tibet, Yunnan, Philippines, Japan, S-China, Taiwan, Ryukyu Isl. Remarks: cancellata agg. = Huperzia laxa (C. Presl) T. Sen & U. Sen; Fern Gazette 11(6): 420, f. 3, a-j (1978) = Huperzia laxa (C. Presl) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247. 1874 = Lycopodium cancellatum Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 27 (1849) = Lycopodium laxum C. Presl; Reliquiae Haenkeanae 1(1): 83 (1825) = Phlegmariurus cancellatus (Spring) Ching; Acta Botanica Yunnanica 4(2): 122 (1982) = Phlegmariurus cancellatus (Spring) Ching; Flora Xizangica 1: 11-12., nom illeg. superfl (1983) = Phlegmariurus cancellatus var. minor Ching (1982) = Urostachys cancellatus (Spring) Herter ex Nessel; Die Bärlappgewächse 140, f. 34 (1939) Huperzia capellae (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution [complete]: Ecuador, Peru, Venezuela, Colombia Remarks: saururus agg. = Lycopodium capellae (Herter) C.V. Morton; Amer. Fern Journal 54: 72 (1964) = Urostachys capellae Herter; Revista Sudamericana de Botánica 10: 114-115 (1954) Huperzia capillaris (Sodiro) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution [complete]: Costa Rica, S-Mexico (Chiapas), Guatemala, Panama, Colombia, Venezuela, Ecuador, Brazil Remarks: linifolia agg. = Huperzia underwoodiana (Maxon) Holub; Folia Geobot. Phytotax. 20: 77 (1985) = Lycopodium capillare Sodiro; Recens. Crypt. Vasc. Quit. 90 (1883) = Lycopodium guatemalense Maxon; Contributions from the U. S. National Herbarium 17(2): 177, t. 9a (1913) = Lycopodium sarmentosum Spring var. rubescens Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 13 (1849) = Lycopodium sarmentosum var. capillare (Sodiro) Sodiro; Cryptogamae vasculares Quitenses 573 (1893) = Lycopodium underwoodianum Maxon; Contr. U. S. Natl. Herb. 13 (1): 41. 1909 [Maxon, Contr. U. S. Natl. Herb. 17 (2): t. 9 (b) (1913]) = Urostachys capillaris (Sodiro) Herter; Estudios Botánicos en la Región Uruguaya 20: 54 (1949) = Urostachys guatemalensis (Maxon) Herter; Repert. Spec. Nov. Regn. Veg. 19: 165 (1923) = Urostachys rubescens (Spring) Herter; Estudios Botánicos en la Región Uruguaya 20: 79 (1949) = Urostachys underwoodianus (Maxon) Herter; Estudios Botánicos en la Región Uruguaya 87 (1949) Huperzia carinata (Desv. ex Poir.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1874) Distribution: India (Tamil Nadu), Nicobars, Philippines, Seychelles, peninsular Malaysia (Penang Isl., Kelantan, Trengganu, Pahan, Singapore), NE-Queensland, Taiwan, SE-Thailand, peninsular Thailand, Fiji (Viti Levu, Vanua Levu, Ovalau), Western Samoa (Aleipata Isl., Savaii, Upolu), New Caledonia, Cook Isl. (Rarotonga), American Samoa (Manua Isl., Tutuila) Remarks: carinata agg. = Huperzia struthioloides (C. Presl) Rothm.; Feddes Repert. 54: 61 (1944) = Lycopodium acrostachyum Hook. & Grev.; Icon. filic. 2: t. 181 (1831) = Lycopodium carinatum Desv. ex Poiret; Encycl. suppl. 3: 555 (1813 [1814]) = Lycopodium carinatum var. minus Tagawa (1964) = Lycopodium cumingii Loddiges (Salomon 1883: 224) [status uncertain = Lycopodium pendulum Roxb.; Calcutta J. Nat. Hist. 4. 472 (1844) = Lycopodium struthioloides C. Presl; Reliq. haenk. 1: 82 (1825) = Lycopodium urostachyum Wall. ex Ridley; J. Straits Branch Roy. Asiat. Soc. 80: 142. 1919 = Phlegmariurus carinatus (Poiret) Ching; Acta Bot. Yunnanica 4: 120 (1982) = Selaginella struthioloides (C. Presl) L. Underw.; Bull. Torrey Bot. Club 25: 132 (1898) = Urostachys carinatus (Poiret) Herter; Bot. Arch. 3: 115 (1923) = Urostachys struthioloides (C. Presl) Herter ex Nessel; Bärlappgewächse 170 (1939) Huperzia catacachiensis (Herter ex Nessel) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution [complete]: Colombia Remarks: saururus agg. = Urostachys catacachiensis Herter ex Nessel; [Bärlappgewächse 99. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 162, t. 10, f. 40 (1940) Huperzia catharinae (Christ) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution: Brazil Remarks: selago agg. = Lycopodium catharinae Christ; Schwacke, Pl. Nov. Mineiras 2: 39 (1900) = Urostachys catharinae (Christ) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 387 (1927) Huperzia cavifolia (C. Chr.) Tard.; Adansonia 10: 18 (1970) Distribution [complete]: Madagascar Remarks: gnidioides agg. = Huperzia smithiana (Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985) = Lycopodium cavifolium C. Chr.; Dansk Bot. Ark. 7: 190. t. 76, f. 6-10 (1932) = Urostachys cavifolius (C. Chr.) Herter; Index Lyc. 55 (1949) = Urostachys smithianus Nessel; Repert. Spec. Nov. Regni Veg. 36: 185, t. 174 (1934) Huperzia ceylanica (Spring) Rothm.; Feddes Repert. 54: 59 (1951) Distribution: India (Meghalaya, Tamil Nadu), Sri Lanka, Java Remarks: selago agg. Conservation status: IUCN I = Lycopodium ceylanicum Spring; Mém. Acad. Sci. Belg. 15 (1): 37. 1843, 24(2): 16 (1850) = Urostachys ceylanicus (Spring) Herter ex Nessel; Bärlappgewächse 52 (1939) Huperzia chamaeleon (Herter) B. Øllgaard; Opera Bot. 92: 169 (1987) Distribution [complete]: Costa Rica Remarks: reflexa agg. = Urostachys chamaeleon Herter; Amer. Fern. J. 48: 82 (1958) = Lycopodium chamaeleon (Herter) C. Morton; Amer. Fern J. 54: 72 (1964) Huperzia chinensis (H. Christ) Ching; Acta Botanica Yunnanica 3(3): 304 (1981) Distribution: E-Siberia, Japan, Korea, China, Amur, Ussuri Remarks: selago agg. = Huperzia chinensis (Christ) Ching; Acta Bot. Yunnanica 3: 304 (1981) = Huperzia chinensis (H. Christ) Ching; Flora Xizangica 1: 9 (1983) = Huperzia selago (L.) Bernh. ex Schrank & C. Mart. ssp. chinensis (Christ) A. & D. Löve = Lycopodium chinense Christ ex Baroni & Christ; Nuovo Giorn. Bot. Ital., n. s. 4 (1): 1011 t. 3, f. 4 (1897) = Lycopodium selago ssp. chinense (Christ) Hulten = Lycopodium selago var. chinense (Christ) Ohwi; Flora of Japan Pterid. 24 (1965) [nom. illeg., non Lycopodium selago ssp. chinense (H. Christ) Hultén = Urostachys chinense (Christ) Herter ex Nessel; Die Bärlappgewächse 27, Abb. 4, f. 2 (1939) Huperzia chiricana (Maxon) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution [complete]: Panama (Volcan Chiriqui), Costa Rica (Volcan Turrialba) Remarks: saururus agg. = Lycopodium chiricanum Maxon; Contr. U.S. Natl. Herb. 17: 176 (1913) = Urostachys chiricanus (Maxon) Herter ex Nessel; Die Bärlappgewächse 45 (1939) Huperzia chishuiensis X.Y. Wang & P.S. Wang; Bulletin of Botanical Research 16(4): 417, f. 1-3 (1996) Distribution: China Huperzia christii (Silveira) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution: Brazil (Minas Gerais) Remarks: reflexa agg. = Lycopodium christii Silveira; Bol. Comm. Geogr. Geol. Minas Geraes 2, 5: 117, t. 1 (1898) = Lycopodium christianum Kümm.; Magyar Bot. Lapok 26: 99 (1927 [1928]) nom. confus. et superfl. = Urostachys christii (Silveira) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 379 (1927) Huperzia cipoensis (Damazio) comb. ined. Distribution: Brazil (Minas Gerais) Remarks: unverified name, heterocarpon agg. = Lycopodium cipoense Damazio; Bull. Soc. Bot. Genbve, ser. 2, 7: 119, f. 1-8 (1915) = Urostachys cipoensis (Damazio) Nessel; Bärlappgewächse 169 (1939) Huperzia clarkiana (Nessel) comb. ined. Distribution: New Guinea Remarks: unverified name, hippuris agg. = Urostachys clarkianus Nessel; Repert. Spec. Nov. Regni Veg. 36: 183, t. 173 Huperzia clausenii (Herter ex Nessel) comb. ined. Distribution: New Guinea Remarks: unverified name, cancellata agg. = Urostachys clausenii Herter ex Nessel; [Bärlappgewächse 135. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 163, t. 10, f. 47 (1940) Huperzia colanensis B. Øllg.; Fieldiana: Botany, New Series 34: 31 (1994) Distribution [complete]: Peru Huperzia columnaris B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 45, f. 5A (1988) Distribution [complete]: Ecuador Remarks: saururus agg. Huperzia comans (Herter ex Nessel) B. Øllg. & P.G. Windisch; Bradea 5: 8 (1987) Distribution: Brazil Remarks: verticillata agg. = Lycopodium comans Christ; Schwacke, Pl. Nov. Mineiras 2. 40. 1900 [non Hook. f (1844) = Urostachys comans Herter ex Nessel; Arch. Bot. Sao Paulo 1: 400. 1927 [nom. nov. for Lycopodium comans Christ [non Hook. f (1844]) = Urostachys pollucis Herter; Index Lyc. 76 (1949 [nom. superfl. for Urostachys comans Nessel) Huperzia compacta (Hook.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1874) Distribution [complete]: Ecuador Remarks: brevifolia agg. = Huperzia jamesonii (Baker) Holub; Folia Geobot. Phytotax. 20: 73 (1985) = Lycopodium compactum Hook.; Icones Plantarum 3: t. 244 (1839) = Lycopodium jamesonii Baker; Handb. fern-allies 9 (1887) = Urostachys compactus (Hook.) Nessel; Die Bärlappgewächse 102 (1939) = Urostachys jamesonii (Baker) Nessel; Die Bärlappgewächse 100 (1939) Huperzia copelandiana (Chou & Bartlett) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution: Sumatra Remarks: macgregorii agg. = Lycopodium petiolatum Copel.; Univ. Calif. Publ. Bot. 14: 377. 1929 [non Herter (1909) = Lycopodium copelandianum R. C.-Y. Chou & Bartlett; Bull. Torrey Bot. Club 74: 369 (1947) Huperzia copelandii (Herter) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution: Solomons Remarks: phlegmaria agg. = Lycopodium filicaulon Copel.; Philipp. J. Sci. 60: 100, t. 2. 1936 [non Lycopodium filicaule Hook. f (1844) = Urostachys copelandii Herter; Estudios Botánicos en la Región Uruguaya 20: 57 (1949) Huperzia coralium (Spring) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution: peninsular Malaysia (Perak), Java Remarks: phlegmaria agg. = Lycopodium coralium Spring; Miquel, Pl. jungh. 3: 273 (1854) = Urostachys coralium (Spring) Herter ex Nessel; Bärlappgewächse 234 (1939) Huperzia coriacea (Rolleri) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988) Distribution [complete]: Colombia Remarks: reflexa agg. = Lycopodium coriaceum Rolleri; Revista Mus. La Plata (N.S.) Bot. 13 (71): 85, t. 5 (B, D) (1981) Huperzia costaricensis (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution [complete]: Costa Rica Remarks: taxifolia agg. = Urostachys costaricensis Herter; Amer. Fern J. 48: 83 (1958) = Lycopodium costaricense (Herter) C. Morton; Amer. Fern 54: 72 (1964) Huperzia crassa (Humb. & Bonpl. ex Willd.) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 60 (1944) Distribution [complete]: Costa Rica, Mexico (Oaxaca), Guatemala, Panama, Colombia, Venezuela, Ecuador, Bolivia, Hispaniola, Peru Remarks: saururus agg. = Huperzia erythraea (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1874) = Huperzia pilgeriana (Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 75 (1985) = Huperzia springii (Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 77 (1985) = Lycopodium bonae-voluntatis (Herter) C. V. Morton; Amer. Fern Journal 54: 72 (1964) = Lycopodium crassum Humb. & Bonpl. ex Willd.; Sp. Pl. 5: 50 (1810) = Lycopodium erythraeum Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 7 (1849) = Lycopodium nesselianum Duek & Lellinger; Amer. Fern Journal 68(4): 120 (1978) = Lycopodium saururus var. crassum (Humb. & Bonpl. ex Willd.) Sodiro; Recensio Cryptogamarum Vascularium Provinciae Quitensis 90 (1883) = Urostachys bonae-voluntatis Herter; Revista Sudamericana de Botánica 10: 112. [1953] (1954) = Urostachys crassus (Humb. & Bonpl. ex Willd.) Herter ex Nessel; Die Bärlappgewächse 75 (1939) = Urostachys crassus var. herteri Nessel; Repert. Spec. Nov. Regn. Veg. 36(12-15): 180, t. 171 (1934) = Urostachys erythraeus (Spring) Herter ex Nessel; Die Bärlappgewächse 74 (1939) = Urostachys kupperi Nessel (1934) = Urostachys orionis Herter; Index Lyc. 73. 1949 [nom. superfl. = Urostachys pilgerianus Herter ex Nessel; Revista Sudamericana de Botánica 6(5-6): 161, t. 9, f. 32 (1940) = Urostachys springii (Kl. & Karsten) Herter ex Nessel; Revista Sudamericana de Botánica 6(5-6): 161, t. 9, f. 31 (1940) var. gelida B. Øllg.; Flora of Ecuador 33: 56 (1988) Distribution [complete]: Ecuador var. manus-diaboli B. Øllg.; Flora of Ecuador 33: 56 (1988) Distribution [complete]: Ecuador Huperzia crebra (Alderw.) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution: New Guinea Remarks: macgregorii agg. = Lycopodium crebrum Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 24: 4 (1917 ("crebre")) = Urostachys creber (Alderw.) Herter ex Nessel; Bärlappgewächse 181 (1939) Huperzia crispata (Ching in H.S. Kung) Ching; Acta Botanica Yunnanica 3(3): 293 (1981) Distribution: Sichuan Remarks: selago agg. = Lycopodium crispatum Ching ex Kung; Acta Phytotaxonomica Sinica 18(2): 236, f. 1: 3 (1980) Huperzia crucis-australis (Herter) comb. ined. Distribution: South America Remarks: unverified name, saururus agg. = Lycopodium crucis-australis (Herter) C. Morton; Amer. Fern J. 54: 72 (1964) = Urostachys crucis-australis Herter; Revista Sudamer. Bot. 10: 119 (1953 [1954]) Huperzia cruenta (Spring) Rothm.; Feddes Repert. 54: 59 (1944) Distribution [complete]: Colombia, Venezuela Remarks: saururus agg. = Lycopodium cruentum Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 34 (1849) = Urostachys cruentus (Spring) Herter ex Nessel; Bärlappgewächse 27 (1939) Huperzia cryptomeriana (Maxim.) R.D. Dixit; Journal of the Bombay Natural History Society 77(3): 541 (1981) Distribution: India (Arunachal Pradesh), China, Japan, Taiwan, Korea, Yakushima Isl. Remarks: hamiltonii agg. = Lycopodium cryptomerianum Maxim.; Bulletin de l'Academie Imperiale des Sciences de St-Petersbourg 15: 231 (1870) = Phlegmariurus cryptomerianus (Maxim.) Ching; Acta Phytotaxonomica Sinica 37(1): 51 (1999) = Urostachys cryptomerianus (Maxim.) Herter ex Nessel; Die Bärlappgewächse 70 (1939) Huperzia cubana (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985) Distribution [complete]: Cuba Remarks: taxifolia agg. = Lycopodium cubanum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 50 (1909) = Urostachys cubanus (Herter) Herter; Repert. Spec. Nov. Regni Veg. 19: 165 (1923) Huperzia cuernavacensis (Underw. & F. E. Lloyd) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Mexico (Chiapas, Sinaloa, Jalisco, Michoacan, Mexico State, Morelos, Oaxaca), Guatemala Remarks: taxifolia agg. = Lycopodium cuernavacensis Underw. & F. E. Lloyd; Bull. Torrey Club 33: 110 (1906) = Urostachys cuernavacensis (Underw. & F. E. Lloyd) Herter ex Nessel; Die Bärlappgewächse 92 (1939) Huperzia cumingii (Nessel) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Ecuador, Colombia Remarks: saururus agg. = Lycopodium cumingii (Nessel) Rolleri; Revista del Museo de La Plata, Nueva Serie, Botanica 13(71): 92 (1980) = Urostachys cumingii Nessel; Repert. Spec. Nov. Regn. Veg. 36(12-15): 178, t. 170 (1934) Huperzia cuneifolia (Hieron.) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Costa Rica, Panama, Colombia, E-Venezuela, N-Peru Remarks: myrsinites agg. = Lycopodium aqualupianum var. obtusifolium Spring; Mémoires de l'Académie Royales des Sciences, Lettres & Beaux Arts de Belgique 24: 31 (1849) = Lycopodium cuneifolium Hieron.; Bot. Jahrb. Syst. 34: 572 (1905) = Urostachys cuneifolius (Hieron.) Herter ex Nessel; Die Bärlappgewächse 252, t. 62, f. 14 (1939) Huperzia curiosa (Herter) Tard.; Adansonia 10: 18 (1970) Distribution [complete]: Madagascar Remarks: squarrosa agg. = Urostachys curiosus Herter; Notul. Syst. (Paris) 15: 355 (1958) Huperzia curvifolia (Kunze) Holub; Folia Geobot. Phytotax. 20(1): 72 (1985) Distribution [complete]: Peru, Ecuador Remarks: verticillata agg. = Lycopodium curvifolium Kunze; Linnaea 9: 5 (1835) = Urostachys curvifolius (Kunze) Nessel; Die Bärlappgewächse 129 (1939) = Lycopodium tenue var. tenuissimum Spring; Mémoires de l'Académie Royales des Sciences, Lettres & Beaux Arts de Belgique 24: 21 (1849) var. parvifolia (Wercklé ex Nessel) B. Øllg.; Novon 3(1): 71 (1993) Distribution [complete]: Costa Rica, Honduras, Panama, Colombia = Lycopodium verticillatum var. parvifolium (Wercklé ex Nessel) Lellinger; Proc. Biol. Soc. Wash. 89(61): 719 (1977) = Urostachys verticillatus var. parvifolius Wercklé ex Nessel; Revista Sudamer. Bot. 6: 163 (1940) Huperzia dacrydioides (Baker) Pichi-Serm.; Webbia 23: 162 (1968) Distribution: Transvaal, Natal, Cape Prov., Zimbabwe, Malawi, Mozambique, Comores, Uganda, Kenya, Tanzania, Sudan, Zaire, Burundi, Ethiopia Remarks: gnidioides agg. = Lycopodium dacrydioides Baker; The Fern Allies 17 (1887) = Urostachys dacrydioides (Bak.) Herter ex Nessel; Bärlappgewächse 188 (1939) Huperzia dalhousiana (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247 (1874) Distribution [complete]: NE-Queensland, peninsular Malaysia (Penang Isl., Pahan, Melaka), Borneo Remarks: dalhousiana agg. = Huperzia caudifolia (Alderw.) Holub; Folia Geobot. Phytotax. 20: 71 (1985) = Huperzia clarae (Bailey) Holub; Folia Geobot. Phytotax. 20: 71 (1985) = Lycopodium caudifolium Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 1: 14 (1911) = Lycopodium clarae Bailey; Queensl. Dept. Agric., Brisbane, Bull. 21 (Bot. Bull. 7): 69 (1893) = Lycopodium dalhousianum Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 25 (1849]) = Urostachys caudifolius (Alderw.) Herter ex Nessel; Bärlappgewächse 185 (1939) = Urostachys clarae (Bailey) Herter ex Nessel; Bärlappgewächse 185 (1939) = Urostachys dalhousianus (Spring) Herter ex Nessel; Bärlappgewächse 185 (1939) Huperzia darwiniana (Herter ex Nessel) B. Øllg.; Fieldiana: Botany, New Series 34: 33 (1994) Distribution [complete]: Peru, Bolivia Remarks: saururus agg. = Urostachys darwinianus Herter ex Nessel; Revista Sudamericana de Botánica 6(5-6): 161 (1940) Huperzia delavayi (H. Christ & Herter in Herter) Ching; Acta Botanica Yunnanica 3(3): 303 (1981) Distribution: Yunnan Remarks: selago agg. = Lycopodium delavayi H. Christ & Herter in Herter; Bot. Jahrb. Syst. 43(1, Beibl. 98): 41 (1909) = Urostachys delavayi (H. Christ & Herter in Herter) Herter ex Nessel; Die Bärlappgewächse 31, Abb. 4, f. 9 (1939) Huperzia delbrueckii (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution: Philippines (Mindanao) Remarks: nummulariifolia agg. = Urostachys delbrueckii Herter; Bot. Arch. 3. 19 (1923) = Lycopodium delbrueckii (Herter) Copel.; Univ. Calif. Publ. Bot. 12: 384 (1931) Huperzia deminuens (Herter) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution: Brazil (Minas Gerais) Remarks: brongniartii agg. Conservation status: IUCN R = Lycopodium deminuens Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 44 (1909) = Urostachys deminuens (Herter) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 384 (1927) Huperzia dentata (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Azores, Madeira Remarks: reflexa agg. = Huperzia selago ssp. dentata (Herter) Valentine; Feddes Repertorium 69: 44 (1964) = Huperzia selago ssp. dentata Tutin; Flora Europaea 1: 3 (1964) = Lycopodium dentatum Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 41 (1909) = Urostachys dentatus (Herter) Herter ex Nessel; Bärlappgewächse 41 (1939) Huperzia desvauxii Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution: Distribution unknown Huperzia dianae (Herter) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution [complete]: Colombia Remarks: brevifolia agg. = Urostachys dianae Herter; Revista Sudamer. Bot. 10: 116 (1953 [1954]) = Lycopodium dianae (Herter) C. Morton; Amer. Fern J. 54: 72 (1964) Huperzia dichaeoides (Maxon) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Costa Rica, Belize, Guatemala, Nicaragua, Panama, E-Colombia, Ecuador Remarks: aqualupiana agg. = Lycopodium dichaeoides Maxon; Proc. Biol. Soc. Wash. 18: 231 (1905) = Urostachys dichaeoides (Maxon) Herter ex Nessel; Die Bärlappgewächse 252 (1939) Huperzia dichotoma (Jacq.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution: Florida (Collier Co., Fakahatchee Strand), Costa Rica, Mexico (Chiapas, San Luis Potosi, Veracruz, Puebla, Oaxaca), Belize, Guatemala, Honduras, El Salvador, Nicaragua, Panama, Galapagos, Puerto Rico, Cuba, Jamaica, Hispaniola, Virgin Isl. (Virgin Gorda), Mexico, to, Brazil, Guadeloupe, Colombia, Ecuador, St. Eustatius, St. Kitts, Dominica, Martinique, St. Vincent, Grenada, British Guyana, Surinam, French Guiana, Bolivia Remarks: dichotoma agg. Trivial names: Hanging Club-Moss = Huperzia dichotoma (Jacq.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) = Huperzia graminea (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) = Huperzia lindeneri (Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985) = Huperzia schlechtendalii (Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985) = Lycopodium barbatum H. Christ; Bulletin de l'Herbier Boissier, sér. 2, 5(3): 254 (1905) = Lycopodium chamaepeuce Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 50 (1909) = Lycopodium dichotomum Jacq.; Enum. Stirp. Vindob. 3 (1762) = Lycopodium gramineum Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 19 (1849) = Phlegmariurus dichotomus (Jacq.) W.H. Wagner & Beitel; Novon 3: 305 (1993) = Plananthus dichotomus (Jacq.) P. Beauv.; Prod. aethéogam. 111 (1805) = Urostachys barbatus (H. Christ) Herter ex Nessel; Die Bärlappgewächse 30 (1939) = Urostachys chamaepeuce (Herter) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 164 (1923) = Urostachys dichotomus (Jacq.) Herter; Beihefte zum Botanischen Centralblatt 39(2): 249 (1922) = Urostachys gramineus (Spring) Herter; Estudios Botánicos en la Región Uruguaya 20: 63 (1949) = Urostachys lindeneri Herter ex Nessel; Revista Sudamericana de Botánica 6(5-6): 164, t. 11, f. 50 (1940) = Urostachys schlechtendalii Nessel; Revista Sudamericana de Botánica 6(5-6): 164, t. 11, f. 51 (1940) Huperzia dielsii (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution: New Guinea Remarks: phyllantha agg. = Lycopodium dielsii Herter; Bot. Jahrb. Syst. 54: 229 (1916) = Urostachys dielsii (Herter) Herter ex Nessel; Bärlappgewächse 213 (1939) Huperzia dura Pichi-Serm.; Webbia 27: 392, fig. 1 (1972 [1973]) Distribution: Tanzania Remarks: gnidioides agg. Huperzia echinata (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247 (1874) Distribution [complete]: Colombia Remarks: taxifolia agg. = Lycopodium echinatum Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 24 (1849) = Urostachys echinatus (Spring) Herter ex Nessel; Bärlappgewächse 178 (1939) Huperzia ellenbeckii (Nessel) Pichi-Serm.; Webbia 23: 162. 1968 Distribution: Ethiopia Remarks: ophioglossoides agg. = Urostachys ellenbeckii Nessel; Repert. Spec. Nov. Regni Veg. 36: 188, t. 175 (1934) Huperzia elmeri (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution: Philippines (Luzon) Remarks: phlegmaria agg. = Urostachys elmeri Herter; Bot. Arch. 3: 16 (1923) Huperzia emeiensis (Ching & H.S. Kung) Ching & H.S. Kung; Acta Botanica Yunnanica 3(3): 299-300 (1981) Distribution: Sichuan Remarks: selago agg. = Lycopodium emeiense Ching & Kung ex Kung; Acta Phytotax. Sin. 18 (2): 235, f. 2 (1980) Huperzia engleri (Herter) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution [complete]: Peru Remarks: brevifolia agg. = Lycopodium engleri Hieron. & Herter ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 45 (1909) = Urostachys engleri (Herter) Nessel; Bärlappgewächse 94 (1939) Huperzia eremorum (Rolleri) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Colombia Remarks: brevifolia agg. Conservation status: IUCN R = Lycopodium eremorum Rolleri; Hickenia 1 (26): 141, f. 1 (1978) Huperzia ericifolia (C. Presl) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Ecuador, Peru, Bolivia Remarks: aqualupiana agg. = Lycopodium ericifolium C. Presl; Reliq. haenk. 1: 77 (1825) = Urostachys phlegmaria var. ericaefolius (C. Presl) Nessel; Die Bärlappgewächse 217 (1939) Huperzia erosa Beitel & W.H. Wagner Distribution [complete]: Hawaii (Kauai, Oahu, Lanai, Maui, Hawaii Isl.) Huperzia erubescens (Brackenr.) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.) Remarks: selago agg. = Lycopodium erubescens Brackenr.; U. S. Expl. exped., Filic. 16: 320, atlas t. 45, f. 1 (1854) = Urostachys erubescens (Brack.) Herter ex Nessel; Bärlappgewächse 61 (1939) Huperzia erythrocaulon (Fée) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution: Brazil, ?Venezuela Remarks: myrsinites agg. = Lycopodium erythrocaulon Fée; Crypt. vasc. Brésil 2: 95, t. 106, f. 2 (1872-73) = Urostachys erythrocaulon (Fée) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 421 (1927) Huperzia espinosana B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 47, f. 7A (1988) ( esp) Distribution [complete]: Ecuador Remarks: saururus agg. Huperzia everettii (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution: S-Sulawesi Remarks: selago agg. = Lycopodium everettii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909) = Urostachys everettii (Herter) Herter ex Nessel; Bärlappgewächse 66 (1939) Huperzia eversa (Poir.) B. Øllgaard; Harling & L. Andersson, Fl. Ecuador 33: 28 (1988) Distribution [complete]: Ecuador, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia Remarks: reflexa agg. = Huperzia ecuadorica (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985) = Lycopodium ecuadoricum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 48 (1909) = Lycopodium eversum Poir.; Lam., Encycl. Suppl. 3: 556 (1814) = Lycopodium polycarpum (Sodiro) Underw. & F. E. Lloyd; Bulletin of the Torrey Botanical Club 33: 105 (1906) [nom. illeg., non Lycopodium polycarpos Kunze (1834) = Lycopodium reflexum Lam. var. polycarpum Sodiro; Recensio Cryptogamarum Vascularium Provinciae Quitensis 90 (1883) = Lycopodium reflexum Willd.; Sp. Pl. Editio quarta ed. 4 5: 52 (1810) [non Lam. (1789) and (1795) [nom. illeg. = Lycopodium rolandii-principis Herter ex Bonap. = Urostachys dingesianus Nessel; Repert. Spec. Nov. Regn. Veg. 39(8): 69, t. 193b (1935) = Urostachys ecuadoricus (Herter) Herter; Estudios Botánicos en la Región Uruguaya 20: 59 (1949) = Urostachys rolandii-principis Herter ex Nessel; Bärlappgewächse 11 7 (1939) Huperzia fargesii (Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: China, Taiwan, Japan Remarks: cancellata agg. = Lycopodium christensenianum Christ & Herter ex Herter; Hedwigia 49: 89 (1909) = Lycopodium fargesii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 48 (1909) = Lycopodium fargesii var. gracile Tagawa; Acta Phytotaxonomica & Geobotanica 14(3): 92 (1951) = Lycopodium fauriei Rosenst.; Hedwigia 56(5): 348 (1915) = Lycopodium tereticaule Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 411. 1911 [non Poiret (1814) = Lycopodium quasiprimaevum Koidz.; Acta Phytotax. Geobot. 1: 32 (1932) = Phlegmariurus fargesii (Herter) Ching; Acta Botanica Yunnanica 4(2): 120-121 (1982) = Phlegmariurus fargesii (Herter) Ching; Flora Xizangica 1: 12 (1983) = Urostachys christensenianus (H. Christ & Herter) Herter ex Nessel; Die Bärlappgewächse 139 (1939) = Urostachys fargesii (Herter) Herter ex Nessel; Die Bärlappgewächse 137 (1939) = Urostachys fauriei (Rosenst.) Herter; Estudios Botánicos en la Región Uruguaya 20: 61 (1949) = Urostachys tereticaulis (Hayata) Herter ex Nessel; Die Bärlappgewächse 137, f. 32 (1939) Huperzia feejeensis (Luerss.) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution: Fiji Remarks: phlegmaria agg. = Lycopodium phlegmarium var. feejeense Luerss.; Fil. graeff. 1: 270 (1871 ("feejeensis")) = Urostachys feejeensis (Luerss.) Herter; Notul. Syst. (Paris) 15: 355. 1958 Huperzia filiformis (Sw.) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Costa Rica, Colombia, Guatemala, Nicaragua, Panama, Venezuela, Ecuador, ?Peru, Bolivia, Guatemala, Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.), NE-Queensland Remarks: verticillata agg. Conservation status: IUCN V, I (Hawaii) = Huperzia polytrichoides (Kaulf.) Trevis.; Die Bärlappgewächse 129 (1939) = Huperzia tuerckheimii (Maxon) Holub; Folia Geobot. Phytotax. 20: 77 (1985) = Huperzia verticillata (L. f.) Trevis. var. filiformis (Sw.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) = Lycopodium acerosum Sw. var. filiforme (Sw.) Spring; Flora Brasiliensis 1(2): 111 (1840) = Lycopodium filiforme Sw.; J. Bot. (Schrader) 1800(2): 114 (1801) [Syn. Fil. 174, 398, t. 4, f. 3. 1806] = Lycopodium polytrichoides Kaulf.; Enumeratio Filicum 6 (1824) = Lycopodium tortile H. Christ; Bulletin de la Société Botanique de Genève 1(5): 235 (1909) = Lycopodium tuerckheimii Maxon; Contributions from the U. S. National Herbarium 13(1): 23, t. 1 (1909) = Lycopodium verticillatum var. filiforme (Sw.) Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 12: 111 (1840) = Phlegmariurus filiformis (Sw.) W. H. Wagner = Urostachys filiformis (Sw.) Herter; Estudios Botánicos en la Región Uruguaya 20: 61 (1949) = Urostachys polytrichoides (Kaulf.) Herter ex Nessel; Die Bärlappgewächse 129 (1939) = Urostachys tuerckheimii (Maxon) Herter; Repert. Spec. Nov. Regni Veg. 19: 163 (1923) Huperzia firma (Mett.) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Colombia, Brazil, Ecuador Remarks: reflexa agg. = Huperzia flexibilis (Fée) B. Øllg.; Opera Bot. 92: 169 (1987) = Lycopodium firmum Mett.; Annales des Sciences Naturelles, Botanique, sér. 5, 3: 307 (1865) = Lycopodium flexibile Fée; Crypt. vasc. Brésil 2: 94p t. 105, f. 3 (1872-73) = Urostachys firmus (Mett.) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 391 (1927) = Urostachys flexibilis (Fée) Herter; Index Lyc. 61 (1949) Huperzia flagellacea (Kuhn) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution: New Guinea Remarks: varia agg. = Lycopodium flagellaceum Kuhn; Forschungsr. Gazelle 4: 15 (1889) = Urostachys flagellaceus (Kuhn) Herter ex Nessel; Bärlappgewächse 237 (1939) Huperzia foliacea (Maxon) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Costa Rica, Panama Remarks: brongniartii agg. = Lycopodium foliaceum Maxon; Smithsonian Misc. Collect. 56(29): 1 (1912) = Urostachys foliaceus (Maxon) Herter ex Nessel; Die Bärlappgewächse 68 (1939) Huperzia foliosa (Copel.) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Fiji (Viti Levu, Vanua Levu, Ovalau, Taveuni) Remarks: phlegmaria agg. = Lycopodium foliosum Copel.; Bernice P. Bishop Mus. Bull. 59: 7, t. 1 (1929) = Urostachys foliosus (Copel.) Herter; Index Lyc. 61 (1949) Huperzia fontinaloides (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution [complete]: Brazil (Santa Catarina), Colombia Remarks: quadrifariata agg. = Lycopodium serpyllifolium Fée; Crypt. vasc. Brésil 1: 222, t. 73, f. 3 (1869) = Urostachys serpyllifolius (Fée) Herter; Index Lyc. 81 (1949) = Huperzia gehrtii (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985) = Urostachys gehrtii Nessel; Repert. Spec. Nov. Regni Veg. 36: 189, t. 177 (1934) = Lycopodium fontinaloides Spring; Flora Brasiliensis 1(2): 112, pl. 5, f. 2 (1840) = Urostachys fontinaloides (Spring) Herter ex Nessel; Archivos de Botânica do São Paulo 1: 401 (1927) Huperzia fordii (Bak.) Dixit; Census Indian Pteridophytes 7 (1984) [comb. nov. Distribution: India (Eastern Himalayas), China, Taiwan, Japan, Ryukyu Isl. Remarks: hamiltonii agg. = Huperzia fordii (Baker) Holub; Folia Geobot. Phytotax. 20(1): 72 (1985) = Lycopodium fordii Bak.; Handbook of the Fern-Allies 17 (1887) = Lycopodium poissonii Herter; Bot. Jahrb. Syst. 43: Beibl. 9851 (1909) = Lycopodium subdistichum Makino; Bot. Mag. (Tokyo) 12: 37 (1898) = Phlegmariurus fordii (Baker) Ching; Acta Bot. Yunnanica 4: 126. 1982 [Acta Phytotax. Sin. 20 (4): 447 (1982]) = Urostachys fordii (Baker) Herter ex Nessel; Bärlappgewächse 69 (1939) = Urostachys poissonii (Herter) Herter ex Nessel; Bärlappgewächse 196 (1939) = Urostachys subdistichus (Makino) Herter ex Nessel; Bärlappgewächse 196 (1939) Huperzia formosana Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution: Taiwan Huperzia friburgensis (Nessel) B. Øllg. & Wind.; Bradea 5: 10 (1987) Distribution: Brazil Remarks: reflexa agg. = Urostachys friburgensis Herter ex Nessel; Arch. Bot. Sao Paulo 1: 391 (1927) = Lycopodium friburgense (Nessel) Rolleri; Revista Mus. La Plata, N. S. 13 (Bot. 78): 191 (1984) Huperzia fuegiana (Roiv.) Holub; Folia Geobot. Phytotax. 20: 72 (1985) Distribution [complete]: Argentina (Santa Cruz Prov.), Tierra del Fuego, Falkland Isl. Remarks: selago agg. Conservation status: IUCN R = Lycopodium fuegianum Roiv.; Ann. Bot. Soc. Zool.-Bot. "Vanamo" 6 (8): 13-15, f. 4-5 (1936) = Urostachys fuegianus (Roiv.) Herter; Index Lyc. 62 (1949) = Urostachys selago var. hessei Herter ex Nessel (1939) [nom. illeg. = Urostachys selago var. aschersonianus Herter ex Nessel (1939) [nom. illeg. Huperzia funiformis (Cham. ex Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17. 248 (1874) Distribution [complete]: Puerto Rico, Cuba, Hispaniola, Jamaica, Costa Rica, Panama, S-Mexico, Colombia, Venezuela, Ecuador, Peru, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent, British Guyana, Trinidad Remarks: taxifolia agg. = Lycopodium funiforme Cham. ex Spring; Bull. Acad. Roy. Sci. Belg. 8: 516. 1841 [Brongniart, Hist. Vég. fossil. 2: 10 (1837)] [nom. inval. = Urostachys funiformis (Cham. ex Spring) Herter; Symbolae Antillarum 9: 387 (1925) Huperzia gagnepainiana (Herter) Tard.; Adansonia 10: 18 (1970) Distribution [complete]: Madagascar Remarks: pecten agg. = Urostachys gagnepainianus Herter; Revista Sudamer. Bot. 8: 23 (1949) Huperzia galapagensis (Hamann) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution [complete]: Galapagos Remarks: verticillata agg. Conservation status: IUCN R = Lycopodium setaceum ssp. galapagense O. Hamann; Botaniska Notiser 127(2): 253 (1974) Huperzia gedeana (Alderw.) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: Java Remarks: selago agg. = Lycopodium gedeanum Alderw.; Malayan ferns & fern allies, Suppl. 1, corr.: 65. 1917, [Bull. Jard. Bot. Buitenzorg ser. 2, 28: 43, t. 8 (1918]) = Urostachys gedeanus (Alderw.) Herter ex Nessel; Bärlappgewächse 45 (1939) Huperzia gigantea (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: Distribution unknown Remarks: unverified name, verticillata agg. = Urostachys verticillatus (L. f.) Herter var. giganteus Herter; Bot. Arch. 3: 13 (1923) = Urostachys giganteus Herter ex Nessel; Bärlappgewächse 124 (1939) Huperzia glauca (Cesati) comb. ined. Distribution: Sarawak Remarks: unverified name, dalhousiana agg. = Lycopodium glaucum Cesati; Atti Accad. Sc. Fis. 7: 35 (1876) = Urostachys glaucus (Cesati) Herter; Revista Sudamer. Bot. 8: 86 (1949) Huperzia gnidioides (L. f.) Trevis.; Atti della Società Crittogamologica Italiana 17: 247 (1875) Distribution: Transvaal, Natal, Cape Prov., Swaziland, Lesotho, Madagascar, Zimbabwe, Malawi, Mozambique, Comores, Tanzania, Mauritius, Réunion, Zaire, Rwanda, Burundi, Seychelles Remarks: gnidioides agg. = Huperzia helmii (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985) = Huperzia stricta (Baker) Tardieu; Adansonia ser. 2 10: 20 (1970) = Lepidotis funiculosa (Lam.) P. Beauv.; Prodr. aethéogam. 108 (1805) = Lycopodium ambiguum Schrad.; Götting. gel. Anz. 1818: 920. 1818 = Lycopodium flagelliforme Schrad.; Götting. Gel. Anz. 1818: 920. 1818 = Lycopodium funiculosum Lam.; Encyclopédie Méthodique, Botanique 3: 689 (1789) = Lycopodium gnidioides L. f.; Suppl. pl. 448 (1782) = Lycopodium gnidioides var. strictum (Bak.) C. Chr. (1932) = Lycopodium pinifolium Kaulf.; Enumeratio Filicum 7 (1824) [non Bl (1828) = Lycopodium strictum Bak.; Journal of Botany, British and Foreign 20: 271. (n.s., vol. 11) (1882) = Plananthus gnidioides (L. f.) Beauv.; Prod. aethéogam. 110 (1805) = Urostachys gnidioides (L. f.) Herter ex Nessel; Die Bärlappgewächse 187 (1939) = Urostachys helmii Nessel; Repert. Spec. Nov. Regni Veg. 36: 186p t. 174 (1934) = Urostachys strictus (Baker) Herter ex Nessel; Die Bärlappgewächse 191, t. 50 (1939) = ?Urostachys funiculus Herter; Revista Sudamer. Bot. 8: 22 (1949) Huperzia goebelii (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: Sumatra, Borneo Remarks: phlegmaria agg. = Urostachys goebelii Nessel; Repert. Spec. Nov. Regni Veg. 36: 188, t. 175 (1934) = Huperzia heroldii (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985) = Urostachys heroldii Nessel; Revista Sudamer. Bot. 6: 167, t. 14, f. 69 (1940) Huperzia goliathensis (Alderw.) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: New Guinea Remarks: selago agg. = Lycopodium goliathense Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 7: 29 (1912) = Urostachys goliathensis (Alderw.) Herter ex Nessel; Bärlappgewächse 44 (1939) Huperzia goudotii (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution [complete]: Colombia, Costa Rica, ?Peru Remarks: brevifolia agg. = Lycopodium goudotii Herter; Bot. Jahrb. Syst. 43. Beibl. 98: 47 (1909) = Urostachys goudotii (Herter) Herter ex Nessel; Bärlappgewächse 106 (1939) Huperzia guangdongensis (Ching) Holub; Folia Geobot. Phytotax. 20(1): 73 (1985) Distribution [complete]: Guangdong Remarks: phlegmaria agg. = Phlegmariurus guangdongensis Ching; Acta Botanica Yunnanica 4(2): 123 (1982) Huperzia gunturensis (Alderw.) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: Java Remarks: carinata agg. = Lycopodium gunturense Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 1: 14 (1911) = Urostachys gunturensis (Alderw.) Herter ex Nessel; Bärlappgewächse 183 (1939) Huperzia haeckelii (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution [complete]: Society Isl. (Tahiti) Remarks: verticillata agg. = Lycopodium haeckelii Herter; Repert. Spec. Nov. Regni Veg. 5: 22 (1908) = Urostachys haeckelii (Herter) Herter ex Nessel; Bärlappgewächse 138 (1939) Huperzia haitensis (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: Haiti Remarks: taxifolia agg. = Urostachys haitensis Herter; Repert. Spec. Nov. Regni Veg. 19: 161 (1923) Huperzia haleakalae (Brackenridge) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution [complete]: +Hawaii (+Maui), Alaska, Canada (Yukon, Alberta, British Columbia), USA (Rocky Mts. south, to, Washington, Montana, Colorado, Wyoming) Remarks: selago agg. = Lycopodium haleakalae Brackenridge; Wilkes, U.S. Expl. Exped. 16: 321 (1854) = Lycopodium selago var. haleakalae (Brackenridge) Warburg = Urostachys haleakalae (Brack.) Herter ex Nessel; Bärlappgewächse 30. 1939 Huperzia hamiltonii (Spreng.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution [complete]: Nepal, India (throughout, in hilly regions), Burma, Sri Lanka, Indochina, Malesia, Thailand (widespread), ?Taiwan Remarks: hamiltonii agg. = Huperzia hamiltonii (Spreng.) T. Sen & U. Sen; Fern Gazette 11(6): 419-420, f. 2, bb-hh (1978) [nom. illeg., non Huperzia hamiltonii (Spreng.) Trevis (1874) = Lycopodium hamiltonii Sprengel; Syst. veg. 5: 429 (index) (1828) = Lycopodium obtusifolium Buch.-Ham. ex D. Don; Prodromus Florae Nepalensis 18-19 (1825) [nom. illeg., non Lycopodium obtusifolium Sw. (1806) = Phlegmariurus hamiltonii (Spreng.) Á. Löve & D. Löve; Taxon 26(2-3): 324 (1977) = Urostachys hamiltonii (Spreng.) Herter ex Nessel; Die Bärlappgewächse 68 (1939) Huperzia hartwegiana (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution [complete]: Mexico (Chiapas), Guatemala, Colombia, Venezuela, Ecuador, N-Peru Remarks: taxifolia agg. = Huperzia caracasica (Herter) Holub; Folia Geobot. Phytotax. 20: 71 (1985) = Lycopodium caracasicum Herter; Hedwigia 49(2-3): 88, t. 3A (1909) = Lycopodium funckii Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 44 (1909) = Lycopodium hartwegianum Spring; Mém. Acad. Roy. Sci. Belgique 24: 14 (1850) = Urostachys caracasicus (Herter) Herter ex Nessel; Die Bärlappgewächse 90 (1939) = Urostachys funckii (Herter) Herter ex Nessel; Die Bärlappgewächse 170, t. 41 (1939) = Urostachys hartwegianus (Spring) Herter ex Nessel; Die Bärlappgewächse 90 (1939) = Urostachys maxonii Herter ex Nessel; Revista Sudamericana de Botánica 6(5-6): 165 (1940) Huperzia hastata B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 39, f. 6A (1988) Distribution [complete]: Ecuador Remarks: saururus agg. Huperzia hellwigii (Warb.) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: New Guinea Remarks: phlegmaria agg. = Lycopodium hellwigii Warb.; Monsunia 1: 97 (1898) = Urostachys hellwigii (Warb.) Herter ex Nessel; Bärlappgewächse 230 (1939) Huperzia hemleri (Nessel) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution [complete]: Brazil (Rio de Janeiro) Remarks: reflexa agg. = Urostachys hemleri Nessel; Arch. Bot. Sao Paulo 1: 389, t. 9 (1927) Huperzia henryi (Baker) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: Yunnan Remarks: hamiltonii agg. = Lycopodium henryi Baker; Bulletin of Miscellaneous Information Kew 1906(1): 15 (1906) = Phlegmariurus henryi (Baker) Ching; Acta Botanica Yunnanica 4(2): 125-126 (1982) = Urostachys henryi (Baker) Herter; Index Lyc. 64 (1949) Huperzia herteriana (Kümmerle) T. Sen & U. Sen; Fern Gazette 11(6): 415-417, f. 1, k-r (1978) Distribution: China, Sikkim, India (Sikkim, West Bengal, Manipur) Remarks: selago agg. = Lycopodium herterianum Kümm.; Magyar Bot. Lapok 26: 99 (1927 [1928]) = Lycopodium lucidulum auct. et Clarke; Trans. Linn. Soc. Lond. Il. Bot. 1-589, t. 84. f. 1 (1880) = Lycopodium sikkimense Herter; Bot. Jahrb. Syst. 43(1, Beibl. 98): 42-43 (1909) Bot. Jahrb. Syst. [nom. illeg., non Lycopodium sikkimense Müll. Hal (1861) = Urostachys herterianus (Kümmerle) Herter; Estudios Botánicos en la Región Uruguaya 20: 64 (1949) = Urostachys sikkimensis (Herter) Herter ex Nessel; Bärlappgewächse 52 (1939) Huperzia heterocarpon (Fée) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: Brazil (Santa Catarina, Sao Paulo), Argentina (Misiones) Remarks: heterocarpon agg. = ?Urostachys major (Nessel) Herter; Index Lyc. 69 (1949) = Urostachys spegazzinii Herter ex Nessel; Arch. Bot. Sao Paulo 1: 412 (1927) = Urostachys portoanus Nessel; Repert. Spec. Nov. Regni Veg. 39: 70, t. 194b (1935) = Lycopodium heterocarpon Fée; Crypt. vasc. Brésil 2: 93 (1872-73) = Urostachys heterocarpon (Fée) Herter; Repert. Spec. Nov. Regni Veg. 19: 164 (1923) = Huperzia longearistata (Christ) Holub; Folia Geobot. Phytotax. 20: 74 (1985) = Lycopodium longearistatum Christ; Schwacke, Pl. Novas Mineiras 2: 40 (1900) = Urostachys longearistatus (Christ) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 414 (1927) Huperzia heteroclita (Desv. ex Poir.) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution [complete]: Ecuador, Peru Remarks: myrsinites agg. = Lycopodium heteroclitum Desv. ex Poiret; Encycl. suppl. 3: 554 (1813 [1814]) Huperzia hexasticha B. Øllg. & Wind.; Bradea 5: 11, fig. 2 (1987) Distribution: Brazil (Sao Paulo) Remarks: quadrifariata agg. Huperzia hildebrandtii (Herter) Tardieu; Adansonia ser. 2 10: 18 (1970) Distribution [complete]: Madagascar = Lycopodium hildebrandtii Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 40 (1909) = Urostachys hildebrandtii (Herter) Herter ex Nessel; Die Bärlappgewächse 28, t. 4, f. 3 (1939) Huperzia hilliana (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: India (Travancore) Remarks: verticillata agg. = Urostachys hillianus Nessel; Repert. Spec. Nov. Regni Veg. 36: 186, t. 174 (1934) Huperzia hippuridea (H. Christ) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution [complete]: Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Mexico (Oaxaca, Chiapas), Guatemala, El Salvador Remarks: brongniartii agg. = Lycopodium hippurideum H. Christ; Prim. Fl. Costaric. 3(1): 56 (1901) = Lycopodium poseidonis (Herter) C. V. Morton; Amer. Fern Journal 54: 42 (1964) = Urostachys hippurideus (H. Christ) Herter ex Nessel; Die Bärlappgewächse 88 (1939) = Urostachys poseidonis Herter; Revista Sudamericana de Botánica 10: 122-123 (1953) Huperzia hippuris (Desv. ex Poiret) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) Distribution: Java Remarks: hippuris agg. = Lycopodium blumeanum Vriese; Tijdschr. Natuurl. Gesch. Physiol. 3: 338 (1844) = Lycopodium hippuris Desv. ex Poiret; Encycl. suppl. 3: 559 (1813 [1814]) = Urostachys hippuris (Poiret) Herter; Index Lyc. 65 (1949) Huperzia hochreutineri (Herter) comb. ined. Distribution: Java Remarks: unverified name, hippuris agg. = Lycopodium clavatum var. hochreutineri Nessel; Bärlappgewächse 300, t. f. 46. 1939 [nom. inval. = Lycopodium hochreutineri Alderw.; [erroneous citation in Herter (1949a: 23) = Lycopodium pinifolium Blume var. hochreutineri Herter; Annuaire Conserv. Bot. Geneve 15/16: 227 (1912) = Urostachys hochreutineri (Herter) Herter ex Nessel; Bärlappgewächse 234 (1939) = Urostachys mercurii Herter; Index Lyc. 69 (1949) Huperzia hoffmannii (Maxon) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988) Distribution [complete]: Costa Rica, Panama Remarks: reflexa agg. = Lycopodium hoffmannii Maxon; Contr. U.S. Natl. Herb. 17(4): 423 (1914) = Urostachys hoffmannii (Maxon) Herter ex Nessel; Die Bärlappgewächse 99 (1939) Huperzia hohenackeri (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution [complete]: Peru, Ecuador, Colombia Remarks: brevifolia agg. = Lycopodium hohenackeri Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 46 (1909) = Urostachys hohenackeri (Herter) Herter ex Nessel; Bärlappgewächse 105 (1939) Huperzia holstii (Hieron.) Pichi-Serm.; Webbia 23: 163 (1968) Distribution: Tanzania, Kenya Remarks: gnidioides agg. = Lycopodium holstii Hieron.; Engler, Pflanzenw. Ost-Afrikas C: 90 (1895) = Urostachys holstii (Hieron.) Herter ex Nessel; Bärlappgewächse 202 (1939) Huperzia homocarpa (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution [complete]: Costa Rica, Panama, Colombia, Venezuela, Ecuador, Hispaniola (Dominican Republic, Haiti) Remarks: dichotoma agg. = Lycopodium homocarpum (Herter) C. Chr.; Kongl. Svenska Vetenskapsakad. Handl., 3. Ser. 16 (2): 92 (1937) = Urostachys homocarpus Herter; Repert. Spec. Nov. Regni Veg. 19: 163 (1923) Huperzia hookeri (W.J. Hook. & Grev.) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution: Nepal Remarks: squarrosa agg. = Lycopodium hookeri Wallich ex Hook. & Grev.; Icon. filic. 2: t. 185. 1831 [1830 or 1831] Huperzia horizontalis (Nessel) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: Borneo Remarks: phlegmaria agg. = Lycopodium aellenii (Herter) Tag.; Acta Phytotax. Geobot. 26: 108 (1974) = Lycopodium horizontale Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 7: 30. 1912 [non C. Presl (1825) = Urostachys horizontales Herter ex Nessel; Bärlappgewächse 225 (1939) Huperzia huberi B. Øllg.; Amer. Fern Journal 79: 153 (1989) Distribution [complete]: Venezuela Huperzia humbertii-henrici (Herter) Tardieu; Adansonia ser. 2 10: 18 (1970) Distribution [complete]: Madagascar Remarks: gnidioides agg. = Urostachys humbertii-henrici Herter; Revista Sudamer. Bot. 8: 23 (1949) Huperzia hupehensis Ching; Acta Bot. Yunnanica 3: 301 (1981) Distribution: W-China Remarks: selago agg. Huperzia hypogaea B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 58, f. 8E (1988) Distribution [complete]: Ecuador, Peru, Colombia Remarks: saururus agg. Huperzia hystrix (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985) Distribution: Ecuador Remarks: saururus agg. = Urostachys hystrix Herter; Revista Sudamer. Bot. 10: 120 (1953 [1954]) = Lycopodium hystrix (Herter) C. Morton; Amer. Fern J. 54: 72 (1964) Huperzia Iajouensis Ching; Acta Bot. Yunnanica 3: 305 (1981) Distribution: S-Tibet Remarks: unverified name Huperzia ignambiensis (Compton) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991) Distribution [complete]: New Caledonia Remarks: verticillata agg. = Lycopodium ignambiense Compton; J. Linn. Soc. Bot. 45: 458 (1922) Huperzia innocentia (Herter) comb. ined. Distribution: South America Remarks: unverified name, saururus agg. = Lycopodium innocentium (Herter) C. Morton; Amer. Fern 54: 72 (1964) = Urostachys innocentium Herter; Revista Sudamer. Bot. 10: 127 (1953 [1954]) Huperzia insularis (Carmich.) Rothm.; Feddes Repert. 54: 60 (1944) Distribution: Tristan da Cunha Remarks: unverified name, saururus agg. = Lycopodium insulare Carmich.; Trans. Linn. Soc. London 12. 509 (1819) = Urostachys insularis (Carmich.) Herter ex Nessel; Bärlappgewächse 81 (1939) Huperzia intermedia Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) [nom. nov. for Lycopodium intermedium Spring, non Blume Distribution [complete]: Brazil, Venezuela (Tepuis), Guadeloupe, British Guyana (Mt. Roraima) Remarks: reflexa agg. = Lycopodium commutatum Spring ex Klotzsch; Linnaea 18: 517 (1844) [nom. inval. = Lycopodium intermedium Spring; C. Martius, Fl. Bras. 1, 2: 111 (1840) [nom. illeg., non Lycopodium intermedium Blume (1828) = Urostachys commutatus (Spring ex Klotzsch) Herter; Estudios Botánicos en la Región Uruguaya 20: 57 (1949), Alt. title: Index Lycopodiarum 57 (1949) = Urostachys intermedius Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 163 (1923) Huperzia itambensis B. Øllg. & Wind.; Bradea 5: 12 (1987) Distribution: Brazil (Minas Gerais) Remarks: brongniartii agg. Huperzia jaegeri (Herter) Pichi-Serm.; Webbia 23: 163 (1968) Distribution: Sierra Leone, Guinea Remarks: hamiltonii agg. = Urostachys jaegeri Herter; Revista Sudamer. Bot. 8: 24 (1949) = Lycopodium jaegeri (Herter) Alston; Mem. Inst. Franc. Afrique Noire 50: 21 (1957) Huperzia juniperistachya (Hayata) Holub; Folia Geobot. Phytotax. 20: 74 (1985) Distribution: Taiwan Remarks: phyllantha agg. = Lycopodium juniperistachyum Hayata; Ic. pl. formos. 4: 132 (1914) = Urostachys juniperistachyus (Hayata) Herter ex Nessel; Bärlappgewächse 70 (1939) Huperzia kajewskii (Copel.) comb. ined. Distribution: Santa Cruz Isl. Remarks: unverified name, phlegmaria agg. = Lycopodium kajewskii Copel.; J. Arnold Arbor. 12: 49 (1931) = Urostachys kajewskii (Copel.) Herter; Index Lyc. 66 (1949) Huperzia kamaensis Ching & S.K. Wu; Acta Botanica Yunnanica 3(3): 300 (1981) Distribution: Tibet, Yunnan Remarks: selago agg. = Huperzia kamaensis Ching & S.K. Wu; Flora Xizangica 1: 9, t. 2, f. 4-6, t. 1, f. 1-3 (1983) [nom. illeg., superfl. Huperzia kangdingensis (Ching) Ching; Acta Botanica Yunnanica 3(3): 294 (1981) Distribution: Sichuan Remarks: selago agg. = Lycopodium kangdingense Ching; Acta Phytotaxonomica Sinica 18(2): 236-237, f. 1: 4, t. 6, f. 3 (1980) Huperzia kaysseri (Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985) Distribution: New Guinea Remarks: unverified name, phlegmaria agg. = Urostachys kaysseri Nessel; [Bärlappgewächse 31, t. 4, f. 10. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 156 (1940) Huperzia kesselringii (Nessel) comb. ined. Distribution: Sulawesi Remarks: unverified name, phlegmaria agg. = Urostachys kesselringii Nessel; Revista Sudamer. Bot. 6: 167, t. 14, f. 67 (1940) Huperzia keysseri (Herter & Schlechter ex Diels) comb. ined. Distribution: Papua New Guinea Remarks: unverified name, phlegmaria agg. = Lycopodium keysseri Herter & Schlechter ex Diels; Bot. Jahrb. Syst. 62: 456 (1929) Huperzia killipii (Herter) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution [complete]: Colombia, Venezuela, French Guiana Remarks: taxifolia agg. = Urostachys killipii Herter; Revista Sudamer. Bot. 10: 128 (1953 [1954]) = Lycopodium killipii (Herter) C. Morton; Amer. Fern J. 54: 72 (1964) Huperzia kuesteri (Nessel) B. Øllg.; Opera Bot. 92: 169 (1987) Distribution [complete]: Ecuador, Peru Remarks: saururus agg. = Urostachys kuesteri Nessel; Repert. Spec. Nov. Regni Veg. 35: 182p t. 172 (1934) Huperzia kunmingensis Ching; Acta Bot. Yunnanica 3: 297 (1981) Distribution: Yunnan Remarks: unverified name, selago agg.

          Huperzia species A to K.
          For Huperzia species L to Z including hybrids go here

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