Family Cheilanthaceae, genus Pellaea;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Pellaea, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

005.00 Class Filicopsida
		Trivial names: Ferns, varens, helechos, fougeres, farnkraut
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
027.000 Order Pteridales
		Statistics: 10 families, 53 genera, 1065-1300 spp. [1358 species and 42 hybrids listed]
035.0000 Family Notholaenaceae
		Statistics: 9 genera, 113-126 spp. [133 species and 8 hybrids listed]
		Revision date: 20.07.2001
		= Hemionitidaceae Pichi Sermolli; Webbia 21: 487 (1966)
036.2000 Subfamily Pellaeoideae
		Revision date: 20.07.2001

036.2001 Pellaea Link; Fil. Spec. Cult. 48, 59 (1841) [nom. cons. Trivial names: Cliff Brakes Statistics: 55-65 spp. SW-USA, Mexico, to Galapagos and Haiti, South Africa, few east to India, China and New Zealand, N-Spain, introduced Azores [67 species and 2 hybrids listed] Revision date: 20.07.2001 = Bakeriopteris O. Ktze.; Rev. Gen. Pl. 2. 807 (1891) = Cincinalis Desvaux; Berl. Mag. 5. 311 (1811) = Crypteris Nuttall; ms. ex Hook., Sp. 2. 149 (1858) = Holcochlaena Baker; Syn. Fil. 153 (1867) = Holodanaea Presl; Suppl. Tent. Pterid. 37 (1845) = Hymenoloma Davenport; Bot. Gazette 21. 262 (1896) = Ormopteris J. Smith; Hist. Fil. 281 (1875) = Pellaeopsis J. Smith; Hist. Fil. 289 (1875) = Platyloma J. Smith; Journ. Bot. 4. 160 (1841) = Pteridella Mett. ex Kuhn; v. Deck. Reis. 3(3): Bot 13 (1879) = Synochlamys Fée; 7 mém. 35, t. 20, f. 4 (1857) = Mildella Trevis.; Rend. Ist. Lombardo II, 9: 810 (1876) [fide Fraser-Jenkins (1997) Pellaea ambigua (Fée) Bak.; Syn. 147 (1867) Pellaea andromedifolia (Kaulf.) Fée; Gen. 129 (1850-52) Trivial names: Coffee Fern Pellaea angolensis Schelpe; Garcia de Orta, Ser. Bot. 3(1): 53 (1976) Pellaea angulosa (Bory) Bak.; Syn. ed. 11. 153 (1874) Pellaea atropurpurea (L.) Link; Fil. Sp. 59 (1841) Trivial names: Purple Cliff Brake Pellaea boivinii Hook.; Sp. Fil. 2: 147. t. 118 A (1858) Pellaea brachyptera (Moore) Baker; Hooker & Baker, Syn. Fil. ed. 2: 477 (1874) Trivial names: Sierra Cliff Brake Pellaea brasiliensis Bak.; Engl. Jahrb. 17. 522 (1893) Pellaea breweri D. C. Eaton; Proc. Amer. Acad. Arts 6: 555 (1865) Trivial names: Brewer's Cliff Brake Pellaea bridgesii Hook.; Sp. 2. 238 (1858) Trivial names: Bridges' Cliff Brake Pellaea calidirupium Brownsey & Lovis; New Zealand J. Bot. 28(3): 197 (1990) Pellaea calomelanos (Swartz) Link; Fil. Sp.: 61 (1841) Trivial names: Hard fern var. leucomelas (Mett. ex Kuhn) J.E.Burrows; South. Afr. Ferns & Fern Allies: 177 (1990) var. swynnertoniana (Sim) Schelpe (1964) Pellaea connectens C. Chr.; Acta Hort. Gothob. 1. 84 t. 18, 19 (1924) Pellaea cordifolia (Sessé & Moc.) A. R. Sm.; Amer. Fern J. 70(1): 26 (1980) Trivial names: Heart-leaved Cliff Brake Pellaea crenata Tryon; Contr. Gray Herb. 143: 68. t.3(E:1-7) (1942) Pellaea cymbiformis Prado; Amer. Fern J. 83(4): 129 (1993) Pellaea doniana (J. Sm.) Hook.; Sp. 2. 137 t. 125 A (1858) Pellaea dura (Willd.) Bak.; Journ. Bot. 327 (1880) var. latipinna Schelpe (1967) var. holstii (Hieron.) Schelpe (1967) Pellaea falcata (R. Br.) Fée; Gen. 129 (1850-52) Trivial names: Sickle fern Pellaea flavescens Fée; Cr. vasc. Br. 1. 44 t. 22 f. 2 (1869) Pellaea gastonyi Windham; Contr. Univ. Michigan Herb. 19: 40 (1993) Pellaea glabella Mett. ex Kuhn; Linnaea 36: 87 (1869) Trivial names: Smooth Cliff Brake ssp. glabella Mett. ex Kuhn; Linnaea 36: 87 (1869) ssp. simplex (Butters) A. & D. Löve; Taxon 26: 325 (1977) ssp. missouriensis (G.J. Gastony) Windham; Contr. Univ. Michigan Herb. 19: 39 (1993) ssp. occidentalis (E.E. Nelson) Windham; Contr. Univ. Michigan Herb. 19: 39 (1993) Pellaea gleichenioides (Gardn.) Christ; Bull. Boiss. II. 2. 545 (1902) Pellaea intermedia Mett. ex Kuhn; Linnaea 38: 84 (1869) Trivial names: Intermediate Cliff Brake Pellaea intramarginalis (Kaulf.) J. Sm.; Cat. Kew Ferns 4 (1856) var. serratifolia Hook. & Baker; Syn. Fil. 149 (1867) Pellaea leonardii (Maxon) comb. ined. Pellaea longipilosa Bonap.; N. Pt. 16. 33 (1924) Pellaea lyngholmii Windham; Contr. Univ. Michigan Herb. 19: 40 (1993) Pellaea mairei Brause; Hedwigia 64. 201 t. 4 C (1914) Pellaea malabarica Madhus. & Jyothi ex Geev.; Indian Fern J. 9(1-2): 39, 1992 publ. 31 Jan. (1993) Pellaea marginata Eat.; U. S. and Mex. Bound. Surv. Bot. 233 (1859) Pellaea mucronata (D. C. Eaton) D. C. Eaton; Emory, Rep. U.S. Mex. Bound. 2(1): 233 (1859) Trivial names: Bird's Foot Cliff Brake ssp. mucronata (D. C. Eaton) D. C. Eaton; Emory, Rep. U.S. Mex. Bound. 2(1): 233 (1859) ssp. californica (Lemmon) Windham; Contr. Univ. Michigan Herb. 19: 42 (1993) Pellaea myrtillifolia Mett. ex Kuhn; Linn. 36. 85 (1869) Pellaea nana (Hook.) Bostock; Fl. Australia 48: 708 (1998) Pellaea nitidula (Hook.) Wall. ex Bak.; Syn. 149 (1867) Pellaea notabilis Maxon; Contr. U. S. Nat. Herb. 10. 500 (1908) Pellaea ovata (Desv.) Weath.; Contr. Gray Herb. 114: 34 (1936) Trivial names: Ovate-leaved Cliff Brake Pellaea pallida Bak.; Syn. 478 (1874) Pellaea paradoxa (R. Br.) Hook.; Sp. 2. 135 t. 111 A (1858) Pellaea patula (Baker) Ching; Acta Phytotax. Sin. 10: 302 (1965) Pellaea paupercula (Christ) Ching; Bull. Fan Mem. Inst. 2. 203 (1931) Pellaea pectiniformis Bak.; Syn. ed. 11. 147 (1874) Pellaea pinnata (Kaulf.) Prantl; Engl. Jahrb. 8. 418 (1882) Pellaea pringlei Dav.; Gard. and Forest 4. 555 t 88 (1891) Pellaea prolifera Schelpe; Bull. Jard. Bot. Nation. Belg. 38: 193, f. 1 (1968) Pellaea pteroides (L.) Prantl; Engl. Jahrb. 3. 420 (1882) Pellaea quadripinnata (Forssk.) Prantl; Engl. Jahrb. 8. 420 (1882) Pellaea riedelii Bak.; Ann. Bot. 6. 213 (1891) Pellaea rinorensis Alderw.; Bull. Dépt. agric. Ind. néerl. 18. 11 (1908) Pellaea robusta (Kuntze) Hook.; Sp. 2. 147 (1858) Pellaea rotundifolia (Forst. f.) Hook.; Sp. 2. 136 (1858) Trivial names: New Zealand Cliff brake, Button fern. Pellaea rufa A.F.Tryon; Ann. Missouri Bot. Gard. 42: 101. t.7 (1955) Pellaea sagittata (Cav.) Link; Fil. Spec. 60 (1841) Pellaea schweinfurthii Hier. ex Diels; Nat. Pfl. 1(4). 267 (1899) Pellaea smithii C. Chr.; Acta Hort. Gothob. 1. 84 t. 18 (1924) Pellaea straminea Ching; Bull. Fan Mem. Inst. 2. 203 t. 17 (1931) Pellaea striata (Desv.) C. Chr.; Dansk Bot. Ark. 7. 117 t. 44 f. 12-15 (1932) Pellaea ternifolia (Cav.) Lk.; Fil. Spec. 59 (1841) Trivial names: Ternate-leaved Cliff Brake ssp. ternifolia (Cav.) Lk.; Fil. Spec. 59 (1841) ssp. arizonica Windham; Contr. Univ. Michigan Herb. 19: 42 (1993) ssp. villosa Windham; Contr. Univ. Michigan Herb. 19: 43 (1993) Pellaea timorensis Alderw. Pellaea tomentosa Bonap.; N. Pt. 6. 64 (1917) Pellaea trichophylla (Baker) Ching; Acta Phytotax. Sin. 10: 302 (1965) Pellaea tripinnata Bak.; Journ. Linn. Soc. 25. 350 (1889) Pellaea truncata Goodd.; Muhlenbergia 8: 94 (1912) Trivial names: Spiny Cliff Brake Pellaea viridis (Forssk.) Prantl; Bot. Jahrb. Syst. 3: 420 (1882) Trivial names: Green Cliff Brake var. glauca (Sim) Schelpe & N.C.Anthony; Contr. Bolus 155 (1982) var. macrophylla (Kunze) Schelpe & N.C.Anthony; Contr. Bolus Herb. 10: 155 (1982) var. involuta (Swartz) Schelpe (1967) var. obscura N.C.Anthony; Contr. Bolus Herb. 10: 154 (1982) Pellaea woodfordii (Wright) C. Chr.; Ind. Suppl. 54 (1913) Pellaea wrightiana Hook.; Sp. Fil. 2: 142 (1858) Trivial names: Wright's Cliff Brake Pellaea x glaciogena W.H.Wagner; Pray & A. R. Sm., Madrono 30(2): 70 (1983) Remarks: Pellaea bridgesii x Pellaea mucronata Pellaea x oaxacana Mickel & Beitel; Pterid. Fl. Oaxaca (Mem. New York Bot. Gard. 46): 271 (1988) Remarks: Pellaea ovata x sagittata Pellaea yunnanensis Ching; Acta Phytotax. Sin. 20(2): 235 (1982)
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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