Family Lycopodiaceae, genus Lycopodiella;
world species list.
By Michael Hassler and Brian Swale.
A list of the species of the genus Lycopodiella, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.
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001.00 Class Lycopsida
Trivial names: Lycophytes
Statistics: 3 orders, 3 families, 6 genera, 1313 spp. [1321 species and 28 hybrids listed]
001.000 Order Lycopodiales
Statistics: 1 family, 483 spp. [487 species and 12 hybrids listed]
001.0000 Family Lycopodiaceae Beauv. ex Mirb.; Hist. Nat. Veg. (Lam. & Mirb.) 4: 293 (1802)
Trivial names: Bärlappgewächse
Statistics: 4 genera, 483 spp. [487 species and 12 hybrids listed]
= Huperziaceae Rothm.; Fedde Repert. 66: 236 (1962) [nom. nov.
= Urostachyaceae Rothm.; Fedde Repert. 54: 58 (1944) [nom. illeg.
= Phylloglossaceae Kze.; Bot. Zeit. 1: 722 (1843)
001.0004 Lycopodiella Holub; Preslia 36: 22 (1964)
Trivial names: Sumpfbärlapp, Moorbärlapp
Statistics: 38 spp. worldwide, mostly New World [38 species and 2 hybrids listed]
Revision date: 20.07.2001
= Lateristachys Holub; Folia Geobot. Phytotax. 18: 440. 1983 [Preslia 47: 104. 1975]
[nom. inval.
= Palhinhaea Franco & Carv., Vasc. ex Carv., Vasc. & Franco; Bol.
Soc. Brot., ser. 2, 41: 24. 1967
= Pseudolycopodiella Holub; Folia Geobot. Phytotax. 18: 441 (1983)
Lycopodiella affinis (Bory) Pichi-Serm.; Webbia 2: 165 (1968)
Distribution: Liberia, Guinea, Sierra Leone, Ivory Coast, Gabon, Nigeria, Cameroun,
Zambia, Zimbabwe, Malawi, Mozambique, Mauritius, Réunion, Burundi, Madagascar,
Zaire, Burundi, Angola, Natal, Cape Prov., Uganda, Tanzania, Burkina Faso, peninsular
Malaysia (Trengganu, Pahan, Johor), Guangdong, India, Sri Lanka, Japan
Remarks: sect. Caroliniana
= Lepidotis radicans ("L., Gmel.") P. Beauv.; Prodr. aethéogam. 109
(1805, status uncertain)
= Lepidotis repens P. Beauv.; Prodr. Aetheog. 109 (1805)
= Lycopodiella caroliniana var. affine (Bory) Schelpe
= Lycopodium affine Bory; Voy. Iles Afrique 2: 204 (1804)
= Lycopodium carolinianum var. affine (Bory) Schelpe (1967)
= Lycopodium radicans "L., Gmel." repens P. Beauv.; Prodr. aethéogam. 109 (1805)
= Lycopodium repens (P. Beauv.) Sw.; Syn. Fil. 180 (1806)
= Pseudolycopodiella affinis (Bory) Holub; Folia Geobot. Phytotax. 20: 79 (1985)
= Lycopodium ericetorum Schrad.; Gött. Gel. Anz. 1818: 920 (1818)
Lycopodiella alopecuroides (L.) Cranfill; Amer. Fern. J. 71: 97 (1981)
Distribution [complete]: S-USA, E-USA, SE-USA (Arkansas, Connecticut, Delaware,
Florida, Georgia, Kentucky, Louisiana, Maryland, Massachusetts, Mississippi,
New Jersey, New York, North Carolina, Pennsylvania, Rhode Island, South Carolina,
Texas, Virginia), Guatemala, Mexico (Chiapas), El Salvador, Colombia, Venezuela,
Ecuador, Peru, Bolivia, Brazil, Paraguay, Cuba, Uruguay, Argentina (Buenos Aires,
Misiones, Corrientes)
Trivial names: Foxtail Club-Moss
Remarks: sect. Lycopodiella
= Lepidotis alopecuroides (L.) Rothm.; Feddes Repert. 54: 66 (1944)
= Lycopodiella matthewsii (Hook.) Holub; Folia Geobot. Phytotax. 18: 441 (1983)
= Lycopodium alopecuroides L.; Sp. Pl. 2: 1100 (1753)
= Lycopodium inundatum L. var. alopecuroides (L.) Tuckerman
= Lycopodium matthewsii Hook.; Icones Plantarum 1: t. 26 (1836)
= Plananthus alopecuroides (L.) P. Beauv.; Prodr. Aetheog. 111 (1805)
var. integerrima (Spring) B. Øllg. & Windisch
Distribution [complete]: British Guyana, Surinam, Costa Rica, Cuba, Mexico, Guatemala,
Venezuela, Colombia, Ecuador, Bolivia, Brazil (Santa Catarina)
= Lycopodium alopecuroides var. integerrimum Spring (1842)
= Lycopodium longipes Hook. & Grev. ex Grev. & Hook.; Bot. Misc. 2: 372 (1831)
Lycopodiella appressa (Chapm.) Cranfill; Amer. Fern Journal 71: 97 (1981)
Distribution [complete]: E-Canada (New Brunswick, Newfoundland, Nova Scotia,
Prince Edward Isl.), S-USA, SE-USA, E-USA (Alabama, Arkansas, Connecticut, Delaware,
Florida, Georgia, Illinois, Kansas, Kentucky, Louisiana, Maine, Maryland,
Massachusetts, Mississippi, Missouri, New Hampshire, New Jersey, New York,
North Carolina, Oklahoma, Pennsylvania, Rhode Island, South Carolina, Tennessee,
Texas, Virginia), Cuba
Trivial names: Southern Club-Moss
Remarks: sect. Lycopodiella
= Lycopodiella bigelowii (Tuck.) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
= Lycopodium appressum (Chapman) Lloyd & Underwood; Bulletin of the
Torrey Botanical Club 27: 153 (1900)
= Lycopodium alopecuroides L. ssp. appressum (Chapman) Clute
= Lycopodium bigelovii Oakes & Tuckerman ex Tuckerman (err. cit. in Spring 1849: 33)
= Lycopodium chapmanii Underwood ex Maxon; Proc. U. S. Natl. Mus. 23: 646. 1901
= Lycopodium inundatum L. var. appressum Chapman; Bot. Gaz. 3: 20 (1878)
= Lycopodium inundatum L. var. bigelowii Tuckerm.; Amer. J. Sci. Arts. 45: 47 (1843)
f. polyclavatum McDonald; Fern Bulletin 9: 9 (1901)
Distribution: North America
= Lycopodium alopecuroides f. polyclavatum (McDonald) Clute; Fern Bulletin
17: 45., under var. adpressum (1909)
= Lycopodium inundatum f. polyclavatum (McDonald) Fernald; Rhodora
42(502): 405 (1940)
Lycopodiella bradei (Herter) B. Øllg.; Opera Bot. 92: 176 (1987)
Distribution: Brazil
Remarks: sect. Campylostachys
= Lycopodium bradei Herter; Revista Sudamer. Bot. 8: 21 (1949)
Lycopodiella brevibracteata (Alderw.) comb. ined.
Distribution: New Guinea
Remarks: unverified name, sect. Campylostachys
= Lycopodium brevibracteatum Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2,
24: 5 (1917)
= Palhinhaea brevibracteata (Alderw.) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
Lycopodiella brevipedunculata (Alderw.) comb. ined.
Distribution: New Guinea
Remarks: unverified name, sect. Caroliniana
= Lycopodium carolinianum L. var. brevipedunculatum Alderw.; Nova Guinea
14: 64. 1924 [non Spring (1842)
= Lycopodium brevipedunculatum (Alderw.) Herter; Index Lyc. 7 (1949)
= Lycopodium carolinianum var. brevipedunculatum Alderw. (1924)
Lycopodiella camporum B. Øllg. & P.G. Windisch; Bradea
5: 24, f. 3 (1987)
Distribution [complete]: Venezuela, Brazil (Minas Gerais), Colombia, Peru, Brazil,
E-Bolivia, British Guyana, Surinam, French Guiana
Remarks: sect. Campylostachys
Lycopodiella carnosa (Silveira) B. Øllg.; Opera Bot. 92: 176 (1987)
Distribution: Brazil
Remarks: sect. Caroliniana
= Lycopodium carnosum Silveira; Comm. Geogr. Minas Gerais 5(2): 119, t. 7, 8 (1898)
= Pseudolycopodiella carnosa (Silveira) Holub; Folia Geobot. Phytotax. 20: 79 (1985)
= Lycopodium goyazense L. Underw. & F. Lloyd; Bull. Torrey Bot. Club 33: 121 (1906)
= Pseudolycopodiella goyazensis (L. Underw. & F. Lloyd) Holub; Folia
Geobot. Phytotax. 18: 442 (1983)
Lycopodiella caroliniana (L.) Pichi-Serm.; Webbia 23(1): 165 (1968)
Distribution [complete]: E-USA, SE-USA (Alabama, Delaware, Florida, Georgia, Kentucky,
Louisiana, Maryland, Massachusetts, Mississippi, New Jersey, New York, North Carolina,
Pennsylvania, South Carolina, Texas, Virginia), Mexico, Nicaragua, Japan (Honshu), Sri Lanka
Remarks: sect. Caroliniana
Trivial names: Slender Club-Moss
= Lepidotis caroliniana (L.) P. Beauv.; Prodr. Aetheog. 108 (1805)
= Lycopodium carolinianum L.; Sp. Pl. 2: 1100 (1753)
= Pseudolycopodiella caroliniana (L.) Holub; Folia Geobot. Phytotax. 18: 442 (1983)
= Lycopodium subinundatum Tag.; Acta Phytotax. Geobot. 10: 75 (1941)
var. meridionalis (Underw. & F.E. Lloyd) B. Øllg. & P.G. Windisch; Bradea 5: 27 (1987)
Distribution [complete]: Puerto Rico, Cuba, Jamaica, Hispaniola, Belize, Guatemala,
Honduras, El Salvador, Nicaragua, Panama, Guadeloupe, Dominica, Trinidad, Colombia,
Mexico, Venezuela, Ecuador, Peru, British Guyana, Surinam, French Guiana, Bolivia
= Lycopodium carolinianum var. meridionalis (Underw. & Lloyd) Nessel;
Archivos de Botânica do São Paulo 1: 431 (1927)
= Lycopodium meridionale Underw. & Lloyd; Bulletin of the Torrey
Botanical Club 33: 121 (1906)
= Pseudolycopodiella meridionalis (Underw. & Lloyd) Holub; Folia
Geobot. Phytotax. 18: 442 (1983)
var. grandifolium (Spring.) comb. ined.
Distribution: Transvaal, Natal, Malawi, Mozambique, Zambia, Zimbabwe, Angola, Tanzania, Zaire
= Lycopodiella sarcocaulon (Kuhn) Pichi-Serm.; Webbia 23: 166 (1968)
= Lycopodium carolinianum var. grandifolium Spring. (1849)
= Lycopodium sarcocaulon Welw. & A. Br. ex Kuhn; Filic. Afr. 210 (1868)
= Pseudolycopodiella sarcocaulon (Kuhn) Holub; Folia Geobot. Phytotax. 18: 442 (1983)
var. paradoxa (Mart.) B. Øllg. & P.G. Windisch; Bradea 5: 27 (1987)
Distribution [complete]: Brazil, Venezuela, Paraguay, Uruguay
= Lycopodium carolinianum var. paradoxum (Mart.) H. Christ; Plantas Novas
Mineiras 2: 42 (1900)?
= Lycopodium paradoxum Mart.; Icones Plantarum Cryptogamicarum 38, t. 20,
f. 2 (1838)
= Pseudolycopodiella paradoxa (Mart.) Holub; Folia Geobot. Phytotax. 18:
442 (1983)
var. tuberosum (Welw. & A. Br. ex Kuhn) comb. ined.
Distribution [complete]: Zambia, Angola
= Lycopodium carolinianum var. tuberosum (Welw. & A. Br. ex Kuhn) Nessel (1939)
= Lycopodium tuberosum A. Braun & Welw. ex Kuhn; Filic. Afr. 211 (1868)
= Pseudolycopodiella tuberosa (Kuhn) Holub; Folia Geobot. Phytotax. 18: 442 (1983)
Lycopodiella cernua (L.) Pichi-Serm.; Webbia 23(1): 166 (1968)
Distribution: Azores, Madeira, *Portugal, *Sicily, *Lebanon, Transvaal, Natal, Cape Prov., Swaziland,
Liberia, Ghana, Nigeria, Cameroun, Bioko Isl. (Fernando Poo), Sao Tomé, Principe Isl., Gabon,
Sudan, Ethiopia, Zaire, Rwanda, Burundi, Uganda, Kenya, Tanzania, Angola, Zambia, Zimbabwe,
Malawi, Mozambique, Sierra Leone, Madagascar, Mauritius, Réunion, Comores, Seychelles,
Capverdes, St. Helena, Senegal, Gambia, Guinea-Bissau, Mali, Ivory Coast, Togo, Benin, Central
African Republic, Equatorial Guinea, Congo, Zaire, Ascension Isl., Japan, Ryukyu Isl., Taiwan,
peninsular Malaysia, Malesia, Borneo (Mt. Kinabalu, etc.), Java, Sikkim, Philippines, India (throughout,
in mountainous regions, Assam, NE-India), Burma, Bangladesh, Thailand, Sri Lanka, Andamans,
Nicobars, Western Australia, Northern Territory, Queensland, New South Wales, New Zealand (North
Isl., South Isl.), Kermadec Isl., Tonga ('Eua, Tongatapu, Late Isl., Kao, Vava'u), Austral Isl. (Rapa Iti,
Rurutu, Tubuai), Marquesas Isl. (Nuku Hiva, Hiva Oa, Ua Huk, Fatu Hiva), Society Isl. (Tahiti), Western
Samoa (Upolu), American Samoa (Tutuila), Southern Marianas (Saipan, Rota Isl., Guam), Northern
Marianas (Pagan, Alamagan), Fiji (Viti Levu, Ovalau), Palau Isl. (Babeldoab, Koror, Nagarakabesang,
Aulupse'el, Peliliu), Micronesia (Yap, Truk, Moen, Dublon, Fefan, Pohnpei, Kosrae), Wallis and Futuna
Isl., Norfolk Isl., Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.), New Caledonia, Cook Isl.
(Rarotonga), Louisiana, S-Georgia, N-Florida, C-Florida, Alabama, Mississippi, South Carolina, Costa
Rica, Galapagos, Cuba, Hispaniola, Jamaica, Puerto Rico, Nicaragua, Mexico (Chiapas, Nayarit, Jalisco,
Mexico State, Oaxaca, Puebla, Veracruz, Hidalgo, Tabasco), Belize, Guatemala, Honduras, El Salvador,
Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil, Paraguay, Trinidad, St. Kitts, Montserrat,
Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent, Grenada, Barbados, Nevis, Venezuela, British
Guyana, Surinam, French Guiana, Uruguay, Argentina (Buenos Aires, Corrientes, Misiones)
Remarks: sect. Campylostachys
= Lepidotis cernua (L.) PB.; Prodr. Aethéogam.: 108 (1805)
= Lycopodiella veigae (Vasc.) Hansen & Sunding
= Palhinhaea veigae Vasc.
= Lycopodium boryanum A. Rich.; Sert. Astrol. 42 (1834)
= Lycopodium capillaceum (Spring) Hieron.; Bot. Jahrb. Syst. 34: 573 (1905)
= Lycopodium cernuum L.; Sp. Pl. 2: 1100 (1753)
= Lycopodium cernuum L. var. capillaceum Spring; Mém. Acad. Roy.
Sci. Belg. 15 [Mon. Lyc. 1]: 80 (1842)
= Lycopodium cernuum var. marianum (Willd.) Bak.
= Lycopodium cernuum var. sikkimense (Müll. Berol.) C.B. Clarke (1880)
= Lycopodium cernuum var. vulcanicum (Blume) Herter; Philippine Journal of
Science 22: 182 (1923)
= Lycopodium cymosum (Fée) L'Herm. ex Hieron.; Bot. Jahrb. Syst. 34: 574 (1905)
= Lycopodium dichotomum Blanco; Fl. Filip. ed. 2.: 570. 1845, ed. 3 (3):
240. 1879 [non Jacq (1762)
= Lycopodium heeschii K. Müll.; Botanische Zeitung. Berlin 19: 164 (1861)
= Lycopodium hupeanum K. Mueller; Bot. Zeit. (Berlin) 19 (24): 165 (1861)
= Lycopodium marianum Willd.; Sp. pl. 5: 31 (1810)
= Lycopodium moritzii K. Müll.; Bot. Zeit. (Berlin) 19 (24): 165 (1861)
= Lycopodium polycephalum Col.; Trans. & Proc. New Zealand Inst. 27: 401 (1895)
= Lycopodium secundum K. Müll.; Bot. Zeit. (Berlin) 19: 164 (1861)
= Lycopodium sikkimense K. Mueller; Bot. Zeit. (Berlin) 19 (24): 164
(1861) [fide Fraser-Jenkins (1997)
= Lycopodium veneris Herter; Index Lyc. 46. 1949 [nom. superfl. for
Lycopodium vulcanicum Blume, by reference to Lycopodium cernuum L. var.
vulcanicum (Blume) Herter (1923c: 182)
= Lycopodium vulcanicum Blume; Enumeratio Plantarum Javae 2: 266 (1828)
= Palhinhaea capillacea (Spring) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
= Palhinhaea cernua (L.) Carv. Vasc. & Franco; Bol. Soc. Brot., ser. 2, 41: 25 (1967)
= Palhinhaea cernua f. sikkimensis (Müll. Berol.) H.S. Kung (1988)
= Palhinhaea cernua var. pendula (Hook.) Nair & Ghosh; [fide Fraser-Jenkins (1997)
= Palhinhaea cernua var. sikkimensis (Müll. Berol.) Ching (1981)
= Palhinhaea cernua var. sikkimensis (Müll. Berol.) Ching (1983)
= Palhinhaea mariana (Willd.) Holub; Folia Geobot. Phytotax. 20: 79 (1985)
= Palhinhaea polycephala (Colenso) Holub; Folia Geobot. Phytotax. 20: 79 (1985)
var. curvata (Sw.) comb. ined.
Distribution [complete]: Puerto Rico, Jamaica, Venezuela, Colombia, Ecuador, Peru,
Bolivia, Costa Rica
= Palhinhaea cernua var. curvata (Sw.) Nair & Ghosh; [fide Fraser-Jenkins (1997)
= Lycopodium curvatum var. cymosum L'Hermin. ex Fée; Hist. foug. Ant. 131 (1866)
= Lycopodium cernuum var. curvatum (Sw.) Hook. & Grev. (1831)
= Lycopodium curvatum Sw.; J. Bot. (Schrader) 1800 (2): 116. 1801 [Syn.
fil. 178, 402 (1806])
Lycopodiella contexta (C. Martius) Holub; Folia Geobot. Phytotax. 18: 441 (1983)
Distribution [complete]: Colombia, Venezuela, Peru, Brazil
Remarks: sect. Caroliniana
= Lepidotis contexta (C. Martius) Rothm.; Feddes Repert. 54: 66 (1944)
= Lycopodium contextum C. Martius; Icon. pl. crypt. 38, t. 20, f. 1 (1834)
= Lycopodium sprucei Baker; Handb. fern-allies 24 (1887)
= Lycopodium alopecuroides var. contextum (Mart.) Baker (1887)
Lycopodiella descendens B. Øllg.; Harling & Andersson, Fl. Ecuador 33:
143, f. 28A (1988)
Distribution [complete]: Ecuador, Peru
Remarks: sect. Campylostachys
Lycopodiella diffusa (R. Br.) B. Øllg.; Opera Bot. 92: 176 (1987)
Distribution [complete]: Tasmania, New Zealand (North Isl., South Isl., Stewart Isl.), Auckland Isl.
Remarks: sect. Lateristachys
= Lateristachys diffusa (R. Br.) Holub; Folia Geobot. Phytotax. 18: 441 (1983)
= Lateristachys ramulosa (Kirk) Holub; Folia Geobot. Phytotax. 18: 441(1983)
= Lepidotis diffusa (R. Br.) Rothm.; Feddes Repert. 54: 67 (1944)
= Lycopodiella ramulosa (Kirk) B. Øllg.; Opera Bot. 92: 176 (1987)
= Lycopodium diffusum R. Br.; Prodr. 165 (1810)
= Lycopodium laterale var. diffusum (R. Br.) Hook. f. (1848)
= Lycopodium ramulosum Kirk; Trans. & Proc. New Zealand Inst. 11: 456,
t. 19 (B) (1879)
Lycopodiella ericina (Cesati) comb. ined.
Distribution: New Guinea (Arfak Mts.)
Remarks: unverified name, sect. Campylostachys
= Lycopodium ericinum Cesati; Rendic. Accad. Sci. Fis. 16: 28, 31 (1877)
Lycopodiella geometra B. Øllg. & Wind.; Bradea 5: 30, fig. 4A (1987)
Distribution: Brazil (Minas Gerais)
Remarks: sect. Lycopodiella
Lycopodiella glaucescens (C. Presl) B. Øllg.; Opera Bot. 92: 176 (1987)
Distribution [complete]: Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru
Remarks: sect. Campylostachys
= Lycopodium glaucescens C. Presl; Reliq. Haenk. 1: 81 (1825)
= Lycopodium spurium var. glaucescens (C. Presl) Nessel; Die Bärlappgewächse 319 (1939)
Lycopodiella hainanense (Yang) comb. ined.
Distribution: Guangdong, Hainan
Remarks: sect. Campylostachys
= Palhinhaea hainanensis Yang; Bull. Bot. Res. North-East. Forest. Inst. 2 (4): 141 (1982)
Lycopodiella hydrophila (Alderw.) comb. ined.
Distribution: New Guinea
Remarks: unverified name, sect. Campylostachys
= Lycopodium hydrophilum Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 24: 4 (1917)
Lycopodiella inundata (L.) J. Holub; Preslia 36: 21 (1964)
Distribution: Austria, Belgium, England, Bulgaria, former Czechoslovakia, Denmark, Finland,
France, Germany, Ireland, Switzerland, Netherlands, Spain, Hungary, Italy, former Yugoslavia,
Portugal, Norway, Poland, Romania, Sweden, N-European Russia, Baltic States, C-European
Russia, W-European Russia, Belarus, Ukraine, Caucasus/Transcaucasus, W-Siberia, C-Siberia,
Japan, Fujian, asiatic Turkey (Anatolia), St. Pierre & Miquelon, W-Canada, E-Canada (Alberta, British
Columbia, New Brunswick, Labrador, Newfoundland, Nova Scotia, Ontario, Prince Edward Island,
Saskatchewan, Quebec), Alaska, NW-USA, NE-USA (Connecticut, Idaho, Illinois, Indiana, Kentucky, Maine,
Maryland, Massachusetts, Michigan, Minnesota, Montana, New Hampshire, New Jersey, Ohio,
Pennsylvania, New York, Rhode Island, Vermont, Virginia, Washington, West Virginia, Wisconsin)
Trivial names: Sumpfbärlapp, Bog Clubmoss
Remarks: sect. Lycopodiella
= Lepidotis incurva Opiz; Kratos 4: 17 (1819)
= Lepidotis inundata (L.) Boern.; Fl. Deutsche Volk: 285 (1912)
= Lepidotis inundata (L.) Opiz; Seznam 195 (1852)
= Lycopodium inundatum L.; Sp. Pl. 2: 1100 (1753)
= Lycopodium palustre Lam.; Fl. franc. 1 (Méth. anal.): (33) (1779)
= Plananthus inundatus (L.) P. Beauv.; Prodr. Aetheog. 111 (1805)
= Lycopodium anderweitii Nessel
= Lycopodium pallidum Nessel
Lycopodiella iuliformis (L. Underw. & F. Lloyd) B. Øllg.; Opera Bot. 92: 176 (1987)
Distribution [complete]: Venezuela (Roraima), N-Brazil, British Guyana, Surinam
Remarks: sect. Caroliniana
= Lycopodium iuliforme L. Underw. & F. Lloyd; Bull. Torrey Bot. Club 33: 120 (1906)
= Pseudolycopodiella iuliformis (L. Underw. & F. Lloyd) Holub; Folia
Geobot. Phytotax. 18: 442 (1983)
var. tatei (A.C. Sm.) B. Øllg.; Amer. Fern Journal 79: 153 (1989)
Distribution [complete]: Venezuela (Cerro Duida), N-Brazil, British Guyana
= Lycopodium duidae A. C. Smith; Gleason, Bull. Torrey Bot. Club. 58: 311 (1931)
= Lycopodium tatei A. C. Smith; Bull. Torrey Bot. Club 57. 180 (1930)
Lycopodiella lateralis (R. Br.) B. Øllg.; Opera Bot. 92: 176 (1987)
Distribution [complete]: New Zealand (North Isl., South Isl.), Chatham Isl., South Australia, Queensland,
Victoria, Tasmania, New South Wales, New Caledonia, ?Philippines
Remarks: sect. Lateristachys
= Lateristachys lateralis (R. Br.) Holub; Folia Geobot. Phytotax. 18: 441 (1983)
= Lepidotis lateralis (R. Br.) Rothm.; Feddes Repert. 54: 67 (1944)
= Lycopodium consimile Col.; T. N. Z. I. 16, 348 (1884)
= Lycopodium laterale R. Br.; Prodr. 1: 165 (1810)
= Lycopodium halconense Copel.; Philipp. J. Sci. 2 (C): 149 (1907)
= Lateristachys halconensis (Copel.) Holub; Folia Geobot. Phytotax. 18: 441 (1983)
Lycopodiella lehmannii (Hieron.) B. Øllg.; Opera Bot. 92: 176 (1987)
Distribution [complete]: Colombia
Remarks: sect. Campylostachys
= Lycopodium lehmannii Hieron.; Bot. Jahrb. Syst. 34: 574 (1905)
Lycopodiella limosa Chinnock; Austrobaileya 3: 665 (1992)
Distribution [complete]: N-Queensland
Conservation status: IUCN R
Lycopodiella margueritae J.G. Bruce; W.H. Wagner & Beitel, Michigan
Bot. 30: 9 (1991)
Distribution [complete]: Michigan
Conservation status: IUCN V, I
Lycopodiella mariana (K. Mueller) comb. ined.
Distribution: Philippines
Remarks: unverified name, sect. Campylostachys
= Lycopodium marianum K. Mueller; Bot. Zeit. (Berlin) 19 (24): 165. 1861[non Willd. (1810)
Lycopodiella pendulina (Hook.) B. Øllg.; Opera Bot. 92: 176 (1987)
Distribution [complete]: Costa Rica, Colombia, Ecuador, Peru, Bolivia, SE-Brazil, Venezuela
Remarks: sect. Campylostachys
= Lycopodiella eichleri (Fée) B. Øllg.; Opera Bot. 92: 176 (1987)
= Lycopodium cernuum var. eichleri (Fée) Nessel; Archivos de
Botânica do São Paulo 1: 446 (1927)
= Lycopodium cernuum var. pendulinum (Hook.) Baker; Handbook of the
Fern-Allies 23 (1887) [nom. illeg., as "pendulum" and incorrectly attributed to Hooker
= Lycopodium eichleri Fée; Crypt. vasc. Brésil 2: 96, t. 106, f. 4 (1872-73)
= Lycopodium pendulinum Hook.; Icones Plantarum 1: t. 90 (1837)
= Palhinhaea eichleri (Fée) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
= Palhinhaea pendulina (Hook.) Holub; Folia Geobot. Phytotax. 20: 79 (1985)
Lycopodiella prostrata (R.M. Harper) Cranfill; Amer. Fern J. 71: 97 (1981)
Distribution [complete]: SE-USA, S-USA (Arkansas, Florida, Georgia, Louisiana,
Mississippi, North Carolina, South Carolina, Texas)
Trivial names: Feather-stem Club-Moss
Remarks: sect. Lycopodiella
= Lycopodium alopecuroides L. var. pinnatum Chapman
= Lycopodium inundatum L. var. pinnatum Chapman; Fl. South. States 600 (1860)
= Lycopodium pinnatum (Chapman) Lloyd & Underwood; Bull. Torrey Bot.
Club 27: 155. 1900 [non Lam. (1792)
= Lycopodium prostratum Harper; Bull. Torrey Bot. Club 33: 229 (1906)
Lycopodiella pungens (Alderw.) comb. ined.
Distribution: Borneo, Sumbawa
Remarks: unverified name, sect. Campylostachys
= Lycopodium pungens Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 20: 26 (1915)
= Lycopodium alderwereltii Kümm.; Magyar Bot. Lapok 26: 99 (1927) [nom. superfl.
Lycopodiella raiateensis (J. Moore) comb. ined.
Distribution [complete]: Society Isl. (Raiatea)
Remarks: sect. Campylostachys
= Lycopodium raiateense J. Moore; Bernice P. Bishop Mus. Bull. 102: 12 (1933)
Lycopodiella riofrioi (Sodiro) B. Øllg.; Opera Bot. 92: 176 (1987)
Distribution [complete]: Costa Rica, E-Colombia, Venezuela, Ecuador, N-Brazil, Peru, Panama
Remarks: sect. Campylostachys
= Lycopodium pensum Lellinger & Mickel; Proc. Biol. Soc. Wash. 89(61): 717, f. 2 (1977)
= Lycopodium riofrioi Sodiro; Cryptogamae vasculares Quitenses 582 (1893)
Lycopodiella salakensis (Treub) comb. ined.
Distribution: Java
Remarks: unverified name, sect. Campylostachys
= Lycopodium salakense Treub; Ann. Jard. Bot. Buitenzorg 7: 141 (1888)
= Palhinhaea salakensis (Treub) Holub; Folia Geobot. Phytotax. 20: 79 (1985)
Lycopodiella serpentina (Kunze) B. Øllg.; Opera Bot. 92: 176 (1987)
Distribution: New Zealand (North Isl.), New Caledonia, Western Australia, South
Australia, Queensland, Victoria, Tasmania, NE-New South Wales
Remarks: sect. Caroliniana
= Lepidotis drummondii (Spring) Rothm.; Feddes Repert. 54. 66 (1944)
= Lycopodium carolinianum Hook. f.; Fl. N. Z. 2, 54 [non L. (1855)
= Lycopodium drummondii Spring; Mon. Lycop. 2, 35 (1848)
= Lycopodium serpentinum Kunze; Plantae Preissianae 2: 108 (1846)
= Pseudolycopodiella serpentina (Kunze) Holub; Folia Geobot. Phytotax. 18: 442 (1983)
Lycopodiella steyermarkii B. Øllg.; Harling & Andersson, Fl. Ecuador 33:
149, f. 21B (1988)
Distribution [complete]: Panama, Colombia, S-Venezuela, Ecuador
Remarks: sect. Campylostachys
Lycopodiella subappressa J.G. Bruce, W.H. Wagner & Beitel; Michigan
Bot. 30: 4 (1991)
Distribution [complete]: Michigan
Conservation status: IUCN E
Lycopodiella suffruticosa (Alderw.) comb. ined.
Distribution: New Guinea
Remarks: unverified name, sect. Campylostachys
= Lycopodium cernuum var. suffruticosum Alderw.; Bull. Jard. Bot.
Buitenzorg, ser. 2, 24: 5 (1917)
= Lycopodium suffruticosum (Alderw.) Herter; Index Lyc. 43 (1949)
= Palhinhaea suffruticosa (Alderw.) Holub; Folia Geobot. Phytotax. 20: 79 (1985)
Lycopodiella tomentosa (Alderw.) comb. ined.
Distribution: New Guinea
Remarks: unverified name, sect. Campylostachys
= Lycopodium tomentosum Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 2, 24: 6 (1917)
= Palhinhaea tomentosa (Alderw.) Holub; Folia Geobot. Phytotax. 20: 79 (1985)
Lycopodiella torta (Sieber ex L. Underw. & F. Lloyd) comb. ined.
Distribution: Martinique
Remarks: sect. Campylostachys
= Lycopodium tortum Sieber ex L. Underw. & F. Lloyd; Bull. Torrey Bot.
Club 33: 118 (1906)
Lycopodiella trianae (Hieron.) B. Øllg.; Opera Bot. 92: 176 (1987)
Distribution [complete]: Colombia, Ecuador
Remarks: sect. Campylostachys
= Lycopodium cernuum var. trianae (Hieron.) Nessel; Die Bärlappgewächse 358 (1939)
= Lycopodium trianae Hieron.; Bot. Jahrb. Syst. 34: 574 (1905)
= Palhinhaea trianae (Hieron.) Holub; Folia Geobot. Phytotax. 20: 79 (1985)
Lycopodiella x brucei Cranfill; Amer. Fern J. 71: 97 (1981)
Distribution: Distribution unknown
Remarks: sect. Lycopodiella
= Lycopodium x brucei (Cranfill) Lellinger; Amer. Fern Journal 74(2): 63 (1984)
Lycopodiella x copelandii (Eiger) Cranfill; Amer. Fern J. 71: 97 (1981)
Distribution: E-USA
Remarks: sect. Lycopodiella, Lycopodium alopecuroides x Lycopodium appressum
= Huperzia x copelandii Eig.
= Lycopodium chapmanii Underwood ex Maxon; [p. p.
= Lycopodium inundatum L. var. elongatum Chapman
= Lycopodium x copelandii Eiger; Biological Review of the City College 18: 21 (1956)
This page is Copyright ©, 2001, 2002 Brian Swale, Michael Hassler. All rights reserved.
For more information, go to © Copyright page
For each species, information on its distribution by country and sometimes by province, region or state
within a country; conservation status (where applicable), and all synonyms with publishing source (other
latin names by which the species has been known) that we could find, and where applicable whether or
not the given name has been verified; are available on the CDROM version of this page.
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