Family Drynariaceae, genus Lepisorus;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Lepisorus, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

005.00 Class Filicopsida
		Trivial names: Ferns
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
050.000 Order Polypodiales
		Statistics: 9 families, 82 genera, 1996-2140 spp. 
		(2293 species and 31 hybrids listed]
		= Order Davalliales
054.0000 Family Drynariaceae Ching; Acta Phytotax. Sin. 16: 1-37 (1978)
		Statistics: 16 genera, 347-370 spp. 
		[444 species and 4 hybrids listed]
		Revision date: 20.07.2001
		Remarks: according to PCR results Polypodiaceae s. lat. is polyphyletic. 
		There are two choices: either include Grammitidaceae, Pleurosoriopsidaceae and 
		Loxogrammaceae in Polypodiaceae, or separate Drynariaceae as family. 
		It is not clear whether or not Drynariaceae is the correct family name.
		= Dipteridaceae subfam. Drynarioideae Crabbe; Jermy & Mickel, Fern Gaz. 11: 156 (1975)
		= Polypodiaceae tribus Drynarieae Subh.Chandra; Brenesia 18: 175 (1980) 
		[without latin descr. or type, et in Fern Gaz. 12(4): 226 (1982), with latin descr. and type
		= Polypodiaceae tribus Lemmaphylleae V.G.Tu; Novosti Sist. Vyssh. Rast. 18: 44 (1981)
		= Polypodiaceae tribus Microsoreae V.G.Tu; Novosti Sist. Vyssh. Rast. 18: 18 (1981)
054.0300 Tribus Lepisoreae

054.0304 Lepisorus (J. Sm.) Ching; Bull. Fan Mem. Inst. Biol. 4: 47 (1933) Statistics: > 80 spp. E- and tropical Asia [96 species listed]. There is taxonomical confusion between this genus and Pleopeltis s. str. Species may be shifted in the future between both genera. Revision date: 20.07.2001 = Paragramma (Blume) Moore (1957) = Grammitis subgen. Paragramma Blume; Epim. 114 (1828) Lepisorus affinis Ching ex Ching & Wang; Acta Phytotax. Sin. 8: 169. t.22(28) (1959) Lepisorus amaurolepidus (Sledge) Bir & Tirkha; Bir & Vasudeva, Journ. Bomb. Nat. Hist. Soc. 68: 192 (1971) Lepisorus angustifrons Tagawa; Acta Phytotax. Geobot. 6: 110 (1936) Lepisorus annuifrons Ching; Bull. Fan Mem. Inst. 4. 70 (1933) Lepisorus atropunctatus Ching (1933) Lepisorus balteiformis (Brause) Hovenkamp; Blumea 43: 110 (1998) Lepisorus bilouensis Ching & Y.X.Lin ex K. H. Shing; Vasc. Pl. Hengduan Mount. 31(6):573 (1993), without latin descr. or type, et in Acta Phytotax. Sin. 1: 166 (1993) Lepisorus boninensis Ching; Bull. Fan Mem. Inst. 4. 76 (1933) Lepisorus calcifer Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 25 (1984) Lepisorus clathratus (Clarke) Ching; Bull. Fan. Mem. Inst. Biol. 4: 71 (1933) Lepisorus coaetaneus Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 12 (1983) Lepisorus confluens W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 55 (1992) Lepisorus contortus (H. Christ in Baroni & H. Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 90 (1933) Lepisorus crassipes Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 18 (1983) Lepisorus crassirhizoma Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 3 (1983) Lepisorus distans (Makino) Ching; Acta Phytotax. Sin. 10: 302 (1965) Lepisorus eilophyllus (Diels) Ching; Bull. Fan Mem. Inst. 4. 65 (1933) Lepisorus elegans Ching & W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 55 (1992) Lepisorus ellipticus Ching; Bull. Bot. Res., Harbin 2(2): 74 (1982) Lepisorus excavatus (Willd.) Ching; Bull. Fan Mem. Inst. Biol. 4 68 (1939) Lepisorus hachijoensis Kurata; Sci. Rep. Yokosuka City Mus. 11: 40 (1965) Lepisorus henanensis Ching & S. K. Wu; Acta Bot. Yunnan. 5(1): 13 (1983) Lepisorus heterolepis (Rosenst.) Ching; Bull. Fan Mem. Inst. 4. 86 (1933) Lepisorus hsiawutaiensis Ching & S. K. Wu; Acta Bot. Yunnan. 5(1): 6 (1983) Lepisorus jinfoshanensis Ching; Bull. Bot. Res., Harbin 3(4): 4 (1983) Lepisorus kansuensis Ching & S. K. Wu; Acta Bot. Yunnan. 5(l): 17 (1983) Lepisorus kawakamii (Hayata) Tagawa; Acta Phytotax. Geobot. 5: 109 (1936) Lepisorus kochenensis Ching; Bull. Fan Mem. Inst. 4. 69 (1933) Lepisorus kuchenensis (Y.C. Wu in Y.C. Wu, K. Wong & Pong) Ching; Bull. Fan Mem. Inst. Biol. 4: 69 (1933) Lepisorus lancifolius Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 75 (1941) Lepisorus lewisii (Baker) Ching; Bull. Fan Mem. Inst. 4. 65 (1933) Lepisorus ligulatus Ching & S. K. Wu; Acta Bot. Yunnan. 5(1): 6 (1983) Lepisorus likiangensis Ching & S. K. Wu; Acta Bot. Yunnan. 5(1): 12 (1983) Lepisorus linearifolius Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 5 (1983) Lepisorus lineariformis Ching & S. K. Wu; Fl. Xizangica 1: 309 (1983) Lepisorus longifolius (BI.) Holtt.; Fl. Malaya II. 151 (1954) Lepisorus longipes Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 25 (1984) Lepisorus longus Ching; Acta Phytotax. Sin. 10: 192 (1965) [nom. nov. Lepisorus loriformis (Wall. ex Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 81 (1933) Lepisorus macrosphaerus (Bak.) Ching; Bull Fan Mem. Inst. Biol. Bot. 4: 73 (1933) var. maximus Ching; Fl. Xizangica 1: 303 (1983) var. minimus Ching; Fl. Xizangica 1: 304 (1983) Lepisorus mamas Hovenkamp; Blumea 43: 112 (1998) Lepisorus maowenensis Ching & S. K. Wu; Acta Bot. Yunnan. 5(1): 14 (1983) Lepisorus marginatus Ching; Fl. Tsinlingensis 2: 233 (1974) [without type). Lepisorus medogensis Ching & Y.X.Lin; Acta Phytotax. Sin. 22(5): 401 (1984) Lepisorus megasorus (C. Chr.) Ching; Bull. Fan Mem. Inst. 4. 79 (1933) Lepisorus mehrae Fraser-Jenkins; New Species Syndrome in Indian Pteridology and the Ferns of Nepal 159 (1997) Lepisorus mikawanus Kurata; Sci. Rep. Yokosuka City Mus. 11: 38 (1965) Lepisorus monilisorus (Hayata) Tagawa; Acta Phytotax. Geobot. 11: 303 (1942) Lepisorus morrisonensis (Hayata) H.Itô; Journ. Jap. Bot. 11: 92 (1935) Lepisorus myriosorus Ching; Wuyi Sci. J. 1- 11 (1981) Lepisorus namegatae (Sa.Kurata) Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 16 (1983) Lepisorus nanchanensis Ching; Bull. Bot. Res., Harbin 3(4): 4 (1983) Lepisorus neolewisii K. H. Shing; Acta Phytotax. Sin. 31(6): 572 (1993) Lepisorus niger Ching; Fl. Xizangica 1: 312 (1983) Lepisorus nudus (Hook.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 83 (1933) Lepisorus nyalamensis Ching & S. K. Wu; Fl. Xizangica 1: 311 (1983) Lepisorus obscurevenulosus (Hayata) Ching; Bull. Fan Mem. Inst. 4. 76 (1933) Lepisorus obtusifrons (Hayata) H.Itô; Journ. Jap. Bot. 11: 93 (1935) Lepisorus oligolepidus (Bak.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 80 (1933) Lepisorus onoei Ching; Bull. Fan Mem. Inst. 4. 87 (1933) Lepisorus oosphaerus (C. Chr.) Ching; Bull. Fan. Mem. Inst. Biol. Bot. 4: 73 (1933) Lepisorus paohuashanensis Ching; Jiang su zhi wu zhi (Rec. Fl. Jiangsu (Fl. Kiang Su) Prov.) 1: 467 (1977) Lepisorus papakensis (Masam.) Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 7 (1983) Lepisorus patungensis Ching & S. K. Wu; Acta Bot. Yunnan. 5(1): 11 (1983) Lepisorus perrierianus Ching; Bull. Fan Mem. Inst. 4. 51 (1933) Lepisorus petiolatus Ching & Y.X.Lin; Acta Phytotax. Sin. 31(5): 475 (1993) Lepisorus pseudoangustus Ching; Fl. Xizangica 1: 310 (1983) Lepisorus pseudolewisii K. H. Shing; Vasc. Pl. Hengduan Mount. 1: 166 (1993) [without latin descr. or type) Lepisorus pseudonudus Ching; Bull. Fan Mem. Inst. Biol. Bol. 4: 83 (1933) Lepisorus pseudoussuriensis Tagawa; Acta Phytotax. Geobot. 5: 110 (1936) Lepisorus pumilus Ching .& S. K. Wu; Acta Bot. Yunnan. 5(1): 16 (1983) Lepisorus pygmaeus Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 24 (1984) Lepisorus schraderi (Mett.) Ching; Bull. Fan Mem. Inst. 4. 51 (1933) Lepisorus sesquipedalis (J. Sm.) Fraser-Jenkins (1992) Lepisorus shansiensis Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 18 (1983) Lepisorus shanyangensis Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 19 (1983) Lepisorus shensiensis Ching & S. K. Wu; Acta Bot. Yunnan. S(1): 14 (1983) Lepisorus simulans Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 23 (1984) Lepisorus sinensis (H. Christ) Ching; Bull. Fan Mem. Inst. 4. 63 (1933) Lepisorus sinicus Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 2 (1983) Lepisorus sinuata (Ching & S.K. Wu) S.L. Yu (1997) Lepisorus sordidus (C. Chr.) Ching; Bull. Fan Mem. Inst, Biol. Bot. 4: 79 (1933) Lepisorus soulieanus (Christ) Ching & S. K. Wu; Acta Bot. Yunnan. 5(1): 11 (1983) Lepisorus subconfluens Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 85 (1933) Lepisorus sublinearis (Bak. ex Takeda) Ching; Bull. Fan Mem., Inst. Biol. Bot. 4: 78 (1933) Lepisorus sungpanensis Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 16 (1983) Lepisorus thaipaiensis Ching & S. K. Wu; Acta Bot. Yunnan. 5(1): 8 (1983) Lepisorus thunbergianus (Kaulf.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 88 (1933) var. subspathulatus (Hook.) Ching (1933) Lepisorus tosaensis (Mak.) H.Itô; Journ. Jap. Bot. 11 93 (1935) Lepisorus uchiyamae (Mak.) H.Itô; Journ. Jap. Bot. 11 93 (1935) Lepisorus undulatus Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 23 (1984) Lepisorus ussuriensis (Regel & Mack.) Ching; Fan Mem. Inst. Biol. 4: 91 (1933) Lepisorus venosus Ching & S. K. Wu; Acta Bot. Yunnan. 5(1,- 15 (1983) Lepisorus virescens Ching & S. K. Wu; Fl. Xizangica 1: 306 (1983) Lepisorus yunnanensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 75 (1941) Lepisorus zosterifolius Ching & Y.X.Lin; Acta Phytotax. Sin. 22(5): 401 (1984)
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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