Family Lycopodiaceae, genus Huperzia; world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Huperzia, giving current name, authority, synonyms, published references, geographic distribution and (perhaps) illustrations.

          Huperzia species L to Z including hybrids.
          For Huperzia species A to K go to here

001.00 Class Lycopsida
Trivial names: Lycophytes, Lycopods
Statistics: 1340 spp. [1339 species and 28 hybrids listed]
001.000 Order Lycopodiales
Statistics: 4 genera, 490 spp. [504 species and 10 hybrids listed]
001.0000 Family Lycopodiaceae Beauv. ex Mirb.; Hist. Nat. Veg. (Lam. & Mirb.) 4: 293 (1802)
Trivial names: Bärlappgewächse, Club-mosses
Statistics: 4 genera, 490 spp. [504 species and 10 hybrids listed]
= Huperziaceae Rothm.; Fedde Repert. 66: 236 (1962) [nom. nov.]
= Urostachyaceae Rothm.; Fedde Repert. 54: 58 (1944) [nom. illeg.]
= Phylloglossaceae Kze.; Bot. Zeit. 1: 722 (1843)

001.0002 Huperzia Bernhardi; Schrad. J. Bot. 1800(2): 126 (1801)
Statistics: ca. 400 spp. nearly worldwide, Europe (1) [409 species and 6 hybrids listed]
Trivial names: Teufelsklaue
Revision date: 20.07.2001; 19.09.2006
= Mirmau Adans.; Fam. Pl. 2: 491. Jul-Aug (1763) [p. p., nom. illegit.]
= Phlegmariurus (Herter) Holub; Preslia 36 (1): 179 21 (1964)
= Plananthus P. Beauv. ex Mirbel; Lam. & Mirbel, Hist. nat. vég. 3: 476, 4: 310 (1802) [nom. superfl., by lectotypification of Huperzia Bernh.]
= Selago Boehmer; Ludwig, Defin. Gen. Pl. ed. Boehmer 484. 1760 [non L. 1753, nec Browne 1756, nec Hill (1757) nom. illeg.]
= Selago J. Hill; Brit. Herb. 533. 1757 (28 jan.) [non L. 1753, nec Browne 1756] nom. inval. (Pichi Sermolli (1971: 140))]
= Selago Schur; Enum. pl. Transsilv. 825. 1866 [non L. 1753, nec Browne 1756, nec Hill (1757])]
= Urostachys Herter; Beih. Bot. Centralbl. 39(2): 249 (1922) [nom. superfl.]
Huperzia laipoensis Ching; Acta Bot. Yunnanica 3: 300 (1981)
Distr.: S-Sichuan
Remarks: selago agg.
Huperzia lajouensis Ching; Acta Botanica Yunnanica 3(3): 305 (1981)
Distr.: Tibet
Remarks: selago agg.
= Huperzia lajouensis Ching; Flora Xizangica 1: 7-8 (1983) [nom. illeg., superfl.]
Huperzia lancifolia (Maxon) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr. [complete]: Panama
Remarks: taxifolia agg.
= Lycopodium lancifolium Maxon; Contr. U.S. Natl. Herb. 17(2): 177 (1913)
= Urostachys lancifolius (Maxon) Herter; Index Lyc. 66 (1949)
Huperzia lauterbachii (E. Pritzel ex Schumann & Lauterb.) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: New Guinea
Remarks: squarrosa agg.
= Lycopodium lauterbachii E. Pritzel ex Schumann & Lauterb.; Fl. Schutzgeb. Südsee 149 (1900)
= Urostachys lauterbachii (Schumann & Lauterb.) Herter ex Nessel; Bärlappgewächse 203 (1939)
Huperzia laxa (Desv.) A. Khokhr.
Distr.: C-Siberia, E-Siberia / Amur / Ussuri
= Lycopodium selago L. ssp. laxum Desv.
Huperzia lechleri (Hieron.) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: Peru, Ecuador
Remarks: brongniartii agg.
= Lycopodium lechleri Hieron.; Bot. Jahrb. Syst. 34: 571 (1905)
= Urostachys lechleri (Hieron.) Herter ex Nessel; Bärlappgewächse 85 (1939)
= Urostachys lechleri (Hieron.) Nessel var. lehmannii Nessel; [Bärlappgewächse 86. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 161, t. 8, f. 35. 1940]
= Urostachys lehmannii (Nessel) Herter; Index Lyc. 67 (1949)
Huperzia lecomteana (Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: Madagascar
Remarks: squarrosa agg.
= Urostachys lecomteanus Nessel; Repert. Spec. Nov. Regni Veg. 36: 1871 t. 174 (1934)
Huperzia ledermannii (Herter) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: New Guinea
Remarks: phlegmaria agg.
= Lycopodium ledermannii Herter; Bot. Jahrb. Syst. 54: 232 (1916)
= Lycopodium phlegmaria L. var. brachystachyum Rosenstock; Repert. Spec. Nov. Regni Veg. 12: 530 (1913)
= Urostachys ledermannii (Herter) Herter ex Nessel; Bärlappgewächse 236 (1939)
Huperzia leipoensis Ching; Acta Botanica Yunnanica 3(3): 300 (1981)
Distr.: China
Huperzia leishanensis X.Y. Wang; Bulletin of Botanical Research 14(4): 355-356, f. 1-3 (1994)
Distr.: Yunnan
Huperzia leitzii (Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: Brazil (Sao Paulo)
Remarks: heterocarpon agg.
= Urostachys leitzii Nessel; Repert. Spec. Nov. Regni Veg. 36: 184, t. 173 (1934)
Huperzia leptodon (Herter) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
Distr.: Colombia
Remarks: reflexa agg.
= Urostachys leptodon Maxon ex Herter; Revista Sudamer. Bot. 10: 120 (1953 [1954])
= Lycopodium leptodon (Herter) C. Morton; Amer. Fern 1. 54: 72 (1964)
Huperzia liangshanica (H.S. Kung) Ching & H.S. Kung; Acta Botanica Yunnanica 3(3): 296 (1981)
Distr.: Sichuan
Remarks: selago agg.
= Lycopodium liangshanicum H.S. Kung; Acta Botanica Yunnanica 2(1): 100-101, f. 1-3 (1980)
Huperzia lignosa (Herter) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: Colombia
Remarks: brevifolia agg.
= Lycopodium lignosum Herter; Bot. Jahrb. Syst. 43: Beibl.- 98: 46 (1909)
= Urostachys lignosus (Herter) Herter ex Nessel; Bärlappgewächse 104 (1939)
Huperzia lindenii (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 247 (1874)
Distr.: Colombia, Ecuador
Remarks: taxifolia agg.
= Huperzia sodiroana (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Huperzia stuebelii (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Huperzia wohlberedtii (Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
= Lycopodium lindenii Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 27 (1849)
= Lycopodium sodiroanum Herter; Hedwigia 49. 91 (1909)
= Lycopodium stuebelii Hieron. et Herter ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 53 (1909 (Feb.))
= Lycopodium wohlberedtii (Nessel) C. Morton; Amer. Fern J. 54: 73 (1964)
= Urostachys lindenii (Spring) Herter ex Nessel; Bärlappgewächse 177 (1939)
= Urostachys sodiroanus (Herter) Herter ex Nessel; Bärlappgewächse 240 (1939)
= Urostachys stuebelii (Herter) Herter ex Nessel; Bärlappgewächse 241 (1939)
= Urostachys wohlberedtii Nessel; Repert. Spec. Nov. Regni Veg. 39: 69, t. 193c (1935)
Huperzia linifolia (L.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
Distr. [complete]: Puerto Rico, Costa Rica, Greater Antilles, Mexico, Chiapas, Belize, Guatemala, Hondruas, Nicaragua, Panama, Bolivia, &, Brazil, Cuba, Nicaragua, Colombia, Venezuela, Ecuador, Greater Antilles, Hispaniola, Martinique, St. Kitts, Guadeloupe, Dominica, St. Lucia, St. Vincent, Grenada, French Guiana
Remarks: linifolia agg.
= Lycopodium linifolium L.; Sp. Pl. 1100 (1753)
= Lycopodium linifolium L. var. subaristatum H. Christ; Bulletin de la Société Botanique de Genève 1(5): 236 (1909)
= Plananthus linifolius (L.) P. Beauv.; Prod. aethéogam. 111 (1805)
= Urostachys linifolia (L.) Herter; Repert. Spec. Nov. Regn. Veg. 19: 165 (1923)
          var. jenmanii (Underw. & F.E. Lloyd) B. Øllg. & P.G. Windisch; Bradea 5: 13 (1987)
Distr. [complete]: Brazil, Colombia, Ecuador, Peru, British Guyana, Surinam, French Guiana
= Huperzia jenmanii (Underw. & F.E. Lloyd) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium jenmanii L. Underw. & F. Lloyd; Bull. Torrey Bot. Club 33: 112 (1906)
= Urostachys jenmanii (L. Underw. & F. Lloyd) Herter ex Nessel; Bärlappgewächse 158 (1939)
          var. tenuifolia (Nessel) B. Øllg.; Flora of Ecuador 33: 90 (1988)
Distr.: Ecuador
= Urostachys linifolius var. tenuifolius Nessel; Revista Sudamericana de Botánica 6(5-6): 164 (1940)
          var. gracile (L'Herm. ex Fée) comb. ined.
Distr.: Jamaica, Hispaniola, N-South America, Guadeloupe, St. Lucia
= Lycopodium linifolium var. gracile L'Herm. ex Fée; Mem. 11: 129 (1866)
Huperzia llanganatensis B. Øllg.; Flora of Ecuador 33: 35 (1988)
Distr. [complete]: Ecuador
Remarks: saururus agg.
Huperzia lockyeri (D. Jones & B. Gray) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distr.: Queensland
Remarks: carinata agg.
Conservation status: IUCN V
= Lycopodium lockyeri Jones & Gray; Austrobaileya 2: 126, f. 1 (1985)
Huperzia loefgreniana (Silveira) B. Øllg. & Wind.; Bradea 5: 13 (1987)
Distr.: Brazil (Sao Paulo)
Remarks: heterocarpon agg.
= Lycopodium loefgrenianum Silveira; Fl. serr. Min. 78, t. 30 (1908)
= Urostachys loefgrenianus (Silveira) Herter; Index Lyc. 68 (1949)
Huperzia longa (Copel.) comb. ined.
Distr.: Solomons
Remarks: squarrosa agg.
= Lycopodium longum Copel.; Philippine Journal of Science 60: 100, t. 3 (1936)
= Urostachys longus (Copel.) Herter; Estudios Botánicos en la Región Uruguaya 20: 68 (1949), Alt. title: Index Lycopodiarum 68 (1949)
Huperzia longyangensis (C.Y. Ma) comb. ined.
Distr. [complete]: China
= Phlegmariurus longyangensis C.Y. Ma (1990)
Huperzia loxensis B. Øllg.; Harling & Andersson: Flora of Ecuador 33: 17, f. 3B (1988)
Distr. [complete]: Ecuador Remarks: brongniartii agg.
Huperzia lucidula (Michx.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
Trivial names: Shining Clubmoss
Distr.: Newfoundland, E-Canada, NE-USA, EC-USA, C-USA, China
Remarks: selago agg.
= Lycopodium lucidulum Michx.; Fl. Bor.-Amer. 2: 284 (1803)
= Lycopodium reflexum (P. Beauv.) Sw.; Syn. fil. 175. 1806 [non Lam. 1792]
= Selago americana Dill. (Herter 1949a: 49) [nom. inval. (polynom.)]
= Urostachys lucidulus (Michx.) Herter ex Nessel; Bärlappgewächse 49. 1939
          var. asiatica Ching; Acta Botanica Yunnanica 3(3): 296 (1981)
Distr.: Distribution unknown
Huperzia luederwaldtii (Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: Brazil
Remarks: brongniartii agg.
= Urostachys luederwaldtii Nessel; Repert. Spec. Nov. Regni Veg. 36: 178, t. 170 (1934)
Huperzia macbridei (Herter) B. Øllg.; Opera Bot. 92: 169 (1987)
Distr.: Peru, Ecuador
Remarks: saururus agg.
= Urostachys macbridei Maxon ex Herter; Revista Sudamer. Bot. 10: 115 (1953 [1954])
= Lycopodium macbridei (Herter) C. Morton; Amer. Fern J. 54: 72 (1964)
Huperzia macgregorii (Baker) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: New Guinea
Remarks: macgregorii agg.
= Lycopodium macgregorii Baker; J. Bot. 28: 109 (1890)
= Urostachys macgregorii (Baker) Herter ex Nessel; Bärlappgewächse 184 (1939)
= Huperzia versteegii (Alderw.) Holub; Folia Geobot. Phytota,x. 20: 77 (1985)
= Lycopodium versteegii Alderw.; Bull. Jard. Bot. Buitenzorg ser. 2, 24: 4 (1917)
= Urostachys versteegii (Alderw.) Herter ex Nessel; Bärlappgewächse 182 (1939)
= Huperzia whartoniensis (Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
= Urostachys whartoniensis Nessel; Repert. Spec. Nov. Regni Veg. 39: 62, t. 190b (1935)
Huperzia macrostachys (Spring) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: India, Sri Lanka
Remarks: phyllantha agg.
= Lycopodium macrostachys Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 30 (1849)
= Urostachys macrostachys (Spring) Herter ex Nessel; Bärlappgewächse 211 (1939)
Huperzia maerhkangensis Ching; Acta Bot. Yunnanica 3. 295 (1981)
Distr.: NW-Sichuan
Remarks: selago agg.
Huperzia mafuluensis (Nessel) comb. ined.
Distr.: New Guinea
Remarks: phlegmaria agg.
Huperzia magnifica (Brownlie) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distr.: Fiji
Remarks: dalhousiana agg.
= Lycopodium magnificum Brownlie; Beih. Nova Hedwigia 55: 22-23, t. 2e f. 3-4 (1977)
Huperzia magnusiana (Herter) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: Philippines
Remarks: squarrosa agg.
= Lycopodium magnusianum Herter; Hedwigia 49(2-3): 91 (1909)
= Urostachys magnusianus (Herter) Herter; Bot. Arch. 3: 14 (1923)
Huperzia mannii (Hillebr.) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distr. [complete]: Hawaii (Kauai, Maui, Hawaii Isl.)
Remarks: myrsinites agg.
Conservation status: IUCN V
= Lycopodium mannii (Hillebrand) Skottsb.; Acta Horti Gothob. 15: 131-132 (1942)
= Lycopodium phlegmaria var. mannii Hbd.; Fl. Hawaiian Isl. 645 (1888)
= Phlegmariurus mannii (Hbd.) W. H. Wagner; Contr. Univ. Michigan Herb. 20: 241 (1995)
Huperzia marsupiiformis (D. Jones & B. Gray) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distr.: NE-Queensland
Remarks: phlegmaria agg.
Conservation status: IUCN V
= Lycopodium marsupiiforme Jones & Gray; Austrobaileya 2: 128, f. 2 (1985)
Huperzia martii (Wawra) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: Brazil
Remarks: heterocarpon agg.
= Lycopodium martii Wawra; Oesterr. Bot. Z. 13: 218 (1863)
= Urostachys martii (Wawra) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 385 (1927)
Huperzia medogensis Ching & Ling; Acta Phytotax. Sin. 22 (3): 193 (1984)
Distr.: Tibet
Remarks: selago agg.
Huperzia megastachya (Baker) Tard.; Adansonia 10: 18 (1970)
Distr.: Madagascar
Remarks: squarrosa agg.
= Lycopodium megastachyum Baker; J. Linn. Soc. Bot. 21: 454 (1885)
= Urostachys megastachyus (Baker) Herter ex Nessel; Bärlappgewächse 210 (1939)
Huperzia melanesica (Brownlie) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distr.: Fiji
Remarks: varia agg.
= Lycopodium melanesicum Brownlie; Beih. Nova Hedwigia 55: 20i t. 2, f. 1-2 (1977)
Huperzia merrillii (Herter) Holub; Folia Geobot. Phytotax. .20: 74 (1985)
Distr.: Luzon
Remarks: hamiltonii agg.
= Urostachys merrillii Herter; Bot. Arch. 3: 15 (1923)
= Lycopodium merrillii (Herter) Copel.; Univ. Calif. Publ. Bot. 12: 384 (1931)
Huperzia mesoamericana B. Øllg.; Novon 3(1): 67, f. 1 (1993)
Distr. [complete]: Panama, Costa Rica
Huperzia mexicana (Herter) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
Distr. [complete]: S-Mexico, Chiapas, Guatemala, Honduras, El Salvador, Greater Antilles
Remarks: brongniartii agg.
= Lycopodium mexicanum Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 49 (1909)
= Urostachys mandiocanus var. mexicanus (Herter) Herter ex Nessel; Die Bärlappgewächse 149 (1939)
= Urostachys mexicanus (Herter) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 164 (1923)
Huperzia mildbraedii (Herter) Pichi-Serm.; Webbia 23: 163 (1968)
Distr.: Cameroun, Tanzania, Principe Isl., Sao Tome
Remarks: gnidioides agg.
= Lycopodium mildbraedii Herter; Hedwigia 49(2-3): 90 (1909)
= Urostachys mildbraedii (Herter) Herter ex Nessel; Bärlappgewächse 188 (1939)
Huperzia mingcheensis (Ching) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: S-Zhejiang
Remarks: hippuris agg.
= Phlegmariurus mingcheensis Ching; Acta Bot. Yunnanica 4: 125 (1982)
= Phlegmariurus mingcheensis var. angustifolius C.Y. Ma; Bulletin of Botanical Research 10(3): 57-58, t. 1 (1990)
Huperzia miniata (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17- 248 (1874)
Distr.: Java, Philippines
Remarks: selago agg.
= Lycopodium miniatum Spring; Bull. Acad. Roy. Sci. Belg. 8: 513 (1841)
= Lycopodium selago var. miniatus (Spring) B.C. Tan & Tolentino; Philippine Journal of Science 116(4): 437-438 (1987)
= Urostachys miniatus (Spring) Herter ex Nessel; Bärlappgewächse 42 (1939)
Huperzia minima (Herter) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: Philippines
Remarks: selago agg.
= Urostachys minimus Herter; Bot. Arch. 3: 13 (1923)
Huperzia minimadenta Cheng; Bull. Bot. Res. North-East. Forest. Inst. 7 (3): 113 (1987)
Distr.: Jiangxi
Remarks: selago agg.
Huperzia minutifolia (Alderw.) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distr.: New Guinea
Remarks: phlegmaria agg.
= Lycopodium coralium Spring var. minutifolium Alderw.; Malayan fern allies 44 (1915)
= Urostachys minutifolius (Alderw.) Herter; Index Lyc. 70 (1949)
= Lycopodium minutifolium (Alderw.) Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 28: 44 (1918)
Huperzia mira (Vareschi) comb. ined.
Distr.: Venezuela
Remarks: brevifolia agg.
= Lycopodium mirum Vareschi; Acta Biol. Venez. 2: 317, f. 23 (1958)
Huperzia mirabilis (Willd.) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distr.: Vietnam
Remarks: phlegmaria agg.
= Lycopodium mirabile Willd.; Sp. pl. 5: 11 (1810)
= Urostachys mirabilis (Willd.) Herter ex Nessel; Bärlappgewächse 225 (1939)
Huperzia miyoshianum (Makino) Ching; Acta Bot. Yunnan 3(3): 303, 304 (1981)
Distr.: E-Siberia / Amur / Ussuri, Japan, Korea, S-Alaska, W-British Columbia, pacific coast south, to, Washington, isolated Alberta
Remarks: selago agg.
= Huperzia selago var. miyoshiana (Makino) Taylor & MacBryde; Canadian Journal of Botany 56(2): 185 (1978)
= Lycopodium miyoshianum Makino; Bot. Mag. (Tokyo) 12: 36 (1898)
= Lycopodium selago ssp. miyoshianum (Makino) Calder & Taylor; Canadian Journal of Botany 43(11): 1388 (1965)
= Lycopodium selago var. miyoshianum (Makino) Makino; Botanical Magazine 16(189): 199 (1902)
= Lycopodium tenuifolium Herter; Bot. Jahrb. Syst. 43(1, Beibl. 98): 41 (1909)
= Urostachys miyoshianum (Makino) Herter ex Nessel; Die Bärlappgewächse 28, Abb. 4, f. 4 (1939)
          var. coreana (Hayata) Ching; Acta Botanica Yunnanica 3(3): 304 (1981)
Distr.: Korea
= Lycopodium miyoshianum var. coreanum Hayata; Icones plantarum formosanarum nec non & contributiones ad floram formosanam. 5: 255 (1915)
= Urostachys coreanus Herter; Index Lyc. 57 (1949)
= Urostachys cryptomerinus (Maxim.) Nessel var. coreanus Herter ex Nessel; [Bärlappgewächse 72, t. 11, f. 8. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 160. 1940]
= Urostachys miyoshiana var. coreanus (Hayata) Herter ex Nessel; Die Bärlappgewächse 28, Abb. 4, f. 5 (1939)
Huperzia mollicoma (Spring) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr. [complete]: Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia, SE-Brazil, C- Brazil
Remarks: verticillata agg.
= Huperzia flaccida (Fée) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
= Lycopodium dichotomum ssp. mollicomum Spring; Flora 21(1): 162 (1838)
= Lycopodium flaccidum Fée; Cryptogames Vasculaires . . . du Brésil 2: 92, t. 106, f. 1 (1872)-73 [nom. illeg., non Lycopodium flaccidum Bory (1833)]
= Lycopodium molllicomum (Spring) Spring; Flora Brasiliensis 1(2): 113 (1840)
= Lycopodium setaceum Lam. var. brasiliense Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 43 (1842)
= Lycopodium williamsii Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33(2): 112 (1906)
= Urostachys flaccidus (Fée) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 164 (1923)
= Urostachys mollicomus (Spring) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 407, t. 25 (1927)
= Urostachys neptuni Herter; Estudios Botánicos en la Región Uruguaya 20: 72 (1949) [nom. illeg. superfl.]
= Urostachys williamsii (Underw. & Lloyd) Herter ex Nessel; Die Bärlappgewächse 164 (1939)
= Urostachys mollicomus (Spring) Nessel var. major Nessel; Repert. Spec. Nov. Regni Veg. 36: 183, t. 172. 1934
Huperzia molongensis (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Ecuador
Remarks: phlegmaria agg.
= Lycopodium molongense Herter; Bot. Jahrb. Syst. 43: Beibl. 98.- 51 (1909)
= Urostachys molongensis (Herter) Herter ex Nessel; Bärlappgewächse 240 (1939)
Huperzia montana (L. Underw. & F. Lloyd) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Jamaica, Cuba
Remarks: brongniartii agg.
= Lycopodium hippurideum var. montanum (Underw. & F.E. Lloyd) Proctor; Amer. Fern Journal 72: 107 (1982)
= Lycopodium montanum Underw. & F.E. Lloyd; Bulletin of the Torrey Botanical Club 33: 107 (1906)
= Urostachys hippurideus var. montanus (Underw. & F.E. Lloyd) Herter ex Nessel; Die Bärlappgewächse 88 (1939)
= Urostachys montanus (Underw. & F.E. Lloyd) Herter; Estudios Botánicos en la Región Uruguaya 20: 70 (1949), Alt. title: Index Lycopodiarum 70 (1949)
Huperzia monticola (Alderw.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Sumatra
Remarks: selago agg.
= Lycopodium monticolum Alderw.; Malayan ferns & fern allies, Suppl. 1, corr.: 65. 1917 [Bull. Jard. Bot. Buitenzorg, ser. 2, 28: 43, t. 7 (1918])
= Urostachys monticolus (Alderw.) Herter ex Nessel; Bärlappgewächse 120 (1939)
Huperzia mooreana (Baker) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Brazil (Bahia)
Remarks: brongniartii agg.
= Lycopodium luetzelburgii Rosenstock; Repert. Spec. Nov. Regni Veg. 20: 95 (1924)
= Huperzia sydowiora (Herter) Rolleri & Deferrari; Notas del Museo de la Plata, Botánica 21(100): 157 (1988)
= Lycopodium sydowiorum Herter; Repert. Spec. Nov. Regn. Veg. 13: 296 (1914)
= Urostachys sydowiorum (Herter) Herter ex Nessel; Archivos de Botânica do São Paulo 1: 390 (1927)
= Lycopodium mooreanum Hort. Sander ex Baker; Gard. Chron., ser. 3, 12: 582 (1892)
= Urostachys mooreanus (Baker) Herter; Index Lyc. 70 (1949)
Huperzia mortonii (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Guatemala
Remarks: dichotoma agg.
= Urostachys mortonii Herter; F. Morton; Repert. Spec. Nov. Regni Veg. 28: 108 (1930)
Huperzia multidichotoma Ching; Acta Bot. Yunnanica 3: 295 (1981)
Distr.: Sichuan
Remarks: selago agg.
Huperzia multifaria (Alderw.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Ternate
Remarks: hippuris agg.
= Lycopodium multifarium Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 5: 226 (1922)
= Urostachys multifarius (Alderw.) Herter ex Nessel; Bärlappgewächse 207 (1939)
Huperzia myriophyllifolia (Hayata) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Taiwan
Remarks: selago agg.
= Lycopodium serratum Thunb. ex Murray var. myriophyllifolium Hayata; Ic. pl. formos. 4: 133 (1914)
= Urostachys myriophyllifolius (Hayata) Herter; Index Lyc. 71 (1949)
Huperzia myrsinites (Lam.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1874)
Distr. [complete]: S-Mexico, Chiapas, Guatemala, Honduras, El Salvador, Nicaragua, Costa Rica, Colombia, Venezuela, Ecuador, Peru, Hispaniola, Trinidad, British Guyana
Remarks: myrsinites agg.
= Lycopodium myrsinites Lam.; Encyclopédie Méthodique, Botanique 3: 654. [1789] (1792)
= Lycopodium patens Willd. ex Spreng.; Systema Vegetabilium ed. 14 4: 12 (1827)
= Lycopodium roraimense Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33: 115 (1906)
= Lycopodium skutchii Maxon; Proc. Biol. Soc. Wash. 46: 159 (1933)
= Plananthus myrsinites (Lam.) P. Beauv.; Prodr. Aetheog. 111 (1805)
= Urostachys myrsinites (Lam.) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 166 (1923)
= Urostachys patens (Willd. ex Spreng.) Herter ex Nessel; Die Bärlappgewächse 245, t. 59, f. 25 (1939)
= Urostachys roraimensis (Underw. & Lloyd) Herter ex Nessel; Die Bärlappgewächse 246 (1939)
Huperzia myrtifolia (G. Forster) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: New Caledonia, Queensland, Tasmania
Remarks: myrtifolia agg.
= Lycopodium myrtifolium G. Forster; Fl. ins. austr. 87 (1786)
= Stachygynandrum myrtifolium (G. Forster) P. Beauv.; Prodr. aethéogam. 113 (1805)
= Urostachys myrtifolius (G. Forster) Herter; Index Lyc. 71 (1949)
Huperzia myrtuosa (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
Distr.: Colombia, Venezuela
Remarks: brevifolia agg.
= Huperzia lellingeri (Rolleri) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Lycopodium lellingeri Rolleri; Hickenia 1 (26): 143 (1978)
= Lycopodium myrtuosum Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 9 (1849)
= Urostachys myrtuosus (Spring) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 387 (1927)
Huperzia nana (C.Y. Ma) comb. ined.; [nom. illeg., non Herter 1814]
Distr.: China
= Phlegmariurus nanus C.Y. Ma (1990)
Huperzia nana (Herter) comb. ined.
Distr.: Ecuador
Remarks: brevifolia agg.
= Lycopodium nanum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 46. 1909 [non Poiret (1814)]
= Urostachys deuteronanus Herter; Index Lyc. 59 (1949)
= Urostachys nanus (Herter) Herter ex Nessel; Bärlappgewächse 104 (1939)
Huperzia nanchuanensis (Ching & H.S. Kung) Ching & H.S. Kung;
Acta Botanica Yunnanica 3(3): 302 (1981)
Distr.: Sichuan
Remarks: selago agg.
= Lycopodium nanchuanense Ching & Kung ex Kung; Acta Phytotax. Sin. 18 (2): 235, f. 5, t. 6, f. 2 (1980)
Huperzia neocaledonica (Nessel) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: New Caledonia
Remarks: hippuris agg.
= Urostachys neocaledonicus Nessel; Repert. Spec. Nov. Regni Veg. 36: 187, t. 175 (1934)
Huperzia nesselii (Brause ex Nessel) Rolleri & Deferrari; Notas del Museo de la Plata,
Botánica 21(100): 156 (1988)
Distr.: Peru
Remarks: saururus agg.
= Urostachys nesselii Brause ex Nessel; [Bärlappgewächse 78, f. 12. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 161 (1940)]
= Lycopodium nesselii (Nessel) Rolleri; Revista Mus. La Plata (N. S.) Bot. 13: 77 (1981)
Huperzia nilagirica (Spring) Dixit; J. Bombay Nat. Hist. Soc. 77 (3): 541 (1981)
Distr.: India (Nilgiri Hills, Tamil Nadu, Kerala)
Remarks: hamiltonii agg.
= Lycopodium niligaricum Spring; Bull. Acad. Sci. Belg. 15 (1): 58 (1843)
Huperzia novae-zelandica (Colenso) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: New Zealand
Remarks: varia agg.
= Lycopodium novae-zelandicum Colenso; Trans. & Proc. New Zealand Inst. 19: 275 (1887)
Huperzia nubigena (Herzog) comb. ined.
Distr.: Bolivia
Remarks: myrsinites agg.
= Lycopodium nubigenum Herzog; Meded. Rijks-Herb. 27: 2 (1915)
= Urostachys nubigenus (Herzog) Herter ex Nessel; Bärlappgewächse 246 (1939)
Huperzia nuda (Nessel) B. Øllg. & Wind.; Bradea 5: 14 (1987)
Distr.: Brazil
Remarks: brongniartii agg.
= Urostachys nudiusculus Herter; Index Lyc. 72. 1949 [nom. superfl. for Urostachys nudus Nessel nudus Christ ex Nessel, Arch. Bot. Sao Paulo 1: 396, t. 14 (1927)]
Huperzia nummulariifolia (Blume) Chambers, Jermy & Crabbe; Brit. Fern Gaz. 10: 176 (1971)
Distr.: peninsular Malaysia (Kedah, Penang Isl., Kelantan, Pahan, Johor), Java, Sumatra, Thailand
Remarks: nummulariifolia agg.
= Lycopodium nummulariifolium Blume; Enum. pl. Javae 2: 263 (1828)
= Phlegmariurus nummulariifolius (Blume) Ching; Acta Bot. Yunnanica 3: 298 (1981)
= Urostachys nummulariifolius (Blume) Herter; Philipp. Sci. 22: 182 (1923)
= Lycopodium rotundifolium Roxb.; Calcutta J. Nat. Hist. 4: 473 (1844)
Huperzia nutans (Brackenr.) Rothm.; Feddes Repert. 54: 62 (1944)
Distr. [complete]: Hawaii (Kauai, Oahu)
Remarks: squarrosa agg.
Conservation status: IUCN E
= Lycopodium nutans Brackenr.; U. S. Expl. exped., Filic. 16: 327, t. 46 (1854)
= Urostachys nutans (Brack.) Herter ex Nessel; Bärlappgewächse 208. 1939
Huperzia nylamensis Ching & S.K. Wu; Acta Botanica Yunnanica 3(3): 305 (1981)
Distr.: Tibet
Remarks: selago agg.
= Phlegmariurus nylamensis (Ching & S.K. Wu) H.S. Kung & L.B. Zhang; Acta Phytotaxonomica Sinica 37(1): 52 (1999)
= Huperzia nylamensis Ching & S.K. Wu; Flora Xizangica 1: 9, t. 2, f. 1-3, t. 1, f. 4-6 (1983) [nom. illeg., superfl.]
Huperzia obovalifolia (Bonap.) comb. ined.
Distr.: Annam
Remarks: obtusifolia agg.
= Lycopodium obovalifolium Bonap.; Notes Pteridol. 14: 190 (1923)
= Urostachys obovalifolius (Bonap.) Nessel; Bärlappgewächse 249 (1939)
Huperzia obscure-denticulata Ching; Acta Bot. Yunnanica 3: 295 (1981)
Distr.: NW-Sichuan
Remarks: selago agg.
Huperzia obtusifolia (P. Beauv.) Rothm.; Feddes Repert. 54: 61 (1944)
Distr.: Mauritius, Reunion
Remarks: obtusifolia agg.
= Lepidotis obtusifolia P. Beauv.; Prodr. aetheogam. 109 (1805)
= Lycopodium obtusifolium (P. Beauv.) Sw.; Syn. fil. 177 (1806)
= Urostachys obtusifolius (P. Beauv.) Herter ex Nessel; Bärlappgewächse 190 (1939)
Huperzia ocanana (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Colombia
Remarks: saururus agg.
= Huperzia moritzii (Nessel) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Urostachys moritzii Herter ex Nessel; [Bärlappgewächse 70. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 1601 t. 9, f. 29 (1940)]
= Lycopodium ocananum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 45 (1909)
= Urostachys ocananus (Herter) Herter ex Nessel; Bärlappgewächse 101 (1939)
= Huperzia arthurii (Herter) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Urostachys arthurii Maxon ex Herter; Revista Sudamer. Bot. 10: 114 (1953 [1954])
= Lycopodium arthurii (Herter) C. Morton; Amer. Fern J. 54: 72 (1964)
= Huperzia trachyloma (Herter) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 157 (1988)
= Lycopodium trachyloma (Herter) C. Morton; Amer. Fern J. 54: 73 (1964)
= Urostachys trachyloma Maxon ex Herter; Revista Sudamer. Bot. 10: 113 (1953 [1954])
Huperzia occidentalis (Clute) Kartesz & Gandhi; Phytologia 70: 201 (1991)
Distr.: SW-Yukon, W-British Columbia, coastal, to, Washington, isolated Alberta, Montana, N-Idaho
= Huperzia occidentalis (Clute) Beitel; Amer. Fern Journal 82(2): 46 (1992) [nom. illeg.]
= Lycopodium lucidulum Michaux f. occidentale Clute; Fern Bull. 11: 13 (1903)
= Lycopodium lucidulum var. occidentale (Clute) L. R. Wilson; Rhodora 34: 170 (1932)
= Lycopodium selago var. occidentale (Clute) B. Boivin; Le Naturaliste Canadien 93: 359 (1966)
Huperzia oceaniana (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Vanuatu
Remarks: myrtifolia agg.
= Lycopodium oceanianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 52 (1909)
= Urostachys oceanianus (Herter) Herter ex Nessel; Bärlappgewächse 237 (1939)
Huperzia oltmannsii (Nessel) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Tahiti
Remarks: phlegmaria agg.
= Urostachys oltmannsii Herter ex Nessel; [Bärlappgewächse 231. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 168, t. 14, f. 71 (1940)]
Huperzia ophioglossoides (Lam.) Rothm.; Feddes Repert. 54: 62 (1944)
Distr.: Transvaal, Zimbabwe, Mozambique, Malawi, trop. Africa, Bioko Isl. (Fernando Poo), Madagascar, Reunion, Comores, Cameroun, Uganda, Kenya, Tanzania, Mauritius, Sudan, Sao Tome
Remarks: ophioglossoides agg.
= Lepidotis longifolia P. Beauv.; Prodr. aethéogam. 109 (1805)
= Lycopodium longifolium (Beauv.) Sw.; Syn. Fil. 177 (1806)
= Lycopodium ophioglossoides Lam.; Encyclopédie Méthodique, Botanique 3: 646 (1789)
= Urostachys ophioglossoides (Lam.) Herter ex Nessel; Die Bärlappgewächse 238 (1939)
Huperzia orgaosana (Herter ex Nessel) comb. ined.
Distr.: Brazil
Remarks: reflexa agg.
= Urostachys orgaosanus Herter ex Nessel; Arch. Bot. Sao Paulo 1: 380 (1927)
Huperzia orizabae (L. Underw. & F. Lloyd) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr. [complete]: S-Mexico, Chiapas, Guatemala, Honduras, El Salvador
Remarks: linifolia agg.
= Lycopodium orizabae Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33: 110 (1906)
= Urostachys orizabae (Underw. & Lloyd) Herter ex Nessel; Die Bärlappgewächse 175 (1939)
Huperzia ovatifolia Ching; Acta Bot. Yunnanica 3: 298 (1981)
Distr.: SE-Yunnan
Remarks: hamiltonii agg.
= Phlegmariurus ovatifolius (Ching) W.M. Chu; Acta Phytotaxonomica Sinica 37(1): 52 (1999)
Huperzia pachyphylla (Herter) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Madagascar
Remarks: obtusifolia agg.
= Huperzia obtusifolia var. pachyphylla (Kuhn ex Herter) C. Chr.; Catalogue des Plantes de Madagascar, Pterid. 69 (1931)
= Lycopodium pachyphyllum Kuhn ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 51. 1909 [Kuhn ex E. Pritzel, in Engler & Prantl., Natürl. Pflanzenfam. l(4): 599 (1901 [nom. inval.])]
= Urostachys pachyphyllus (Herter) Herter ex Nessel; Bärlappgewächse 190 (1939)
Huperzia parksii (Copel.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Fiji
Remarks: verticillata agg.
= Lycopodium parksii Copel.; Bernice P. Bishop. Mus. Bull. 59: 8, t. 2 (1929)
= Urostachys parksii (Copel.) Herter ex Nessel; Bärlappgewächse 124 (1939)
Huperzia parvifolia (Nessel) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
Distr.: Brazil
Remarks: reflexa agg.
= Urostachys parvifolius Raddi ex Nessel; Arch. Bot. Sao Paulo 1: 395, t. 10, f. 2 (1927)
Huperzia patentissima (Alderw.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: New Guinea
Remarks: phlegmaria agg.
= Lycopodium patentissimum Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 24: 4 (1917)
= Urostachys patentissimus (Alderw.) Herter ex Nessel; Bärlappgewächse 226 (1939)
Huperzia pearcei (Baker) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Bolivia
Remarks: reflexa agg.
= Lycopodium pearcei Baker; Handbook of the Fern-Allies 14 (1887)
= Urostachys pearcei (Baker) Herter ex Nessel; Bärlappgewächse 73 (1939)
Huperzia pecten (Baker) Tard.; Adansonia 10: 18 (1970)
Distr.: Madagascar
Remarks: pecten agg.
= Lycopodium pecten Baker; J. Linn. Soc. Bot. 15: 421 (1876)
= Urostachys pecten (Baker) Herter ex Nessel; Bärlappgewächse 67 (1939)
Huperzia perreriana Tard.; Adansonia 10: 20, t. 2, fig. 14 (1970)
Distr.: Madagascar
Remarks: phlegmaria agg.
Huperzia petiolata (C. B. Clarke) Dixit; J. Bombay Nat. Hist. Soc. 77 (3): 541 (1981)
Distr.: India (Meghalaya, Khasia), Yunnan
Remarks: hamiltonii agg.
= Lycopodium hamiltonii Sprengel var. petiolatum C. B. Clarke; Trans. Linn. Soc. London, ser. 2; Bot. 1: 590 (1880 (Aug.))
= Lycopodium petiolatum (C. B. Clarke) Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 36. 1909, by reference to Baker (Handb. fern-allies 10 (1887))
= Lycopodium petiolatum (C.B. Clarke) Christ; Bulletin de l'Académie Internationale de Géographie, Botanique 11: 271., H. Christ? (1902)
= Phlegmariurus hamiltonii var. petiolatus (C.B. Clarke) Ching; Acta Botanica Yunnanica 4(2): 126-127 (1982)
= Phlegmariurus petiolatus (C.B. Clarke) H.S. Kung & L.B. Zhang; Acta Phytotaxonomica Sinica 37(1): 45-47 (1999)
= Urostachys hamiltonii var. petiolatus (C.B. Clarke) Herter ex Nessel; Die Bärlappgewächse 68 (1939)
= Urostachys petiolatus (C.B. Clarke) Herter ex Nessel; Die Bärlappgewächse 196, see Engl. Bot Jahrb. 34, 98 (1909). to check basi, Nessel lists as Baker, Handb (1887)
Huperzia petrovii Siplivinsky; Novosti Sist. Vyssh. Rast. 10: 346 (1973)
Distr.: Russia, W-Siberia, C-Siberia, E-Siberia / Amur / Ussuri
Remarks: selago agg.
= Huperzia appressa auct.; [p. p.]
= Lycopodium appressum (Desv.) V. Petrov; Flora Iakutiae 37 (1930) excl. basionymo & pl. arct. [non Lloyd & Underw. p. p.]
= Lycopodium selago L. ssp. appressum (Desv.) Hult.; [p. p.]
Huperzia pflanzii (Nessel) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100) 156 (1988)
Distr. [complete]: Costa Rica, Bolivia
Remarks: brevifolia agg.
= Lycopodium pflanzii (Nessel) Lellinger; Proc. Biol. Soc. Wash. 89 (61): 717 (1977)
= Urostachys pflanzii Nessel; Repert. Spec. Nov. Regn. Veg. 36(12-15): 181, t. 171 (1934)
Huperzia phlegmaria (L.) Rothm.; Feddes Repert. 54: 62 (1944)
Distr.: India (West Bengal, Sikkim, Assam, Arunachal Pradesh, Kerala, Andamans, Nicobars, peninsular Malaysia, Borneo (Mt. Kinabalu, etc.), Malesia, trop. Asia, Taiwan, Japan, Queensland, China, trop. Africa, Principe Isl., Sao Tome, Uganda, Tanzania, Malawi, Christmas Isl. (Austr.), Sumatra, Thailand
Remarks: phlegmaria agg.
Conservation status: IUCN E (North Marianas)
= Lepidotis phlegmaria (L.) P. Beauv.; Prodr. aethéogam. 109 (1805)
= Lycopodium filiforme Roxb.; Calcutta J. Nat. Hist. 4: 473. 1844 [non Sw. 1800]
= Lycopodium phlegmaria L.; Sp. pl. 1101 (1753)
= Phlegmariurus phlegmaria (L.) Holub; Preslia 36(1): 21 (1964)
= Phlegmariurus phlegmaria (L.) Sen & Sen; Fern Gaz. 11 (6): 421. f. 4 a (1978)
= Urostachys phlegmaria (L.) Herter ex Nessel; Die Bärlappgewächse 215 (1939)


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          var. divaricatum Blume; Enum. pl. Javae 262 (1828)
Distr.: Java
= Lycopodium divergens Alderw.; Malayan fern allies 45 (1915)
          var. tardieuae (Herter) Tardieu; Adansonia ser. 2 10: 18 (1970)
Distr.: Madagascar
= Urostachys tardieuae Herter; Notulae Systematicae. Herbier du Museum de Paris 15: 355 (1958)
= Huperzia tardieuae (Herter) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Huperzia phlegmarioides (Gaudich.) Rothm.; Feddes Repert. 54: 62 (1944)
Distr.: Queensland, New Caledonia, Moluccas, Fiji
Remarks: phlegmarioides agg.
= Lycopodium (Phlegmaria) phlegmarioides Gaudich.; Voy. Uranie 1: 281, t. 23 (1828 (16 Aug))
= Urostachys phlegmarioides (Gaudich.) Herter ex Nessel; Bärlappgewächse 247 (1939)
= Huperzia kandavuensis (Nessel) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
= Urostachys kandavuensis Nessel; Repert. Spec. Nov. Regni Veg. 39: 67, t. 192a (1935)
= Huperzia pseudophlegmaria (Kuhn) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium pseudophlegmaria Kuhn; Forschungsr. Gazelle 4: 16 (1889)
= Urostachys pseudophlegmaria (Kuhn) Herter ex Nessel; Bärlappgewächse 237 (1939)
Huperzia phylicifolia (Desv. ex Poir.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Ecuador, Chile, Galapagos Remarks: myrsinites agg.
= Lycopodium congestifolium Spring; Bull. Acad. Roy. Sci. Belg. 8: 518 (1841)
= Lycopodium phylicifolium Desv. ex Poiret; Encycl. suppl. 3: 546 (1813 [1814] ("phylicaefolium"))
= Lycopodium subulatum var. pastoense Baker; Handbook of the Fern-Allies 21 (1887)
= Urostachys phylicifolius (Poiret) Herter ex Nessel; Bärlappgewächse 246 (1939)
Huperzia phyllantha (Hook. & Arn.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: India (Uttar Pradesh, Arunachal Pradesh, West Bengal, Sikkim, Assam, Manipur, Meghalaya, Kerala, Tamil Nadu), Arctic and Alpine Zones of both hemispheres, mountains of tropical Asia, Africa, America, Cape Prov., Mascarenes, Polynesia, peninsular Malaysia (Penang Isl., Kelantan, Perak, Negeri Sembilan, Singapore), Borneo (Mt. Kinabalu, etc.), Hawaii (Kauai, Oahu, Molokai, Maui, Lanai, Hawaii Isl.)
Remarks: phyllantha agg.
= Lycopodium phyllanthum Hook. & Arn.; Bot. Beechey Voy. 102 (1832 (Oct-Nov.))
= Phlegmariurus phyllanthus (Hook. & Arn.) Dixit; J. Bombay Nat. Hist. Soc. 77 (3): 541 (1980 [1981])
= Urostachys phyllanthum (Hook. & Arn.) Herter; Philipp. J. Sci. 22: 181. 1923
= Huperzia parvifolia (Raddi ex Nessel) Rolleri & Deferrari; Notas del Museo de la Plata, Botánica 21(100): 156 (1988)
= Huperzia parvifolia (Raddi) Rothm.; Feddes Repert. 54: 60 (1944) [nom. inval.]
= Lycopodium parvifolium "Raddi" (Herter 1909a: 34, Rolleri 1981: 81) [nom. inval.]
= Lycopodium pachystachyon Spring; Bull. Acad. Roy. Sci. Belg. 8: 518. 1841 [non Lycopodium pachystachyum Desv. ex Poiret (1814)]
= Lycopodium parvifolium Raddi ex Rolleri; Notas Mus. La Plata, Bot. 13(71): 81 (1981), nom. inval.
= Urostachys pachystachyon (Spring) Herter ex Nessel; Bärlappgewächse 211 (1939)
= Urostachys pachystachyon (Spring) Rothm.; Feddes Repert. 54: 62 (1944)
= Urostachys parvifolius Raddi ex Nessel; Archivos de Botânica do São Paulo 1: 395, t. 10, f. 2 (1927)
Huperzia picardae (H. Christ ex Krug) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Haiti
Remarks: taxifolia agg.
= Lycopodium picardae Christ ex Krug; Urban; Bot. Jahrb. Syst. 24: 148 (1897)
= Urostachys picardae (Krug) Herter; Repert. Spec. Nov. Regni Veg. 19: 166 (1923)
Huperzia pichiana Tard.; Adansonia 10: 21, t. 2, fig. 5-8 (1970)
Distr.: Madagascar
Remarks: verticillata agg.
Huperzia pilgeriana (Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distr.: Peru
Remarks: saururus agg.
= Urostachys pilgerianus Herter ex Nessel; Revista Sudamericana de Botánica 6(5-6): 161, t. 9, f. 32., as Urostachys "Pilgerianus" (1940)
Huperzia pinifolia Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247 (1874)
[nom. nov. for Lycopodium pinifolium Blume, non Kaulf.]
Distr.: Borneo (Mt. Kinabalu, etc.)
Remarks: hippuris agg.
= Lycopodium pinifolium Blume; Enum. pl. javae 2: 264. 1828 [non Kaulf (1824)]
= Urostachys pinifolius (Blume) Herter; Bot. Arch. 3: 16 (1923)
Huperzia piscium (Herter) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distr.: Thailand
= Lycopodium piscium (Herter) Tag. & Iwatsuki; Acta Phytotax. Geobot. 22: 103 (1967)
= Urostachys piscium Herter; Index Lyc. 75 (1949)
Huperzia pithyoides (Schldl. & Cham.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr. [complete]: Costa Rica, S-Mexico, Chiapas, Guatemala, Honduras, El Salvador, Nicaragua, Colombia, Venezuela, Greater Antilles
Remarks: wilsonii agg.
= Lycopodium gigas Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 50 (1909)
= Lycopodium mandiocanum Raddi var. mexicanum Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 45 (1842)
= Lycopodium pithyoides Schlecht. & Cham.; Linnaea 5: 623 (1830)
= Urostachys gigas (Herter) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 164 (1923)
= Urostachys pithyoides (Schlecht. & Cham.) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 164 (1923)
Huperzia pittieri (H. Christ) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr. [complete]: Isla del Coco
Remarks: linifolia agg.
= Lycopodium pittieri H. Christ; Bulletin de la Société Botanique de Genève 1(5): 236 (1909)
= Urostachys pittieri (H. Christ) Herter; Estudios Botánicos en la Región Uruguaya 20: 75 (1949)
Huperzia polaris (Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Antipodes Isl.
Remarks: varia agg.
= Urostachys varius (R. Br.) Nessel var. polaris Herter ex Nessel; Revista Sudamer. Bot. 6: 165, t. 12, fig. 59. 1940 [non Lycopodium varium R. Br. var. polaris Kirk (18849)
= Urostachys polaris (Nessel) Herter; Index Lyc. 76 (1949)
Huperzia polycarpos (Kunze) B. Øllg.; Opera Bot. 92: 169 (1987)
Distr. [complete]: Costa Rica, Panama, Colombia, Ecuador, Peru
Remarks: dichotoma agg.
= Lycopodium cuatrecasasii (Herter) C. V. Morton; Amer. Fern Journal 54: 72 (1964)
= Lycopodium polycarpos Kunze; Linnaea 9: 5., orth. var (1834)
= Urostachys cuatrecasasii Herter; Revista Sudamericana de Botánica 10: 123 (1953)
= Urostachys polycarpos (Kunze) Herter; Estudios Botánicos en la Región Uruguaya 20: 76 (1949)
Huperzia polyclada (Sodiro) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
Distr.: Ecuador
Remarks: saururus agg.
= Lycopodium polycladum Sodiro; Crypt. vasc. quit. 561. 1893 [Anales Univ. Quito 12 (81): 316 (1895])
= Urostachys polycladus (Sodiro) Herter ex Nessel; Bärlappgewächse 73 (1939)
Huperzia polydactyla B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 64, f. 11B (1988)
Distr. [complete]: Ecuador
Remarks: brevifolia agg.
Huperzia polylepidetorum B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 42, f. 6B (1988)
Distr. [complete]: Ecuador
Remarks: saururus agg.
Huperzia pomperyana (Herter) comb. ined.
Distr.: New Caledonia
Remarks: myrtifolia agg.
= Lycopodium balansae var. pomperyanum Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 52 (1909)
= Urostachys pomperyanus (Herter) Herter; Index Lyc. 76 (1949)
Huperzia porophila (F.W. Lloyd & L. Underwood) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Alabama, Illinois, Indiana, Iowa, Kentucky, Minnesota, Missouri, Ohio, Pennsylvania, Tennessee, Virginia, West Virginia, Wisconsin, C-Appalachians, W-Tennessee, W-Kentucky, &, around, isolated Iowa/Minnesota/Wisconsin
Trivial names: Rock Club-Moss
Remarks: selago agg.
= Huperzia selago (L.) Bernh. ex Schrank & Mart. ssp. lucidula var. porophila (Lloyd & Underw.) Love & Love
= Lycopodium lucidulum Michx. var. porophilum (Lloyd & Underwood) Clute; The Fern Allies 262 (1905)
= Lycopodium porophilum Lloyd & Underw.; Bull. Torrey Bot. Club 27(4): 150 (1900)
= Lycopodium selago L. var. porophila (Lloyd & Underwood) Clute
= Urostachys lucidulus (Michx.) Herter var. porophilus (Lloyd & Underwood) Nessel; [non Lam.]
= Urostachys porophilus (Lloyd & Underwood) Herter
Huperzia pringlei (Underw. & F.E. Lloyd) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr. [complete]: S-Mexico, Chiapas, Guatemala, Honduras, El Salvador
Remarks: verticillata agg.
= Lycopodium fontinaloides var. mexicanum Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 49 (1842)
= Lycopodium pringlei Underw. & F.E. Lloyd; Bulletin of the Torrey Botanical Club 33: 109 (1906)
= Urostachys pringlei (Underw. & Lloyd) Herter ex Nessel; Die Bärlappgewächse 131 (1939)
Huperzia prolifera (Blume) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
Distr.: Java, Queensland
Remarks: hippuris agg.
= Lycopodium proliferum Blume; Enum. pl. Javae 2: 265 (1828)
= Urostachys prolifer (Blume) Herter; Index Lyc. 77 (1949)
Huperzia pruinosa (Herter) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Peru
Remarks: phlegmaria agg.
Conservation status: IUCN R
= Lycopodium pruinosum Hieron. et Herter ex Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 52 (1909)
= Urostachys pruinosus (Herter) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 420 (1927)
Huperzia pseudovaria (Brownlie) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distr.: New Caledonia
Remarks: myrtifolia agg.
Conservation status: IUCN R
= Lycopodium pseudovarium Brownlie; Ptéridophytes, in Aubréville, Fl. Nouv.-Calédonie 3.- 23, t. 1, f. 13-15 (1969)
Huperzia pulcherrima (Hook.& Grev.) Pichi-Serm.; Webbia 24: 719 (1970)
Distr.: India (NW-Himalayas, E-India), Nepal, Bhutan, China, Taiwan, Japan
Remarks: hippuris agg.
= Huperzia pulcherrima (Wall. ex Hook. & Grev.) Sen & Sen; Fern Gaz. 11 (6): 419. f 2 i-r (1978)
= Huperzia setacea Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
= Huperzia taiwanensis (Ching) Holub; Folia Geobot. Phytotax. 20(1): 77 (1985)
= Lycopodium pulcherrimum Wall. ex Hook. & Grev.; Icones Filicum 1(2): t. 38 (1827)
= Lycopodium setaceum Buch.-Ham. ex D. Don; Prodromus Florae Nepalensis 18 (1825) [nom. illeg., non Lycopodium setaceum Lam. (1789)]
= Phlegmariurus pulcherrimus (Hook. & Grev.) Löve & Löve; Taxon 26: 324 (1977)
= Phlegmariurus taiwanensis Ching; Acta Bot. Yunnanica 4: 124 (1982 [nom. superfl. for Lycopodium pulcherrimum Hook. & Grev. by reference to Hayata (1914: 132))]
= Urostachys amphorae Herter; Index Lyc. 51 (1949 [nom. superfl. for Urostachys pulcherrimus (Hook. & Grev.) Nessel)]
= Urostachys pulcherrimus (Hook. & Grev.) Herter ex Nessel; Bärlappgewächse 72 (1939)
= Urostachys setaceus (Hamilt. ex D. Don) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 408. 1927 [Bärlappgewächse 141 (19399)
= Urostachys setaceus var. brasiliensis Herter ex Nessel; Arquivos de Botânica do Estado de São Paulo 1: 408., see MerMert., Monographie, Lyc. 1:43 (1842) for basionym (1927)
Huperzia pungentifolia (Silveira) B. Øllg.; Opera Bot. 92: 169 (1987)
Distr.: Brazil (Minas Gerais, Sta. Catarina), Haiti
Remarks: brongniartii agg.
= Urostachys hennebergorum Nessel; Repert. Sp. Nov. Regni Veg. 48: 169, t. 316, f. 1 (1940)
= Lycopodium ouropretanum Christ; Schwacke, Pl. Nov. Mineiras 2: 39, t. 4. 1900 [sens. Bull. Herb. Boissier, ser. 2, 2: 702 (1902])
= Lycopodium pungentifolium Silveira; Bol. Comm. Geogr. Geol. Minas Geraes 2, 5: 119, t. 4 (1898)
= Urostachys pungentifolius (Silveira) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 394 (1927)
Huperzia quadrifariata (Bory) Rothm.; Feddes Repert. 54: 60 (1944)
Distr.: Brazil (Sta. Catarina, Rio de Janeiro)
Remarks: quadrifariata agg.
= Lycopodium quadrangulare Spring, in. C. Martius; Fl. Bras. l(2): 112-113, t. 5, f. 1 (1840)
= Urostachys quadrangularis (Spring) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 404. 1927
= Lycopodium quadrifariatum Bory; Duperrey, Voy. monde; Bot. 1: 245 (1829 (Aug.))
= Urostachys quadrifariatus (Bory) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 405 (1927)
= Huperzia aschersonii (Herter) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Lycopodium aschersonii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 53 (1909)
= Urostachys aschersonii (Herter) Herter ex Nessel; Arch. Bot. Sao Paulo 1. 422 (1927)
Huperzia quasipolytrichoides (Hayata) Ching; Acta Botanica Yunnanica 3(3): 299 (1981)
Distr.: Taiwan
Remarks: selago agg.
= Huperzia reflexo-integra (Hayata) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium quasipolytrichoides Hayata; Icones plantarum formosanarum nec non & contributiones ad floram formosanam. 5: 252-254, f. 89 (1915)
= Lycopodium reflexo-integrum Hayata; Icones plantarum formosanarum nec non & contributiones ad floram formosanam. 5: 254-255, f. 90 (1915)
= Urostachys quasipolytrichoides (Hayata) Herter ex Nessel; Die Bärlappgewächse 64-65, Abb. 8, f. 37 (1939)
= Urostachys reflexo-integer (Hayata) Herter ex Nessel; Bärlappgewächse 64 (1939)
          var. rectifolia (J.F. Cheng) H.S. Kung & L.B. Zhang;
Acta Phytotaxonomica Sinica 36(6): 528 (1998)
Distr.: Distribution unknown
= Huperzia whangshanensis var. rectifolia J.F. Cheng; Fl. Jiangxi 1: 27, 505, f. 12 (1993)
Huperzia recurvifolia Rolleri; Notas del Museo de la Plata, Botánica 21(10, f. 1-33): 210 (1989)
Distr. [complete]: Venezuela, Brazil (Bahia), British Guyana
Huperzia reflexa (Lam.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
Distr. [complete]: Puerto Rico, Costa Rica, Galapagos Isl., Greater Antilles, Mexico, Guatemala, Honduras, El Salvador, Paraguay, &, Bolivia, Cuba, Nicaragua, Colombia, Venezuela, Ecuador, Galapagos, Peru, Brazil, Greater Antilles, Martinique, Guadeloupe, Dominica, St. Vincent, British Guyana (Mt. Roraima)
Remarks: reflexa agg.
= Huperzia bifida (Humb. & Bonpl. ex Willd.) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
= Huperzia densifolia (Baker) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
= Huperzia mexiae (Copel.) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
= Huperzia reflexa (Lam.) Rothm.; Repert. Spec. Nov. Regn. Veg. 54: 60., hom. illeg. superfl (1944)
= Huperzia reflexa var. bifida (Humb. & Bonpl. ex Willd.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
= Lycopodium bifidum Humb. & Bonpl. ex Willd.; Sp. Pl.. Editio quarta 5: 53 (1810)
= Lycopodium densifolium (Baker) Underw. & F.E. Lloyd; Bulletin of the Torrey Botanical Club 33: 106 (1906)
= Lycopodium mexiae Copel.; Univ. California Publ. Bot. 19: 294, t. 47 (1941)
= Lycopodium reflexum Lam.; Encyclopédie Méthodique, Botanique 3: 653 (1792)
= Lycopodium reflexum var. caesium Vareschi; Flora de Venezuela 1(1): 31 (1968), nom. inval. (no type indicated)
= Lycopodium reflexum var. densifolium Baker; Handbook of the Fern-Allies 11 (1887)
= Lycopodium reflexum var. majus Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 26 (1842)
= Lycopodium reversum C. Presl; Reliquiae Haenkeanae 1(1): 82 (1825)
= Lycopodium rigidum Sw.; J. Bot. (Schrader) 1800 (2): 114. 1801 [Syn. fil. 176, 400. 1806] [non J. Gmelin (1791)]
= Lycopodium squarrosum Sw.; Prodr. 137. 1788 [non G. Forster 1786, nec Larn (1792)]
= Lycopodium stellae-polaris (Herter) C.V. Morton; Amer. Fern Journal 54: 72 (1964)
= Plananthus reflexus (Lam.) P. Beauv.; Prodr. Aetheog. 100 (1805)
= Plananthus squarrosus (Sw., Lam., J. Gmelin) P. Beauv.; Prod. aethéogam. 112 (1805) [nom. confus., combination based on different basionyms]
= Urostachys bifidus (Humb. & Bonpl. ex Willd.) Herter ex Nessel; Die Bärlappgewächse 110, t. 24, f. 4 (1939)
= Urostachys densifolius (Baker) Herter; Index Lyc. 58. 1949
= Urostachys jergii Nessel; Repert. Spec. Nov. Regn. Veg. 39: 70, t. 194a (1935)
= Urostachys reflexus (Lam.) Herter; Beihefte zum Botanischen Centralblatt 39: 249 (1922)
= Urostachys reflexus var. densifolius (Baker) Herter ex Nessel & Hoehne; Archivos de Botânica do São Paulo 1(4): 394 (1927)
= Urostachys reversus (C. Presl) Herter; Estudios Botánicos en la Región Uruguaya 20: 78 (1949), Alt. title: Index Lycopodiarum 78 (1949)
= Urostachys stellae-polaris Herter; Revista Sudamericana de Botánica 10: 121 (1954)
          var. minor (Spring) B. Øllg.; Flora of Ecuador 33: 26 (1988)
Distr. [complete]: Honduras, Costa Rica, Ecuador, Peru, Bolivia, Brazil, Trinidad
= Lycopodium brutum Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 47 (1909)
= Lycopodium reflexum var. minus Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 26 (1842)
= Urostachys brutus (Herter) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 162 (1923)
= Urostachys reflexus (Lam.) Herter var. brutus (Herter) Herter ex Nessel; Die Bärlappgewächse 116 (1939)
= Urostachys reflexus var. minor (Spring) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 393 (1927)
Huperzia regnellii (Maxon) B. Øllg. & Wind.; Bradea 5: 16 (1987)
Distr.: Brazil (Minas Gerais)
Remarks: brongniartii agg.
Conservation status: IUCN R
= Lycopodium regnellii Maxon; Contr. U. S. Natl. Herb. 17 (4): 424, t. 23 (1914)
= Urostachys regnellii (Maxon) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 381 (1927)
Huperzia ribourtii (Herter) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Tahiti
Remarks: obtusifolia agg.
= Lycopodium ribourtii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 53 (1909)
= Urostachys ribourtii (Herter) Herter ex Nessel; Bärlappgewächse 250 (1939)
Huperzia rigida (J. Gmelin) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr. [complete]: Hispaniola, Jamaica, Guadeloupe, Martinique, St. Vincent
Remarks: reflexa agg.
= Lycopodium rigidum J. Gmelin; Syst. nat. II [Syst. Veg.1 (2): 128] (1791)
= Plananthus rigidus (J. Gmelin) P. Beauv.; Prod. aethéogam. 111 (1805)
= Urostachys rigidus (J. Gmelin) Herter; Repert. Spec. Nov. Regni Veg. 19: 163 (1923)
Huperzia rimbachii (Sodiro) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Ecuador
Remarks: saururus agg.
= Lycopodium rimbachii Sodiro; Crypt. vasc. quit. 566. 1893 [Anales Univ. Quito 12 (81): 321 (1895])
= Urostachys rimbachii (Sodiro) Herter ex Nessel; Bärlappgewächse 102 (1939)
Huperzia ringshausenii (Nessel) comb. ined.
Distr.: Colombia
Remarks: reflexa agg.
= Urostachys ringshausenii Nessel; Repert. Spec. Nov. Regni Veg. 39: 68, t. 193a (1935)
Huperzia riobambensis (Herter) B. Øllg.; Opera Bot. 92: 169 (1987)
Distr. [complete]: Colombia
Remarks: reflexa agg.
= Urostachys affinis (Grev. & Hook.) Nessel var. riobambensis Nessel; [Bärlappgewächse 98 (1939] (Revista Sudamericana de Botánica 6(5-6): 162, t. 10, f. 38 (1940) [nom. inval.]
= Urostachys riobambensis (Nessel) Herter; Index Lyc. 79 (1949)
= Lycopodium castoris (Herter) C. Morton; Amer. Fern J. 54: 72 (1964)
= Urostachys castoris Herter; Revista Sudamer. Bot. 10: 111 (1953 [1954])
Huperzia robusta (Klotzsch) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr. [complete]: Venezuela
Remarks: phlegmaria agg.
= Huperzia phelpsii (Vareschi) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium phelpsii Vareschi; Acta Botanica Venezuelica 1: 84 (1966)
= Lycopodium robustum Klotzsch; Linnaea 18: 518 (1844)
= Urostachys robustus (Klotzsch) Herter ex Nessel; Bärlappgewächse 212 (1939)
Huperzia rosenstockiana (Herter) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Ecuador
Remarks: taxifolia agg.
= Lycopodium cassandrae (Herter) C. Morton; Amer. Fern J. 54: 72 (1964)
= Lycopodium rosenstockianum Herter; Hedwigia 49: 90 (1909)
= Urostachys cassandrae Herter; Revista Sudamer. Bot. 10: 116 (1953 [1954])
= Urostachys rosenstockianus (Herter) Herter ex Nessel; Bärlappgewächse 174 (1939)
Huperzia rostrifolia (Silveira) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Brazil (Minas Gerais)
Remarks: reflexa agg.
= Lycopodium rostrifolium Silveira; Bol. Comm. Geogr. Geol. Minas Geraes 2, 5: 118, t. 2 (1898)
= Urostachys rostrifolius (Silveira) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 379 (1927)
Huperzia rubicaulis S.K. Wu & X. Cheng; Acta Phytotaxonomica Sinica 23(5): 400, f. 1 (1985)
Distr.: Qinghai, Tibet, Yunnan
Remarks: selago agg.
Huperzia rubra (Cham.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247 (1874)
Distr.: Brazil
Remarks: brongniartii agg.
= Lycopodium rubrum Cham.; Linnaea 8: 389 (1833)
= Urostachys ruber (Cham.) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 381 (1927)
Huperzia rubrica (C. Chr.) Tardieu; Adansonia ser. 2 10: 18 (1970)
Distr.: Madagascar
Remarks: ophioglossoides agg.
= Lycopodium ophioglossoides Lam. var. rubricaule C. Chr.; Dansk Bot. Ark. 7: 192, t. 77, f. 9-14 (1932)
= Urostachys rubricus Herter; Revista Sudamer. Bot. 8: 25 (1949)
Huperzia rubricaulis (Alderw.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: New Guinea
Remarks: selago agg.
= Lycopodium rubricaule Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 24: 3 (1917)
= Urostachys rubricaulis (Alderw.) Herter ex Nessel; Bärlappgewächse 44 (1939)
Huperzia rufescens (Hook.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1874)
Distr.: Ecuador
Remarks: brevifolia agg.
= Lycopodium rufescens Hook.; Icones Plantarum 1: tab 36 (1837)
= Urostachys rufescens (Hook.) Nessel; Die Bärlappgewächse 103 (1939)
Huperzia rupicola (Alderw.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Brazil
Remarks: squarrosa agg.
= Lycopodium rupicolum Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 16: 39 (1914)
= Urostachys rupicolus (Alderw.) Herter ex Nessel; Bärlappgewächse 203 (1939)
Huperzia salvinioides (Herter) Holub; Folia Geobot. Phytotax. 20. 76 (1985)
Distr. [complete]: Philippines, Taiwan, Japan
Remarks: nummulariifolia agg.
= Lycopodium salvinioides (Herter) Tag.; Acta Phytotax. Geobot. 8: 20 (1939)
= Phlegmariurus salvinioides (Herter) Ching; Acta Bot. Yunnanica 4: 122 (1982)
= Urostachys formosanus Herter ex Nessel; [Bärlappgewächse 229. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 167, excl. fig. 1940]
= Urostachys formosanus Herter ex Nessel; Die Bärlappgewächse 229 (1939)
= Urostachys salvinioides Herter; Bot. Arch. 3. 18 (1923)
Huperzia samoana (Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Samoa
Remarks: hippuris agg.
= Urostachys samoanus Herter ex Nessel; [Bärlappgewächse 125, f. 29e 5. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 163 (1940)]
Huperzia sargassifolia (Liebm.) comb. ined.
Distr.: Mexico (Oaxaca)
Remarks: selago agg.
= Lycopodium sargassifolium Liebm.; Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1847: 41 (1847)
Huperzia sarmentosa (Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
Distr.: Ecuador
Remarks: verticillata agg.
= Lycopodium sarmentosum Spring; Mém. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 13 (1849)
= Urostachys sarmentosus (Spring) Herter; Repert. Spec. Nov. Regni veg. 19: 165 (1923)
Huperzia saururoides (Bory & d'Urv.) Rothm.; Feddes Repert. 54: 59 (1951)
Distr.: Falkland Isl.
Remarks: selago agg.
= Lycopodium selago L. var. saururoides Bory & D'Urville ex Baker; Handb. fern-allies 9. 1887 [Bory in D'Urville, Mém. Soc. Linn. Paris 4: 597, pl. 4 (1825 [1826]) [nom. inval.]
= Urostachys saururoides (Bory et D'Urville ex Baker) Herter ex Nessel; Bärlappgewächse 30 (1939)
Huperzia saururus (Lam.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 249 (1874)
Distr.: Marion Isl., Prince Edward Isl., Réunion, Madagascar, Mauritius, Rodriguez Isl., St. Helena, Nightingale Isl., Tristan da Cunha, Gough Isl., Iles Crozet, Iles Kerguelen, New Amsterdam Isl., Natal, Cape Prov., Lesotho, Zimbabwe, Malawi, Uganda, Kenya, Tanzania, Ethiopia, Cameroun, Argentina (Jujuy, Buenos Aires, Catamarca, Cordoba, Salta, San Luis Prov., Tucuman), Peru, Bolivia, Uruguay, Ascension
Remarks: saururus agg.
Conservation status: IUCN I, Ex/E (Ascension), V (St. Helena), I (Tristan da Cunha)
= Huperzia andina (Rosenst.) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Huperzia axillaris (Roxb.) Rothm.; Feddes Repert. 54: 60 (1944)
= Huperzia sanctae-barbarae (Rolleri) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
= Lycopodium andinum Rosenst.; Repert. Spec. Nov. Regn. Veg. 5: 239 (1908)
= Lycopodium axillare Roxb.; Beatson, Tracts St. Helena 312 (1816)
= Lycopodium elongatum Lagasca ex Sw.; Syn. fil. 175 (1806)
= Lycopodium sanctae-barbarae Rolleri; Darwiniana 16: 129, f. l(D-E)o t. 11 t. 3(B) (1970)
= Lycopodium saururus Lam.; Encycl. 3: 653 (1789 [1792])
= Plananthus saururus (Lam.) P. Beauv.; Prod. aethéogam. 111 (1805)
= Selago indica Dill. (Herter 1949a: 49) [nom. inval. (polynom.)]
= Urostachys andinus (Rosenst.) Nessel; Die Bärlappgewächse 76 (1939)
= Urostachys axillaris (Roxb.) Herter ex Nessel; Bärlappgewächse 82 (1939)
= Urostachys elongatus (Sw.) Herter; Estudios Botánicos en la Región Uruguaya 20: 60 (1949)
= Urostachys saururus (Lam.) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 162 (1923)
Huperzia scabrida B. Øllg.; Harling & Andersson, Fl. Ecuador 33. 60, f. 8A (1988)
Distr. [complete]: Ecuador
Remarks: saururus agg.
Huperzia schlechteri (E. Pritzel) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: New Caledonia
Remarks: squarrosa agg.
= Lycopodium schlechteri E. Pritzel; Schlechter, Bot. Jahrb. Syst. 39: 14 (1906)
= Urostachys schlechteri (E. Pritzel) Herter ex Nessel; Bärlappgewächse 207 (1939)
Huperzia schlimii (Herter) B. Øllg.; Opera Bot. 92: 169 (1987)
Distr.: Colombia
Remarks: saururus agg.
= Lycopodium schlimii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 45 (1909)
= Urostachys schlimii (Herter) Herter ex Nessel; Bärlappgewächse 91 (1939)
Huperzia schmidtchenii (Hieron.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Colombia
Remarks: brevifolia agg.
= Lycopodium schmidtchenii Hieron.; Bot. Jahrb. Syst. 34: 570 (1905)
= Urostachys schmidtchenii (Hieron.) Herter ex Nessel; Bärlappgewächse 103 (1939)
Huperzia selago (L.) Bernhardi ex Schrank & C. F. P. Martius; Hort. Reg. Monac.: 3 (1829)
Trivial names: Tannenbärlapp, Northern Firmoss, Fir Clubmoss
Distr.: Caucasus/Transcaucasus, W-Siberia, C-Siberia, E-Siberia / Amur / Ussuri, C-Asia, Taiga Zone from Yukon to Newfoundland, NE-USA, Albania, Austria, Azores, Belgium, England, Bulgaria, former Czechoslovakia, Denmark, Faroer Isl., Finland, France, Germany, Ireland, Switzerland, Netherlands, Spain, Hungary, Italy, Iceland, former Yugoslavia, Malta, Norway, Poland, Romania, Spitsbergen, Sweden, Baltic States, C-European Russia, E-European Russia, N-European Russia, W-European Russia / Belarus / Ukraine, asiatic Turkey (Anatolia), Victoria, Tasmania, Taiwan, Japan, Yakushima Isl., Sikkim, mountains of, C-Brazil, NW-India, NE-India, Nepal
Remarks: selago agg.
= Huperzia dixitii Mandal & S. Ghosh; [fide Fraser-Jenkins (1997)]
= Huperzia selago (L.) Bernh.; Journal für die Botanik 2: 121-35 (1801)
= Lycopodium abietiforme S. F. Gray; Brit. Pl. 2: 22 (1821)
= Lycopodium densum Lam.; Fl. franc. 1 (Meth. anal.): (33) (1779)
= Lycopodium recurvum Kitaibel ex Willd.; Sp. pl. 5: 50 (1810)
= Lycopodium selago L.; Sp. Pl.: 1102 (1753)
= Mirmau selago (L.) H. P. Fuchs; Verh. Naturf. Ges. Basel 66: 43-44 (1955)
= Plananthus fastigiatus Opiz; Seznam 202. 1852, et Böh. Gew. 114 (1823) (Adelbert MS) [nom. superfl.]
= Plananthus recurvus (Kitaibel ex Willd.) Opiz; Seznam 202 (1852)
= Plananthus selago (L.) P. Beauv.; Prod. aetheogam. 112 (1805)
= Selago vulgaris Dill. (Herter 1949a: 49) [nom. inval.]
= Selago vulgaris Schur; Enum pl. Transsilv. 825 (1866)
= Urostachys selago (L.) Herter; Botanisches Centralblatt 39(2): 249 (1922) [nom. illegit.]
          var. sulcata Ching; Acta Botanica Yunnanica 3(3): 303 (1981)
Distr.: Distribution unknown
          var. patens (Beauv.) Trevisan
Distr.: Japan, North America
= Lycopodium selago ssp. patens (Beauv.) Calder & Taylor
= Lycopodium selago var. patens (Beauv.) Desv.
= Plananthus patens P. Beauv. (1805)
Huperzia sellifolia B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 68, f. 12C (1988)
Distr.: Ecuador, Colombia
Remarks: brevifolia agg.
= Urostachys compactus f. minor Herter; Revista Sudamericana de Botánica 10: 119-120 (1954)
Huperzia sellowiana (Herter) B. Øllg.; Opera Bot. 92: 169 (1987)
Distr.: Brazil
Remarks: brongniartii agg.
= Lycopodium sellowianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 44 (1909)
= Urostachys sellowianus (Herter) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 385 (1927)
Huperzia serpentiformis (Herter) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 156 (1988)
Distr.: Colombia
Remarks: brevifolia agg.
= Lycopodium serpentiforme Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 46 (1909)
= Urostachys serpentiformis (Herter) Herter ex Nessel; Bärlappgewächse 104 (1939)
Huperzia serrata (Thunb. ex Murray) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
Distr.: India (West Bengal, Sikkim, Arunachal Pradesh, Assam, Meghalaya, Manipur, Tamil Nadu), Nepal, Burma, Sri Lanka, China, Japan, Malesia, Fiji, Samoa, tropical America, peninsular Malaysia (Pahan), Borneo (Mt. Kinabalu, etc.), E-Siberia / Amur / Ussuri, Taiwan, Cuba, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Hubei, Hunan, Jiangxi, Jilin, Liaoning, Shaanxi, Sichuan, Japan, Korea, Vietnam, Japan, China, Hawaii (Kauai, Oahu, Molokai, Lanai, Hawaii Isl.), Thailand
Remarks: selago agg.
= Huperzia selago (L.) Bernh. ex Schrank & C. Mart. ssp. serrata (Thunb.) A. & D. Löve
= Huperzia serrata (Thunb.) Rothm.; Feddes Repert. 54 (1): 59 (1944)
= Huperzia serrata f. longipetiolata (Spring) Ching; Acta Botanica Yunnanica 3(3): 294 (1981)
= Huperzia shangsiensis (Chun Y. Yang) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
= Lycopodium serratum Thunb. ex Murray; Systemat Vegetabilium. Editio decima quarta 944 (1784), also Thunb., Fl. Jap. 341, t. 38 (1784)
= Lycopodium serratum var. javanicum (Sw.) Makino; Botanical Magazine 12(131): 13 (1898)
= Lycopodium serratum var. longipetiolatum Spring; Monogr. Lycop. 2: 18 (1850)
= Lycopodium serratum var. myriophyllifolium Hayata; Icones plantarum formosanarum nec non & contributiones ad floram formosanam. 4: 133 (1914)
= Lycopodium serratum var. thunbergii Makino; Botanical Magazine 12(131): 12 (1898)
= Phlegmariurus shangsiensis Chun Y. Yang; Acta Phytotaxonomica Sinica 22(1): 87-88, f. 1 (1984)
= Plananthus serratus (Thunb. ex Murray) P. Beauv.; Prod. aethéogam. 100, 111 (1805)
= Urostachys javanicus (Sw.) Herter; Index Lyc. 66 (1949)
= Urostachys myriophyllifolius (Hayata) Herter; Estudios Botánicos en la Región Uruguaya 20: 71 (1949)
= Urostachys serratus (Thunb. ex Murray) Herter; Bot. Arch. 3: 13 (1923)
= Urostachys serratus (Thunb. ex Murray) Herter var. sachalinensis Nessel; Repert. Spec. Nov. Regni Veg. 36: 179, t. 171. 1934
= Urostachys sachalinensis (Nessel) Herter; Index Lyc. 80. 1949
          f. intermedia (Nakai) Ching; Acta Botanica Yunnanica 3(3): 294 (1981)
Distr.: China, Japan
= Lycopodium serratum f. intermedium Nakai; Botanical Magazine 39(463): 196-197 (1925)
          var. integrifolium (Matsuda) comb ined.
Distr.: China
= Lycopodium serratum Thunb. ex Murray var. integrifolium Matsuda; Bot. Mag. (Tokyo) 25: 124 (1911)
= Lycopodium integrifolium (Matsuda) Matsuda & Nakai ex Nakai; Bot. Mag. (Tokyo) 28: 102. 1914 [non Goldie (1822)]
          var. longipetiolatum Spring
Distr.: SE-Asia, common, Japan
= Lycopodium javanicum Sw.; J. Bot. (Schrader) 1800 (2): 114. 1801 [Syn. Fil. 175, 399 (1806])
          var. myriophyllifolium Hayata
Distr.: Distribution unknown **
Huperzia setifolia (Alderw.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Borneo
Remarks: phlegmaria agg.
= Lycopodium setifolium Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 16: 40 (1914)
= Urostachys setifolius (Alderw.) Herter ex Nessel; Bärlappgewächse 233 (1939)
Huperzia sieberiana (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
Distr. [complete]: Martinique, St. Kitts, St. Vincent, ?Haiti
Remarks: reflexa agg.
= Lycopodium sieberianum Spring; Flora 21: 153 (1838)
= Lycopodium squarrosum Lam.; Encycl. 3. 653. 1792 [non G. Forster 1786, nec Sw. (1788)]
= Urostachys sieberianus (Spring) Herter; Repert. Spec. Nov. Regni Veg. 19: 162 (1923)
Huperzia sieboldii (Miq.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Taiwan, Japan, Korea (Cheju Isl.), China
Remarks: cancellata agg.
= Lycopodium sieboldii Miq.; Ann. Mus. Bot. Lugduno-Batavum 3: 184 (1867)
= Phlegmariurus sieboldii (Miq.) Ching; Acta Bot. Yunnanica 4: 121 (1982)
= Phlegmariurus sieboldii (Miq.) Ching; Flora Xizangica 1: 12 (1983), nom superfl. [nom. illeg., non Lycopodium squarrosum G. Forst (1786)]
= Urostachys sieboldii (Miq.) Herter ex Nessel; Bärlappgewächse 138 (1939)
Huperzia silveirae (Nessel) B. Øllg. & Wind.; Bradea 5: 17 (1987)
Distr.: Brazil (Minas Gerais)
Remarks: brongniartii agg.
= Urostachys silveirae Herter ex Nessel; Arch. Bot. Sao Paulo 1: 386 (1927)
Huperzia simonii (Nessel) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Distr.: Nova Pomerania
Remarks: phlegmaria agg.
= Urostachys simonii Nessel; Revista Sudamer. Bot. 6: 167, excl. fig (1940)
Huperzia sintenisii (Nessel) Holub; Folia Geobet. Phytotax. 20: 76 (1985)
Distr.: Puerto Rico, Hispaniola
Remarks: reflexa agg.
= Lycopodium sintenisii (Herter) Maxon ex C. Chr.; Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16 (2): 91 (1937)
= Urostachys sintenisii Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 161 (1923)
Huperzia societensis (J. Moore) comb. ined.
Distr.: Society Isl. (Raiatea)
= Lycopodium societense J. Moore; Bernice P. Bishop Mus. Bull. 102: 12 (1933)
Huperzia socratis (Herter) Rolleri & Deferrari; Notas del Museo de la Plata,
Botánica 21(100): 157 (1988)
Distr.: Colombia
Remarks: brongniartii agg.
= Lycopodium socratis (Herter) C. Morton; Amer. Fern 1. 54: 72 (1964)
= Urostachys socratis Herter; Revista Sudamer. Bot. 10: 117 (1953 [1954])
Huperzia somai (Hayata) Ching; Acta Bot. Yunnanica 3: 301 (1981)
Distr. [complete]: Taiwan, Japan, Philippines (N-Luzon)
Remarks: selago agg.
= Lycopodium chinense var. somai (Hayata) Masam.; Memoirs of the Faculty of Science and Agriculture Taihoku Imperial University 4: 114 (1934)
= Lycopodium selago var. somai (Hayata) Masam.; Short Fl. Form. 32 (1936)
= Lycopodium somai Hayata; Ic. pl. formos. 5: 255, f. 91 (1915)
= Urostachys somai (Hayata) Herter ex Nessel; Bärlappgewächse 54 (1939)
Huperzia sooiana (Lawalrée) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Zaire
Remarks: phlegmaria agg.
= Phlegmariurus sooianus LawaIrée; Acta Bot. Acad. Sci. Hung. 19: 195, f. 1 (1973)
Huperzia sotae (Rolleri) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Argentina (Jujuy, Salta, Tucuman), Bolivia
Remarks: taxifolia agg.
= Lycopodium sotae Rolleri; Darwiniana 16: 130, f. 1 (A-C), t. 2, t. 3(A) (1970)
Huperzia spongiosa (Rolleri) Rolleri & Deferrari; Notas Mus. La Plata, Bot. 21 (100): 157 (1988)
Distr.: Colombia
Remarks: reflexa agg.
= Lycopodium spongiosum Rolleri; Revista Mus. La Plata (N. S.) Bot. 14 (87): 6, f. 3-4 (1985)
Huperzia sprengeri (Nessel) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Tahiti
Remarks: selago agg.
= Urostachys sprengeri Nessel; Repert. Spec. Nov. Regni Veg. 36: 179, t. 171 (1934)
Huperzia squarrosa (C.Forster) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247 (1874)
Distr.: India (West Bengal, Sikkim, Meghalaya, Assam, Kerala), Nepal, Burma, Bangladesh, Sri Lanka, Malesia, Philippines, Polynesia, Seychelles, Mascarenes, Tahiti, peninsular Malaysia (Kedah, Kelantan, Perak, Pahan), Borneo (Mt. Kinabalu, etc.), China, Taiwan, Queensland, Madagascar, Mauritius, Java, Thailand
Remarks: squarrosa agg.
= Huperzia epiceifolia (Desv. ex Poiret) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
= Huperzia squarrosa (G. Forst.) Rothm.; Repert. Spec. Nov. Regn. Veg. 54: 62 (1944)
= Lycopodium epiceifolium Desv. ex Poiret; Encycl. suppl. 3: 559 (1813 [1814])
= Lycopodium forsteri Poiret; Encycl. suppl. 3: 545 (1813 [1814]) [nom. superfl.]
= Lycopodium gnidioides Blanco; Fl. Filip. 824. 1837, ed. 2.569. 1845, ed. 3 (3): 239 (1879)
= Lycopodium madagascariense Desv.; [nom. inval. (Spring 1842: 34)]
= Lycopodium squarrosum Forst.; Prod. 86 (1786)
= Lycopodium squarrosum G. Forster; Fl. ins. austr. 479 (1786)
= Lycopodium pseudo-squarrosum Pampan.; Bull. R. Soc. Tosc. Orticult. 111, 13: 99, t. 2 (1908)
= Phlegmariurus squarrosus (G. Forster) Löve & Löve; Taxon 26: 324 (1977)
= Plananthus squarrosa (Sw.) P. Beauv.; Prodr. Aetheog. 112 (1805)
= Urostachys epiceaefolius (Desv.) Herter ex Nessel; Die Bärlappgewächse 202 (1939)
= Urostachys epiceaefolius var. acutifolius (Desv.) Herter ex Nessel; Die Bärlappgewächse 292 (1939)
= Urostachys epiceifolius (Poiret) Herter ex Nessel; Bärlappgewächse 202 (1939)
= Urostachys madagascariensis (Desv. ex Nessel) Herter; Estudios Botánicos en la Región Uruguaya 20: 68 (1949), Alt. title: Index Lycopodiarum 68 (1949)
= Urostachys madagascariensis (Nessel) Herter; Index Lyc. 68 (1949)
= Urostachys squarrosus (G. Forst.) Herter; Botanisches Archiv 3: 14 (1923)
= Urostachys squarrosus (G. Forster) Herter var. prolifer (Blume) Nessel f. madagascariensis Nessel
= Urostachys squarrosus var. proliferus (Blume) Herter ex Nessel; Bärlappgewächse 200 (1939)


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          f. hippurioides Alderw.
Distr. [complete]: Queensland
Huperzia staudtii (Nessel) Pichi-Serm.; Webbia 23: 163 (1968)
Distr.: Cameroun
Remarks: phlegmaria agg.
= Lycopodium staudtii (Nessel) C. Adams & Alston; Bull. Brit. Mus. (Nat. Hist.) Bot. 1 (6): 183 (1955)
= Urostachys adolfi-friedericii Herter ex Nessel; Die Bärlappgewächse 226 (1939) [nom. inval., without Latin description]
= Urostachys staudtii Nessel; Repert. Spec. Nov. Regni Veg. 36: 189, t. 175 (1934)
Huperzia stemmermanniae (Medeiros & Wagner) comb. ined.
Distr. [complete]: Hawaii
= Phlegmariurus stemmermanniae Medeiros & Wagner; Amer. Fern Journ. 86(3): 89-97 (1996)
Huperzia suberecta (Lowe) Tard.; Adansonia 10: 20 (1970)
Distr. [complete]: Madeira
Remarks: selago agg.
= Huperzia selago ssp. suberecta (Lowe) Toledo & Vasc.; Bol. Soc. Brot., ser. 2 41: 23 (1967)
= Lycopodium selago ssp. suberectum (Lowe) Romariz; Revista da Facultade de Ciências : 2.a Serie : C : Ciências Naturais 3: 71, t. 1, f. 3 (1953)
= Lycopodium selago var. suberectum (Lowe) C. Chr.; Dansk Botanisk Arkiv 7: 188 (1932)
= Lycopodium suberectum Lowe; Trans. Catnbridge Philos. Soc. 4 (1): 9 (1831)
= Urostachys suberectus (Lowe) Herter ex Nessel; Bärlappgewächse 41 (1939)
Huperzia subfalciformis (Alderw.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: New Guinea
Remarks: phlegmarioides agg. = Lycopodium subfalciforme Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 28: 44 (1918)
= Urostachys subfalciformis (Alderw.) Herter ex Nessel; Bärlappgewächse 227 (1939)
Huperzia subintegra (Hillebr.) Beitel & W.H. Wagner
Distr. [complete]: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)
Huperzia submoniliformis (Alderw.) comb. ined.
Distr.: New Guinea
Remarks: phlegmaria agg.
= Lycopodium horizontale Alderw. var. submoniliforme Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 24: 4 (1917)
= Urostachys submoniliformis (Alderw) Herter; Index Lyc. 84 (1949)
Huperzia subtrifoliata (Brownlie) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distr.: Fiji
Remarks: phlegmaria agg.
= Lycopodium subtrifoliatum Brownlie; Beih. Nova Hedwigia 55: 24 (1977)
Huperzia subtubulosa (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Santo Domingo
Remarks: taxifolia agg.
= Urostachys subtubulosus Herter; Repert. Spec. Nov. Regni Veg. 19: 166 (1923)
Huperzia subulata (Desv. ex Poir.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr. [complete]: Costa Rica, Honduras, Panama, Colombia, Ecuador, Peru, Argentina (Salta, Tucuman), Bolivia
Remarks: myrsinites agg.
= Lycopodium ewanii (Herter) C. V. Morton; Amer. Fern Journal 54: 72 (1964)
= Lycopodium subulatum Desv. ex Poir.; Encyclopedie Methodique. Botanique ... Supplement 3: 544. [1813] (1814)
= Urostachys ewanii Herter; Revista Sudamericana de Botánica 10: ,126-127 (1954)
= Urostachys subulatus (Desv. ex Poir.) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 420 (1927)
Huperzia subulifolia (Hook. & Grev.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
Distr.: India (West Bengal, Sikkim, Meghalaya, Uttar Pradesh), Nepal, Sri Lanka
Remarks: verticillata agg.
= Lycopodium subulifolium Wall. ex Hook. & Grev.; lcon. Fil. t. 49 (1831)
= Urostachys subulifolius (Hook. & Grev.) Herter; Index Lyc. 84 (1949)
Huperzia sumatrana (Alderw.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Sumatra
Remarks: selago agg.
= Lycopodium sumatranum Alderw.; Bull. Jard. Bot. Buitenzorg ser. 2, 28: 44 (1918)
= Urostachys sumatranus (Alderw.) Herter ex Nessel; Bärlappgewächse 66 (1939)
Huperzia sutchueniana (Herter) Ching; Acta Bot. Yunnanica 3: 297 (1981)
Distr.: E-China
Remarks: selago agg.
= Lycopodium serratum var. alpestre H. Christ; Bulletin de l'Académie Internationale de Géographie, Botanique 16(199-201): 141 (1906)
= Lycopodium sutchuenianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909)
= Urostachys sutchuenianus (Herter) Herter ex Nessel; Bärlappgewächse 53 (1939)
Huperzia tafanensis (Nessel) comb. ined.
Distr.: Papua New Guinea
= Urostachys tafanensis Nessel; Repert. Spec. Nov. Regni Veg. 39: 61, t. 189b (1935)
Huperzia tahkuanensis Ching; Acta Botanica Yunnanica 3(3): 297 (1981)
Distr.: China
Remarks: selago agg.
Huperzia takingensis Ching; Acta Botanica Yunnanica 3(3): 300-301 (1981)
Distr.: China
Remarks: selago agg.
Huperzia talamancana B. Øllg.; Novon 3(1): 67, f. 2 (1993)
Distr. [complete]: Costa Rica, Panama
Huperzia talamauana (Alderw.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Sumatra
Remarks: phyllantha agg.
= Lycopodium talamauanum Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2,28: 45 (1918)
= Urostachys talamauanus (Alderw.) Herter ex Nessel; Bärlappgewächse 212 (1939)
Huperzia talpiphila B. Øllg.; Harling & Andersson, Fl. Ecuador 33: 44, f. 9A (1988)
Distr. [complete]: Ecuador
Remarks: saururus agg.
Huperzia tauri (Herter) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distr.: Java
Remarks: phlegmaria agg.
= Lycopodium apiculatum Spring; Miquel, Pl. Jungh. 3: 272 1854 [non Poiret 1814]
= Urostachys tauri Herter; Index Lyc. 85 (1949)
= Urostachys apiculatus (Spring) Herter ex Nessel; Bärlappgewächse 227 (1939)
Huperzia taxifolia (Sw.) Trevis. (1874)
Distr. [complete]: Puerto Rico, Costa Rica, Greater Antilles, Cuba, Nicaragua, S-Mexico, Chiapas, Belize, Guatemala, Honduras, El Salvador, Colombia, Venezuela, Ecuador, Greater Antilles, Galapagos, Jamaica, Saba Isl., St. Kitts, Nevis, Montserrat, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent, Grenada, British Guyana, Surinam, French Guiana
Remarks: taxifolia agg.
= Huperzia passerinoides (Kunth) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
= Huperzia passerinoides var. nitens (Schlecht. & Cham.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
= Lycopodium herminieri Spring; Bull. Acad. Roy. Sci. Belg. 8514. 1841
= Lycopodium nitens Schlecht. & Cham.; Linnaea 5: 623 (1830)
= Lycopodium passerinoides Kunth; Nova Genera & Sp. Pl. 1: 41 (1815)[1816]
= Lycopodium schwendeneri Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 50 (1909)
= Lycopodium taxifolium Sw.; Nova Genera & Sp. Pl. seu Prodromus 138 (1788)
= Lycopodium taxifolium var. passerinoides (Kunth) Baker; Handbook of the Fern-Allies 16 (1887)
= Phlegmariurus taxifolius (Sm.) Love & Love; Taxon 26(2-3): 324 (1977)
= Plananthus taxifolius (Sw.) P. Beauv.; Prod. aethéogam. 112 (1805)
= Urostachys bruelkei Nessel; Repert. Spec. Nov. Regn. Veg. 36(12-15): 184, t. 173 (1934)
= Urostachys herminieri (Spring) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 162 (1923)
= Urostachys nitens (Schltdl. & Cham.) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 164 (1923)
= Urostachys passerinoides (Kunth) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 417 (1927)
= Urostachys passerinoides var. nitens (Schlecht. & Cham.) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 418 (1927)
= Urostachys rubiginosus Herter ex Nessel; Revista Sudamericana de Botánica 6(5-6): 162, t. 8, f. 36 (1940)
= Urostachys schwendeneri (Herter) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 165 (1923)
= Urostachys schwendeneri (Herter) Herter var. spongiosus Nessel; Revista Sudamer. Bot. 6: 165 (1940)
= Urostachys spongiosus (Nessel) Herter; Index Lyc. 83 (1949)
= Urostachys schwendeneri var. nitens (Schlecht. & Cham.) Herter ex Nessel; Die Bärlappgewächse 166 (1939)
= Urostachys taxifolius (Sm.) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 162 (1923)
Huperzia tenuicaulis (Underw. & Lloyd) B. Øllg.; Opera Bot. 92: 169 (1987)
Distr. [complete]: Puerto Rico, Guadeloupe, Dominica, British Guyana
Remarks: linifolia agg.
= Urostachys tenuicaulis (Underwood & Lloyd) Herter; Repert. Spec. Nov. Regni Veg. 19: 165 (1923)
= Lycopodium tenuicaule Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33: 113 (1906)
Huperzia tenuis (Humb. & Bonpl. ex Willd.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
Distr. [complete]: Costa Rica, Panama, Colombia, Venezuela, Ecuador, N-Peru
Remarks: verticillata agg.
= Lycopodium tenue Humb. & Bonpl. ex Willd.; Sp. Pl., Editio quarta 5: 55 (1810)
= Urostachys pax-hominibus Herter; Index Lyc. 74 (1949) [nom. superfl. for Lycopodium tenue Willd.]
= Urostachys tenuis (Humb. & Bonpl. ex Willd.) Herter ex Nessel; Archivos de Botânica do São Paulo 1(4): 401 (1927)
Huperzia terrae-guilelmii (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: NIrian Jaya
Remarks: phlegmaria agg.
= Lycopodium terrae-guilelmii Herter; Bot. Jahrb. Syst. 54: 229 (1916)
= Urostachys terrae-guilelmii (Herter) Herter ex Nessel; Bärlappgewächse 231 (1939)
Huperzia tetragona (Hook. & Grev.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1874) **
Distr.: Ecuador
Remarks: saururus agg.
= Lycopodium catharticum Hook.; Annals of Natural Hisotry 1: 430-431, t. 14 (1838)
= Lycopodium myrsinites var. minus Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles 15: 29 (1842)
=Lycopodium tetragonum Hook. & Grev.; Icones Filicum 1: t. 109 (1829)
= Lycopodium tetragonum Hook. & Grev.; Icon. filic. 1: t. 109 (1829)
= Lycopodium tetragonum var. patulum Spring; Mémoires de l'Académie Royales des Sciences, Lettres & Beaux Arts de Belgique 24: 12 (1849)
= Urostachys catharticus (Hook.) Herter ex Nessel; Bärlappgewächse 135 (1939)
= Urostachys tetragonus (Hook. & Grev.) Herter ex Nessel; Bärlappgewächse 135 (1939)
Huperzia tetrapterygia (Herter) comb. ined.
Distr. [complete]: Queensland
Remarks: phlegmarioides agg.
= Lycopodium tetrapterygium (Herter) F.M. Bailey; Proc. Roy. Soc. Queensland 1: 150. 1884
= Urostachys tetrapterygius (Bailey) Herter; Index Lyc. 86 (1949)
Huperzia tetrasticha (Kunze) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Java, peninsular Malaysia (Kelantan, Pahan, Negeri Sembilan), Sulawesi
Remarks: carinata agg.
= Lycopodium tetrastichum Kunze; Bot. Zeit. (Berlin) 6: 99 (1848)
= Lycopodium sarasinorum Christ; Verh. Naturf. Ges. Basel 11: 34, t. 2o f. 11-12 (1894)
= Urostachys sarasinorum (Christ) Herter ex Nessel; Bärlappgewächse 182 (1939)
= Urostachys tetrastichus (Kunze) Herter ex Nessel; Bärlappgewächse 183 (1939, by reference to Alderw (1915a: 35))
Huperzia thwaitesii (Nessel) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Java
Remarks: selago agg.
= Urostachys thwaitesii Herter ex Nessel; [Bärlappgewächse 52. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 158, t. 8, f. 14 (1940)]
Huperzia tibetica (Ching) Ching; Acta Bot. Yunnanica 3: 304 (1981)
Distr.: Yunnan, Tibet
Remarks: selago agg.
= Lycopodium tibeticum Ching; Bull. Fan Mem. Inst. Biol. Bot. Ser. 10: 17 (1940)
= Huperzia tibetica (Ching) Ching; Flora Xizangica 1: 7 (1983) [nom. superfl.]
Huperzia toppingii (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Philippines
Remarks: squarrosa agg.
= Urostachys toppingii Herter; Bot. Arch. 3: 15 (1923)
Huperzia tournayana (LawaIrée) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Zaire
Remarks: phlegmaria agg.
= Phlegmariurus tournayanus Lawalrée; Acta Bot. Acad. Sci. Hung. 19: 197, f. 2 (1973)
Huperzia transiens (Alderw.) comb. ined.
Distr.: Sumatra
Remarks: phlegmaria agg.
= Lycopodium transiens Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 5: 227 (1922)
= Urostachys transiens (Alderw.) Herter ex Nessel; Bärlappgewächse 143 (1939)
Huperzia transilla (Baker) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Ecuador
Remarks: brevifolia agg.
= Lycopodium transilla Sodiro ex Baker; J. Bot. 15: 168 (1877)
= Urostachys transilla (Baker) Herter ex Nessel; Bärlappgewächse 106 (1939)
Huperzia treitubensis (Silveira) B. Øllg.; Opera Bot. 92: 170 (1987)
Distr.: Brazil (Minas Gerais)
Remarks: brongniartii agg.
= Lycopodium treitubense Silveira; Bol. Comm. Geogr. Geol. Minas Geraes 2, 5: 118, t. 3 (1898)
= Urostachys treitubensis (Silveira) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 380 (1927)
= Lycopodium hoehnei (Nessel) Rolleri; Revista Mus. La Plata (N. S.) Bot. 13: 77 (1981)
= Urostachys hoehnei Nessel; Arch. Bot. Sao Paulo 1- 383, t. 6 (1927)
= Lycopodium inflexum Silveira; Bol. Comm. Geogr. Geol. Minas Geraes 2,5: 118, t. 2 (1898) [non (P. Beauv.) Sw. (1806)]
= Urostachys inflexus Herter ex Nessel; Arch. Bot. Sao Paulo 1: 384. 1927 [nom. nov. for Lycopodium inflexum Silveira [non (P. Beauv.) Sw. (1806])]
= Urostachys capri Herter; Index Lyc. 55 (1949) [nom. superfl.]
Huperzia trifoliata (Copel.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Fiji
Remarks: phlegmaria agg.
= Lycopodium trifoliatum Copel.; Bernice P. Bishop Mus. Bull. 59: 7, t. 1 (1929)
= Urostachys trifoliatus (Copel.) Herter ex Nessel; Bärlappgewächse 224 (1939)
Huperzia trigona (C. Chr.) Tard.; Adansonia 10: 20 (1970)
Distr.: Madagascar
Remarks: obtusifolia agg.
= Lycopodium trigonum C. Chr.; Dansk Bot. Ark. 7.191, t. 76, f. 1-5 (1932)
= Urostachys humbertii Nessel; Repert. Sp. Nov. Regni Veg. 48: 170, t. 317 ("307"), f. 3 (1940)
= Urostachys trigonus (C. Chr.) Herter; Index Lyc. 87 (1949)
Huperzia tubulosa (Maxon) B. Øllg.; Annals of the Missouri Botanical Garden 79: 697 (1992)
Distr. [complete]: Costa Rica, Panama
Remarks: taxifolia agg.
= Lycopodium tubulosum Maxon; Contributions from the U. S. National Herbarium 17(2): 178, t. 10 (1913)
= Urostachys tubulosus (Maxon) Herter ex Nessel; Die Bärlappgewächse 177 (1939)
Huperzia ulei (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Brazil (Rio de Janeiro)
Remarks: brongniartii agg.
= Urostachys ulei Herter; Repert. Spec. Nov. Regni Veg. 19162 (1923)
Huperzia ulicifolia (Sw.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
Distr.: Distribution unknown
Remarks: squarrosa agg.
= Lycopodium ulicifolium Sw.; Syn. fil. 177 (1806)
= Urostachys ulicifolius (Sw.) Herter ex Nessel; Bärlappgewächse 204 (1939)
Huperzia ulixis (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Colombia
Remarks: saururus agg.
= Urostachys ulixis Herter; Revista Sudamer. Bot. 10: 115 (1953 [1954])
= Lycopodium ulixis (Herter) C. Morton; Amer. Fern 1. 54: 73 (1964)
Huperzia unguiculata B. Øllg.; Harling & Andersson: Fl. Ecuador 33: 26, f. 2B (1988)
Distr.: Ecuador
Remarks: reflexa agg.
Huperzia urbanii (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Ecuador
Remarks: reflexa agg.
= Lycopodium urbanii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 47 (1909)
= Urostachys urbanii (Herter) Herter ex Nessel; Bärlappgewächse 110 (1939)
Huperzia vanikorensis (Copel.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Santa Cruz Islands, Solomons, Vanuatu
Remarks: carinata agg.
= Lycopodium vanikorense Copel.; Journal of the Arnold Arboretum 12: 49 (1931)
= Urostachys vanikorensis (Copel.) Herter; Index Lyc. 87 (1949)
Huperzia varia (R. Br.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247 (1874)
Distr.: Tasmania, New Zealand, Chatham Isl., Antipodes Isl., Auckland Isl., Lord Howe Isl., New Ireland Remarks: varia agg.
Trivial name: Iwituna (Maori)
= Huperzia billardieri (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 247. 1874
= Lycopodium billardierei Spring; Mon. Lycop. 1, 56 (1843)
= Lycopodium novae-zelandicum Colenso
= Lycopodium flagellaria A. Rich.; Essai Fl. N. Z., 60 [non Bory (1832)]
= Lycopodium phlegmaria A. Cunn.; Compan. bot. Mag. 2, 361 [non L. (1837)]
= Lycopodium varium R. Br.; Prodr. 65 (1810)
= Lycopodium varium var. billardieri (Spring) T. Kirk; T. N. Z. I. 16, 377 (1884)
= Urostachys billardieri (Spring) Herter ex Nessel; Bärlappgewächse 194 (1939)
= Urostachys varius (R. Br.) Herter ex Nessel; Bärlappgewächse 192 (1939)

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Huperzia venezuelanica (Herter) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distr.: Venezuela
Remarks: saururus agg.
= Lycopodium venezuelanicum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909)
= Urostachys venezuelanus (Herter) Herter ex Nessel; Bärlappgewächse 74 (1939)
= Huperzia schneei (Vareschi) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Lycopodium schneei Vareschi; Acta Bot. Venez. 1: 83, f. 1 (1966)
Huperzia vernicosa (Grev. & Hook.) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
Distr.: India (Tamil Nadu, Korala), Sri Lanka
Remarks: selago agg.
= Lycopodium vernicosum Hook. & Grev. ex Grev. & Hook.; Bot. Misc. 2: 364 (1831)
= Phlegmariurus vernicosus (Grev. & Hook.) Löve & Löve; Taxon 26: 324 (1977)
= Urostachys vernicosus (Grev. & Hook.) Herter ex Nessel; Bärlappgewächse 61 (1939)
Huperzia verticillata (L. f.) Trevisan; Atti della Società Crittogamologica Italiana 17: 248 (1875)
Distr.: Puerto Rico, Greater Antilles, trop. Am, Greater Antilles, Mexico, to, Brazil, Trinidad, Saba Isl., Guadeloupe, Dominica, Martinique, St. Lucia, Transvaal, Natal, Cape Prov., Swaziland, trop. Africa, Mascarenes, Zimbabwe, Malawi, Mozambique, Reunion, Kenya, Tanzania, Transkei, Lesotho, NE- Oranje Free State, Madagascar, Comores, Bioko Isl. (Fernando Poo), Sao Tome
Remarks: verticillata agg.
= Lycopodium setaceum Lam.; Encycl. 3: 653 (1789 [1792])
= Plananthus setaceus (Lam.) P. Beauv.; Prod. aethéogam. 112 (1805)
= Lycopodium portoricense L.M. Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33: 108 (1906)
= Lycopodium verticillatum L. f.; Supplementum Plantarum 448 (1782)
= Plananthus verticillatus (L. f.) P. Beauv.; Prod. aethéogam. 112 (1805)
= Stachygynandrum verticillatum (L. f.) P. Beauv.; Prodr. aethéogam. 113 (1805)
= Urostachys portoricensis (Underwood & Lloyd) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 163 (1923)
= Urostachys verticillatus (L. f.) Herter; Beihefte zum Botanischen Centralblatt 39: 249 (1922)
Huperzia vorwerkii (Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
Distr.: India (Himalaya)
Remarks: selago agg.
= Urostachys vorwerkii Nessel; Repert. Spec. Nov. Regni Veg. 36: 178, t. 170 (1934)
Huperzia vrieseana (Spring) comb. ined.
Distr.: Java
Remarks: phlegmaria agg.
= Lycopodium vrieseanum Spring; Mem. Acad. Roy. Sci. Belg. 24 [Mon. Lyc. 2]: 33 (1849)
= Urostachys vrieseanus (Spring) Herter ex Nessel; Bärlappgewächse 236 (1939)
Huperzia warneckei (Nessel) Pichi-Serm.; Webbia 23: 163 (1968)
Distr.: Cameroun, Uganda, Tanzania, Sao Tome
Remarks: phlegmaria agg.
= Urostachys warneckei Herter ex Nessel; [Bärlappgewächse 213. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 166, t. 13, f. 64 (1940)]
= Lycopodium warneckei (Nessel) Alston; Bol. Soc. Brot.. ser. 2A, 30: 20 (1956)
Huperzia watsoniana (Maxon) B. Øllg.; Annals of the Missouri Botanical Garden 79: 701 (1992)
Distr. [complete]: El Salvador, Costa Rica, Panama
Remarks: verticillata agg.
= Lycopodium watsonianum Maxon; Smithsonian Miscellaneous Collections 56(29): 3, t. 3 (1912)
= Urostachys watsonianus (Maxon) Herter ex Nessel; Bärlappgewächse 159 (1939)
Huperzia weberbaueri (Hieron. & Herter ex Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
Distr.: Peru, Ecuador
Remarks: saururus agg.
= Huperzia papillata (Rolleri) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium papillatum Rolleri; Amer. Fern Journal 65(1): 3 (1975)
= Urostachys weberbaueri Herter ex Nessel; [Bärlappgewächse 99. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 162, t. 10p f. 41 (1940)]
Huperzia weddellii (Herter) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
Distr.: Peru, Ecuador
Remarks: brongniartii agg.
= Lycopodium weddellii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 45 (1909)
= Urostachys weddellii (Herter) Herter ex Nessel; Bärlappgewächse 91 (1939)
Huperzia whangshanensis Ching & P.C. Chiu; Acta Botanica Yunnanica 3(3): 299 (1981)
Distr.: S-Anhwei
Remarks: selago agg.
Huperzia whitfordii (Herter) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
Distr.: Luzon, Philippines
Remarks: squarrosa agg
. = Urostachys whitfordii Herter; Bot. Arch. 3: 14 (1923)
= Lycopodium whitfordii (Herter) Copel.; Univ. Calif. Publ. Bot. 12: 384 (1931)
Huperzia wilsonii (Underw. & F.E. Lloyd) B. Øllg.; Opera Bot. 92: 170 (1987)
Distr. [complete]: Puerto Rico, Guatemala, Nicaragua, Costa Rica, Panama, S-Mexico, Jamaica, Colombia, Venezuela, Ecuador, Peru, Greater Antilles, Guadeloupe, Saba Isl., Guadeloupe, Martinique, St. Lucia, St. Vincent, Surinam, Argentina (Misiones, Salta, Tucuman), Brazil, Paraguay, Uruguay
Remarks: wilsonii agg.
= Huperzia lindaviana (Herter) Holub; Folia Geobot. Phytotax. 33: 97 (1988)
= Huperzia mandiocana (Raddi) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248. 1874
= Lycopodium pseudo-mandiocanum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 49 (1909)
= Urostachys pseudo-mandiocanus (Herter) Herter ex Nessel; Arch. Bot. Sao Paulo 1: 411 (1927)
= Lycopodium andinum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 49. 1909 [non Rosenstock (1908)]
= Lycopodium arcanum Maxon; Field Museum of Natural History, Botanical Series 17(4): 310, t. 3 (1938)
= Lycopodium lindavianum Herter; Hedwigia 49(2-3): 90., as Lycopodium "Lindavianum" (1909)
= Lycopodium mandiocanum Raddi; Plantarum Brasiliensium Nova Genera 1: 77-78, t. 4 (1825)
= Lycopodium stamineum Maxon; Smithsonian Miscellaneous Collections 56(29): 2-3, t. 2 (1912)
= Lycopodium trichodendron Herter; Bot. Jahrb. Syst. 43(Beibl. 98): 49 (1909)
= Lycopodium wilsonii Underw. & Lloyd; Bulletin of the Torrey Botanical Club 33: 111 (1906)
= Urostachys lindavianus (Herter) Herter ex Nessel; Bärlappgewächse 146 (1939)
= Urostachys mandiocanus (Raddi) Herter; Repert. Spec. Nov. Regni Veg. 19: 164 (1923)
= Urostachys mandiocanus (Raddi) Herter var. filifolius Herter; Revista Sudamericana de Botánica 10: 124 (1954)
= Urostachys stamineus (Maxon) Herter ex Nessel; Die Bärlappgewächse 148 (1939)
= Urostachys trichodendron (Herter) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 163 (1923)
= Urostachys wilsonii (Underw. & Lloyd) Herter; Repert. Spec. Nov. Regn. Veg. 19(11-13): 163 (1923)
Huperzia x bartleyi (Cusick) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distr.: Ohio, E-USA
Remarks: selago agg.
Remarks: Lycopodium lucidulum x Lycopodium porophilum
= Lycopodium x bartleyi Cusick; Amer. Fern Journal 77(3): 100 (1987)
Huperzia x buttersii (Abbe) comb. ined.
Distr.: E-USA, E-Canada, Minnesota
Remarks: selago agg.
Remarks: Lycopodium lucidulum x Lycopodium selago
= Lycopodium x buttersii Abbe; Rhodora 55(651): 91-92 (1953)
Huperzia x calquistii
Distr. [complete]: Hawaii (Maui)
Remarks: Huperzia erubescens x Huperzia subintegra
Huperzia x gillettii Beitel & W.H. Wagner
Distr. [complete]: Hawaii (Maui, Kauai, Oahu, Molokai, Lanai, Hawaii Isl.)
Remarks: Huperzia serrata x Huperzia subintegra
Huperzia x medeirosii Beitel & W.H. Wagner
Distr. [complete]: Hawaii (Maui)
Remarks: Huperzia haleakalae x Huperzia subintegra
Huperzia x sulcinervia (Spring) Trevisan; Atti Soc. Ital. Sci. Nat. 17: 248 (1874)
Distr.: Hawaii (Oahu, Lanai, Hawaii Isl.), E-Asia
Remarks: Huperzia erubescens x Huperzia serrata
= Huperzia sulcinervia (Spring) Rothm.; Feddes Repert. 54: 59 (1951)
= Lycopodium x helleri Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909)
= Urostachys helleri (Herter) Herter ex Nessel; Bärlappgewächse 53 (1939)
= Lycopodium sulcinervium Spring; Bull. Acad. Roy. Sci. Belg. 8: 514 (1841)
= Urostachys sulcinervius (Spring) Herter ex Nessel; Bärlappgewächse 54 (1939)
Huperzia xiphophylla (Baker) Tardieu; Adansonia ser. 2 10: 20 (1970)
Distr.: Madagascar
Remarks: squarrosa agg.
= Lycopodium xiphophyllum Baker; Journal of the Linnean Society, Botany 22: 535 (1887)
= Urostachys xiphophyllus (Baker) Herter ex Nessel; Die Bärlappgewächse 72, t. 11, f. 9 (1939)
Huperzia yakusimensis (Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
Distr.: Japan
Remarks: selago agg.
= Lycopodium yakusimensis Herter ex Nessel; [Bärlappgewächse 65. 1939 [nom. inval.], Revista Sudamer. Bot. 6: 160, t. 8, f. 27 (1940)]
Huperzia yandongensis (Ching & Zhang) comb. ined.
Distr.: Zhejiang
Remarks: hamiltonii agg.
= Phlegmariurus yandongensis Ching & Zhang; Bull. Bot. Res. North-East. Forest. Inst. 3 (3): 2 (1983)
Huperzia yunnanensis (Ching) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
Distr.: Yunnan
Remarks: cancellata agg.
= Phlegmariurus yunnanensis Ching; Acta Bot. Yunnanica 4: 121 (1982)
Huperzia zollingeri (Herter) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
Distr.: Java;
Remarks: selago agg.
= Lycopodium zollingeri Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 48 (1909)
= Urostachys zollingeri (Herter) Herter ex Nessel; Bärlappgewächse 120 (1939)