Family Hymenophyllaceae, genus Hymenophyllum;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Hymenophyllum, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

      Hymenophyllum species A to H. Species I to Z here.

005.00 Class Filicopsida
		Trivial names: Ferns
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.

009.0000 Family Hymenophyllaceae Link; Handb. Erken. Gew. 3: 36 (1833)	
		Statistics: 8 genera, 250-300 spp.	
		= Hymenophyllaceae subfam. Cardiomanioideae K.lwats.; Acta Phytotax. Geobot. 35 (4-6): 
		177 (1984)

009.0004 Hymenophyllum J. E. Sm.; Mém. Acad. Turin 5: 4 (1793) Statistics: 330-350 spp. tropics / south-temperate, mostly neotropical [350 species and 1 hybrid listed]. Meringium, Mecodium and Sphaerocionium are frequently treated as separate genera, but very inconsistently in the various parts of the world. Unfortunately the re-inclusion of Mecodium leads to unavailable combinations in China. Revision date: 20.07.2001 = Acanthotheca Nakai; Bot. Mag. Tokyo 40. 242 (1926) = Amphipterum (Copeland) Presl; Epim. Bot. 258 (1849-51) = Apteropteris (Copel.) Copeland; Philip. Journ. Sc. 67: 34 (1938) = Buesia (Morton) Copeland; Philip. Journ. Sc. 67: 47 (1938) = Buesia Morton; Bot. Gaz. 93. 336 (1932) = Charitophyllum v. d. Bosch; Pl. Jungh. 1. (18) (1856) = Chilodium Presl; Hymenoph. 23 (1843) = Cycloglossum Presl; Hym. 32 (1843) = Dermatophlebium Presl; Epim. Bot. 258 (1849-51) = Diplophyllum v. d. Bosch; Versl. Meded. Konink. Akad. Wetens. Afd. Natuurk. 11: 318 (1861) [non Lehm. 1814 = Euphorophyllum van den Bosch; Plant. Jungh. 1. (19) (1856) = Hemicyatheon (Domin) Copeland; Philip. Journ. Sc. 67: 27 (1938) = Hymenoglossum Presl; Hymenoph. 35 (1843) = Hymenophyllum Sect. Sphaerocionium (Presl) Morton (1947) = Hymenophyllum subgen. Mecodium Copel.; Phil. Jour. Sci. 64: 93 (1937) = Hymenophyllum subgen. Meringium (Presl) Copel.; Philip. Journ. Sc. 64: 14 (1937) = Hymenophyllum subgen. Sphaerocionium (Presl) C. Chr.; Ind. Fil. Suppl. 3: 5 (1934) = Hymenophyllum subgen. Sphaerocionium (Presl) Copel.; Philip. Journ. Sc. 64: 164 (1937) = Leptocionium C. Presl; Hymenophyllaceae 26 (1843) = Mecodium Presl; Epim. Bot. 258 (1849-51) = Mecodium Presl ex Copeland; Phil. Jour. Sci. 67: 17 (1938) = Meringium C. Presl; Hymenophyllaceae 24 (1843) = Myriodon (Copel.) Copeland; Philip. Journ. Sc. 67: 47 (1938) = Myrmecostylum Presl; Hymenoph. 27 (1843) = Pachyloma Van den Bosch; Versl. Akad. Wet. Amsterdam 11. 318 (1861) = Ptychomanes Hedw.; Fil. Gen. et Spec. fasc. II (1800) = Ptychophyllum Presl; Hymenoph. 28 (1843) = Rosenstockia Copeland; Gen. Fil. 36 (1947) = Sphaerocionium C. Presl; Abh. Böhm. Ges. V. 3: 33 (1843) = Tetralasma Philippi; Linnaea 30. 208 (1860) Hymenophyllum abruptum Hook.; Sp. 1. 88 t. 31 B (1844) Hymenophyllum acanthoides (Bosch) Rosenst.; Bull. Jard. bot. Buit. II. nr. 11. 25 (1911) Hymenophyllum acrocarpum (Ching) comb. ined. Hymenophyllum aculeolatum v.d.Bosch; Ned. kr. Arch. 5(3). 179 (1863) Hymenophyllum adiantoides v.d.Bosch; Ned. kr. Arch. 5(3). 184 (1863) Hymenophyllum affine Brackenr.; Expl. Exp. 16. 265 t. 37 f. 1 (1854) Hymenophyllum alishanense DeVol; Taiwania 15: 283, f. 6 (1970) Hymenophyllum alveolatum C. Chr.; Ind. 356 (1905) Hymenophyllum amabile Morton; Journ. Wash. Acad. Sci. 22. 63 f. 1 (1932) Hymenophyllum andinum v.d.Bosch; Ned. kr. Arch. 4. 397 (1859) Hymenophyllum angulosum Christ; Phil. Journ. Sci. Bot. 3. 269 (1908) Hymenophyllum angustum v. d. Bosch; Ned. kr. Arch. 5(3). 183 (1863) Hymenophyllum antillense (Jenm.) Jenm.; Bull. Dep. Jam. II. 18. 6 (1890) Hymenophyllum apiculatum Mett. ex Kuhn; Linn. 36. 391 (1868) Hymenophyllum applanatum (Gray & Williams) comb. ined. Hymenophyllum archboldii (Copel.) C. Morton; Contrib. U.S. Nation. Herb. 38: 167 (1968) Hymenophyllum asplenioides Sw.; Schrad. Journ. 1800(2). 98 (1801) Hymenophyllum atrovirens Col.; Tasm. Journ. 2. 186 (1844) Hymenophyllum australe Willd.; Sp. 5. 527 (1810) Trivial names: Austral filmy ferm Hymenophyllum austrosinicum Ching; Hongkong Nat. 7: 87, 318 (1936) Hymenophyllum axillare Sw.; Schrad. Journ. 1800(2). 101 (1801) Hymenophyllum badium Hook. et Grev.; Icon. Fil. t. 76 (1828) Hymenophyllum baileyanum Domin; Bibl. Bot. 86. 21 t. 2 f. 2-3 (1913) Hymenophyllum bamlerianum Rosenst.; Fedde Repert. 10. 323 (1912) Hymenophyllum barbatum (v. d. Bosch) Bak.; Syn. Fil. 68 (1868) Hymenophyllum bartlettii (Copel.) C. Morton; Contrib. U.S. Nation. Herb. 38: 167 (1968) Hymenophyllum batuense Rosenst.; Bull. Jard. bot. Buit. II. nr. 11. 22 (1911) Hymenophyllum bicolanum Copel.; Philip. Journ. Sc. 64: 31. t.10 (1937) Hymenophyllum bismarckianum Christ; Schum. & Laut., Nachtr. Fl. deut. Südsee 34 (1905) Hymenophyllum bivalve (Forst.) Sw.; Schrad. Journ. 1800(2). 99 (1801) Hymenophyllum blandum Racib.; Pterid. Buit. 20 (1898) Hymenophyllum bontocense Copel.; Philip. Journ. Sc. 64: 33. t.12 (1937) Hymenophyllum bougainvillense J.W. Grimes; Kew Bull. 34(2): 233 (1979) Hymenophyllum brachyglossum A.Br. ex Kze.; Bot. Zeit. 227 (1847) Hymenophyllum brachypus Sod.; Cr. vasc. quit. 28 (1893) Hymenophyllum brassii C. Chr.; Brittonia 2: 273 (1937) Hymenophyllum breve Rosenst.; Fedde Rep. 20. 89 (1924) Hymenophyllum brevidens Alderw.; Bull. Jard. bot. Buit. II. nr. VII. 20 (1912) Hymenophyllum bryophilum C. Chr.; Brittonia 2: 274 (1937) Hymenophyllum campanulatum Christ; Phil. Journ. Sci. Bot. 2. 156 (1907) Hymenophyllum caparaoense Brade; Arq. Jard. Bot. Rio Janeiro 11: 21. t.1 (1951) Hymenophyllum capense Schrad.; Gött. gel. Anz. 911 (1818) Hymenophyllum capillaceum Roxb.; Beats., St. Helena 311 (1816) Hymenophyllum capillare Desv.; Prod. 333 (1827) var. alternialatum (Pichi-Serm.) Faden Hymenophyllum capurroi Sota; Darwiniana 17: 51 (1972) Hymenophyllum cardunculus C. Chr.; Mitt. Inst. Hamburg 7. 144 (1928) Hymenophyllum caudiculatum Mart.; Ic. Cr. Bras. 102 t. 67 (1834) var. productum (K. Presl) C. Chr. Hymenophyllum cernuum Gepp; Gibbs, Dutch N. W. New Guinea 68 (1917) Hymenophyllum cincinnatum Gepp; Gibbs, Dutch N. W. New Guinea 68 (1917) Hymenophyllum compactum Bonap.; N. Pt. 16. 16 (1925) Hymenophyllum consanguineum Morton; Contr. U.S. Nation. Herb. 29: 163 (1947) Hymenophyllum contractile Sod.; Cr. vasc. quit. 22 (1893) Hymenophyllum copelandianum Alderw.; Bull. Jard. bot. Buit. II. nr. VII. 19 (1912) Hymenophyllum copelandii C. Morton; Contrib. U.S. Nation. Herb. 38: 173 (1968) [nom. nov. Hymenophyllum corrugatum Christ; Bull. Boiss. II. 8. 508 (1903) var. elongatum Christ Hymenophyllum crassipetiolatum Stolze; Amer. Fern J. 66(1): 15 (1976) Hymenophyllum crispatoalatum Hayata; Ic. Fl. Formosa 6. 256 (1915) Hymenophyllum crispatum Wall. ex Hook. & Grev.; lcon. t. 77 (1828) Hymenophyllum crispum H.B.K.; Nov. Gen. & Sp. 1. 26 (1815) var. bipinnatisectum Morton (1947) Hymenophyllum cristatum Hook. & Grev.; Ic. Fil. t. 148 (1829) Hymenophyllum cumingii Presl; Hym. 32, 56 t. 11 A (1843) var. rariforme C. Chr. & Skottsb. var. calyciforme E. Br. (1931) Hymenophyllum cupressiforme Lab.; Nov. Holl. pl. spec. 2. 102 t. 250 f. 2 (1806) Trivial names: Common filmy fern Hymenophyllum darwinii Hook. f.; v.d.Bosch, Ned. kr. Arch. 53. 157 (1863) Hymenophyllum delicatulum Sehnem; Sellowia 7: 304. t. l(6) (1956) Hymenophyllum deltoideum C. Chr.; Cat. Pl. Mad. Pter. 18 (1932) (nomen), Dansk Bot, Ark. 7: 10 t. 2 f. 4-5 (1932) Hymenophyllum demissum (Forst. f.) Swartz; Schrad. Journ. 1800(2). 100 (1801) Trivial names: Irirangi (Maori) Hymenophyllum dentatum Cav.; Descr. 276 (1802) Hymenophyllum denticulatum Sw.; Schrad., Journ. Bot. 1800 (2): 100 (1801) Hymenophyllum dependens Morton; Contr. U. S. Nation.Herb. 29: 179 (1947) Hymenophyllum deplanchei Mett. ex Kuhn; Linn. 35. 393 (1868) Hymenophyllum devolii Lai Hymenophyllum dicranotrichum (Presl) Sadeb.; Nat. pfl. 1(4). 110 (1899) Hymenophyllum dilatatum (Forst.) Sw.; Schrad. Journ. 1800(2). 100 (1801) Hymenophyllum dimidiatum Mett. ex Kuhn; Linn. 35. 393 (1868) Hymenophyllum diversilabium (Copel.) C. Morton; Contrib. U.S. Nation. Herb. 38: 173 (1968) Hymenophyllum diversilobum (Presl) Fée; 11 mém. 118 (1866) Hymenophyllum eboracense Croxall; Austral. J. Bot. 23: 518 (1975) Hymenophyllum edanoi (Copel.) C. Morton; Contrib. U.S. Nation. Herb. 38: 173 (1968) Hymenophyllum edentulum (v.d.Bosch) C. Chr.; Ind. 360 (1905) Hymenophyllum elberti Rosenst.; Meded. Rijks Herb. Leiden nr. 14: 31 (1912) Hymenophyllum elegans Spreng.; Syst. 4. 133 (1827) Hymenophyllum elegantulum Bosch; Ned. kr. Arch. 4. 408 (1859) Hymenophyllum ellipticosorum Alderw.; Nova Guinea 14. 27 (1924) Hymenophyllum elongatum J.W. Grimes; Kew Bull. 34(2): 233 (1979) Hymenophyllum emarginatum Sw.; Schrad. Journ. 1800(2). 101 (1801) Hymenophyllum epiphyticum J. W. Moore; Bishop Mus. Bull. 102. 5 (1933) Hymenophyllum exsertum Wall.; List no. 170 1828 (nom. nud.) ex Hook.; Sp. Fil. 1: 109. t. 38A (1844) Hymenophyllum falklandicum Bak.; Syn. 68 (1867) Hymenophyllum farollonense Hier.; Engl. Jahrb. 34. 430 (1904) Hymenophyllum feejeense Brackenr.; Expl. Exp. 16. 266 t. 37 f. 2 (1854) Hymenophyllum fendlerianum Sturm; Fl. bras. 12. 291 (1859) Hymenophyllum ferax v.d.Bosch; Ned. kr. Arch. 4. 392 (1859) Hymenophyllum ferrugineum Colla; Herb. Pedem. 6: 190 (1836), et Mem. Ac. Torino 39. 30 (1836) var. donatii Looser; 1939 Hymenophyllum firmum Alderw.; Nova Guinea 14. 28 (1924) Hymenophyllum flabellatum Lab.; Nov. Holl. pl. spec. 2. 101 t. 250 f. 1 (1806) Trivial names: Shiny filmy fern Hymenophyllum flaccidum Bak.; Syn. ed. 1. 451 (1868) [non v.d.Bosch (1859) Hymenophyllum flexuosum A. Cunn.; Hook., Comp. Bot. Mag. 2. 369 (1836) Hymenophyllum foersteri Rosenst.; Fedde Repert. 12. 165 (1918) Hymenophyllum fragile (Hedw.) Morton; Contr. U. S. Nation. Herb. 29: 172 (1947) Hymenophyllum fuciforme Sw.; Syn. 148 (1806) Hymenophyllum fucoides (Sw.) Sw.; Schrad. Journ. 1800(2). 99 (1801) var. pedicellatum (Klotzsch) Hieron. (1904) var. calodictyon (Bosch) Stolze; Fieldiana, Bot. n.s. 20: 56 (1989) var. chachapoyense Stolze; Fieldiana, Bot. n.s. 20: 57 (1989) Hymenophyllum funckii v.d.Bosch; Ned. kr. Arch. 5(3). 166 (1863) Hymenophyllum fuscum (Bl.) v.d.Bosch; Hym. Jav. 62 t. 51-52 (1861) Hymenophyllum geluense Rosenst.; Fedde Repert. 6. 372 (1908) Hymenophyllum glaziovii Bak.; Hook., Ic., pl. t. 1612 (1886) Hymenophyllum gollmeri v.d.Bosch; Ned. kr. Arch. 5(3). 151 (1863) Hymenophyllum gongboense (Ching) comb. ined. Hymenophyllum gorgoneum Copel.; Philip. Journ. Sc. 64: 60. t.26 (1937) Hymenophyllum gracilescens Domin; Bibl. Bot. 86. 23 t. 1 f. 2-3 (1913) Hymenophyllum gracilius Copel.; Bishop Mus. Bull. 93. 7 t. 3 (1932) Hymenophyllum hainanense (Ching) comb. ined. Hymenophyllum halconense Cop.; Phil. Journ. Sci. Bot. 2. 144 (1907) Hymenophyllum hallierii Rosenst.; Bull. Jard. bot. Buit. II. nr. 11. 23 (1911) Hymenophyllum hamuliferum Alderw.; Bull. Buit. II no. 28. 29 (1918) Hymenophyllum heimii Tardieu; Bull. Soc. Bot. France 84: 538. f.1 (1938) Hymenophyllum helicoideum Sod.; Cr. vasc. quit. 20 (1893) Hymenophyllum hemidimorphum R.C.Moran & B.Ollg.; Nordic J. Bot. 15(2): 181 (1995) Hymenophyllum hemipteron Rosenst.; Fedde Rep. 22. 4 (1925) Hymenophyllum herterianum Brause; Engl. Jahrb. 50. 43 (1920) Hymenophyllum herzogii Rosenst.; Meded. Rijks Herb. Leiden nr. 19. 5 (1913) Hymenophyllum hieronymi (Brause) C. Chr.; Ind. Suppl. III. 114 (1934) Hymenophyllum hirsutum (L.) Sw.; Schrad. Journ. 1800(2). 99 (1801) Hymenophyllum hirtellum Sw.; Schrad. Journ. 1800(2). 101 (1801) var. gratum (Fée) Proctor (1966) var. vicentinum Proctor (1966) Hymenophyllum holochilum (v.d.Bosch) C. Chr.; Ind. 362 (1905) Hymenophyllum horizontale Morton; Contr. U. S. Nation. Herb. 29: 181 (1947) Hymenophyllum hosei Copel.; Phil. Journ. Sci. 12C. 46 (1917) Hymenophyllum howense Brownlie; Pacif. Sc. 14: 244 (1960) Hymenophyllum humatoides (Copel.) comb. ined. Hymenophyllum humbertii C. Chr.; Arch. Bot. (Caen) 2. Bull. mens. 209 (1928), Dansk Bot. Ark. 7. 10 t. 2 f. 6-8 (1932) Hymenophyllum humboldtianum Fourn.; Ann. sc. nat. V. 18. 265 (1873) Hymenophyllum hygrometricum (Poir.) Desv.; Prod. 333 (1827)
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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