Family Polypodiaceae, genus Polypodium;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Polypodium, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

005.00 Class Filicopsida
		Trivial names: Ferns, varens, helechos, fougeres, farnkraut
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
050.000 Order Polypodiales
		Statistics: 9 families, 82 genera, 1996-2140 spp. 
		(2293 species and 31 hybrids listed]
		= Order Davalliales
058.0000 Family Polypodiaceae Bercht. & J.C. Presl; Prirozen. Rostlin. 1: 272 (1820) s. str.
		Statistics: 19 genera, 397-446 spp. 
		[450 species and 19 hybrids listed]
		Revision date: 20.07.2001
		Remarks: according to PCR results Polypodiaceae s. lat. is polyphyletic. 
		There are two choices: either include Grammitidaceae, Pleurosoriopsidaceae and 
		Loxogrammaceae in Polypodiaceae, or separate Drynariaceae as family, as done here.
		= Platyceriaceae Ching; Sunyatsenia 5 (1940)
		= Polypodiaceae subfam. Lepisorioideae Ching; Acta Phytotax. Sin. 16(3): 17 (1978) [nom. nov.
		= Polypodiaceae subfam. Pleopeltidoideae Nayar
		= Polypodiaceae tribus Aglaomorpheae Subh. Chandra; Brenesia 18: 175 (1980) 
		[without latin descr. or type, et in Fern Gaz. 12(4): 226 (1982), with latin descr. and type
		= Polypodiaceae tribus Colyseae V.G.Tu; Novosti Sist. Vyssh. Rast. 18: 29 (1981)
		= Polypodiaceae tribus Pleopelteae V.G.Tu; Novosti Sist. Vyssh. Rast. 18: 38 (1981)
058.1000 Subfamily Polypodioideae
		Remarks: many authors treat the following genera as one large genus Polypodium s. lat.
		= Tribus Polypodieae

058.1008 Polypodium L.; Sp. Pl. 2: 1082 (1753) [s. str. Statistics: 150-170 spp., mainly American tropics, very few in Old World [181 species and 7 hybrids listed]. Revision date: 20.07.2001 Trivial names: Tüpfelfarn = Anaxetum Schott; Gen. Fil. t. 1 (1834) = Catenularia Zippel ms. ex Mett.; Polyp. 37 (1857) = Cephalosorium Moore; Ind. Fil. L\XV (1857) = Chilopteris Presl; Tent. Pterid. 208 (1836) = Chrysopteris Link; Fil. Sp. 120 (Sept. 1841) [nom. superfl. = Clathropeltis Kuhn; v. Decken's Reisen 88. Bot. 51 (1879) = Coelopteris A. Braun ms. ex Mett.; Polyp. 50 (1857) = Cyrtomiphlebium Hooker; Spec. Fil., 6. 15 (1863) = Cystidium J. Sm. ms. ex Moore; Ind. 278 (1861) = Diagramma Blume; Enum. pl. Jav. 114 (1828) = Dicranopteris BI. ex Endl.; Gen. Suppl. 1. 1346 (1841) = Dictyogramma Presl; Epim. Bot. 148 (1849-51) = Dictyophlebia Gris.; Abh. Ges. Gött. 7. 283, 286 (1857) = Eleutherophlebia Gris.; Abh. Gött. Ges. Wiss. 7. 282. 285 (1857) = Goniolepicystis Diels; Nat. Pflanzenfam. 14. 323 (1899) = Lepicystis J. Sm.; Lond. Journ. Bot. 1: 195 (1842) = Leptoselliguea C. Chr.; Gardens Bull. S. S. 7. 309 (1934) = Leptostegia Zipp. ms. ex Mett.; Pol. 50 (1857) = Marginaria Bory; Dict. Class. d'Hist. Nat. 6: 587 (1824) = Marginariopsis Moore; Index LXXIII (1857) = Micropteris J. Smith; Hist. Fil. 186 (1875) = Microterus Presl; Epim. bot. 124 (1849-51) = Phlebiolepicystis Diels; Nat. Pfl. 1(4). 321 (1899) = Phlebodiopsis Moore; Ind. Fil. LXXVII (1857) = Phyllitidis J. Smith; Journ. Bot. 3. 397 (1841) = Phymatodes Presl; Tent. Pter. 195 (1836) = Pleurogonium Presl; Tent. Pterid. 187 (1836) = Pseudovittaria C. Chr.; Dansk Bot. Ark. 6. no. 3. 45 (1929) = Synammia C. Presl; Tent. Pterid. 212 (1836) = Trichocalymma Zenker ms. ex Kze.; Linn. 24: 251 (1851) Polypodium abitaguae Hook.; Sp. 4: 206 (1862) Polypodium adnatum Kunze; Kl. Linn. 20: 395 (1847) Polypodium aequale Maxon; Contr. U.S. Natl. Herb. 8: 273 (1903) Polypodium aichmophyllum Alston; Journ. of Bot. 77: 287 (1939) Polypodium alansmithii R.C. Moran; Ann. Missouri Bot. Gard. 77: 845 (1990) Polypodium alavae A.R. Sm.; Amer. Fern J. 70: 21 (1980) Polypodium alborufulum Brade; Arq. Jard. Bot. Rio Janeiro 11: 29. t.102 (1951) Polypodium amorphum Suksdorf; Werdenda 1: 16 (1927) Trivial names: Irregular Polypody Polypodium angustialatum Bonap.; N. Pt. 14. 151 (1924) Polypodium antillense Maxon; Proc. Biol. Soc. Wash. 43. 83 (1930) Polypodium apagalolepis Sota; Rev. Mus. La Plata, Bot. 10: 133, f. 22 (1967) Polypodium appalachianum Haufler & Windham; Amer. Fern J. 81: 18 (1991) Trivial names: Appalachian Polypody Polypodium arcanum Maxon; Proc. Biol. Soc. Washington 51 38 (1938) Polypodium argentinum Maxon; Contr. U. S. Nat. Herb. 17. 688 (1916) Polypodium argyrolepis Sod.; Cr. vasc. quit. 349 (1839) Polypodium arthropodium Fée; 8 mém. 88 (1857) Polypodium attenuatum Humb. & Bonpl. ex Willd.; Sp. Pl. 5: 191 (1810) Polypodium aturense Maury; Jour. de Bot. 3: 129. f. 3 (1889) Polypodium balaonense Hieron; Bot. Jahrb. Syst. 34: 529 (1905) Polypodium balliviani Rosenst.; Fedde Repert. 9. 344 (1911) Polypodium bombycinum Maxon; Contr. U.S. Natl. Herb. 17: 592 (1916) Polypodium bradei Sota; Rev. Mus. La Plata, Bot. 9: 266, f. 7, 8A, B (1965) Polypodium bryopodum Maxon; Contr. U. S. Nat. Herb. 17. 668 (1916) Polypodium caceresii Sod.; Cr. vasc. quit. 360 (1893) Polypodium californicum Kaulfuss; Enum. Filic. 102 (1824) Trivial names: California Polypody Polypodium calirhiza S.A. Whitmore & A.R. Smith; Madrono 38: 235 (1991) Polypodium cambricum L.; Sp. Pl.: 1086 (1753) Trivial names: Südlicher Tüpfelfarn ssp. cambricum L.; Sp. Pl.: 1086 (1753) ssp. serrulatum (Schlecht. ex Arcang.) Pichi-Serm.; Webbia 40(1): 49 (1986) ssp. azoricum (Vasc.) E.Nardi; Webbia 33(2): 431 (1979) ssp. macaronesicum (Bobrov) Fraser-Jenkins; Greuter & al., Med-Checklist 1: n 056-24-980-16-56 (1981) Polypodium camptocarpum (Fée) C. Chr.; Ind. 186 (1905), 515 (1906) Polypodium catharinae Langsd. & Fisch.; Ic. Fil. 9 t. 9 (1810) Polypodium caudiceps (Moore) Nichols.; Ill. Dict. Gard. 4: 592 (1888) Polypodium chiapense A.M. Evans & A.R. Sm.; Amer. Fern J. 70: 23 (1980) Polypodium chionolepis Sod.; Cr. vasc. quit. 375 (1893) Polypodium chnoophorum Kze.; Flora 1839(1), Beibl. 34 (1839) Polypodium christensenii Maxon; Contr. U.S. Nat. Herb. 13: 10 (1909) Polypodium cinerascens Lindm.; Ark. för Bot. 1: 238. t. 11, f. 6 (1903) Polypodium collinsii Maxon; Contr. U. S. Nat. Herb. 17. 583 t. 41 (1916) Polypodium colysoides Maxon & Copel.; Univ. Calif. Publ. Bot. 19: 292 (1941) Polypodium conterminans Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd. ser. 5, 1: 201 (1849) Polypodium cryptocarpon Fée; 8 mém. 88 (1857) Polypodium crystalloneuron Rosenst.; Fedde Repert. 11. 67 (1912) Polypodium dasypleuron Kze.; Linn. 9: 43 (1834) Polypodium decorum Brackenr.; Expl. Exp. 16: 7, t. 2, f. 2 (1854) Polypodium decrescens Christ; Ann. Jard. Buit. II. 4: 35 (1904) Polypodium diplotrichum Mickel & Beitel; Mem. New York Bot. Gard. 46: 297 (1988) Polypodium dissimile L.; Syst. Nat. ed. 10, 2: 1325 (1759) Polypodium dulce Poir. in Lam.; Encycl. 5: 523 (1804) Polypodium eatonii Baker in Hook. & Baker; Syn. Fil. ed. 2, 511 (1874) Polypodium echinolepis Fée; Mém. Foug. 8: 87 (1857) Polypodium eperopeutes Mickel & Beitel; Mem. New York Bot. Gard. 4: 298 (1988) Polypodium espinosae Weatherby; Contr. Gray Herb. 85. 15 t. 2 (1929) Polypodium exsul Mett. ex Kuhn; Linn. 36. 138 (1869) Polypodium fauriei Christ; Bull. Boiss. 4: 672 (1896) Polypodium feuillei Bertero; Mercurio Chileni, 745. Julio (1829) var. ibanezii Looser Polypodium fimbriatum Maxon; Contr. U. S. Nat. Herb. 17. 596 (1916) Polypodium fissidens Maxon; Contr. U.S. Natl. Herb. 8: 275 (1903) Polypodium flagellare H. Christ; Bull. Herb. Boissier 4: 660 (1896) Polypodium fournieri C. Chr.; Ind. 528 (1966) Polypodium fraternum Schltdl. & Cham.; Linnaea 5: 608 (1830) Polypodium fraxinifolium Jacq.; Collectanea 3: 187 (1791) Polypodium friedrichsthalianum Kunze; Farrnkräuter 2: 55 (1850) Polypodium funckii Mett.; Pol. 57, n. 72 (1857) Polypodium furfuraceum Schltdl. & Cham.; Linnaea 5: 607 (1830) Polypodium furfuratum Sessé & Moc.; Fl. Mexic. od.2. 239 (1894) Polypodium fuscopetiolatum A.R. Sm.; Amer. Fern J. 70: 24 (1980) Polypodium gauthieri (Fée) C. Chr.; Ind. 326 (1905), 529 (1906) Polypodium giganteum Desv.; Mem. Soc. Linn. Paris 6: 236 (1827) Polypodium glaberulum Mickel & Beitel; Pterid. Fl. Oaxaca (Mem. New York Bot. Gard. 46): 300 (1988) Polypodium glandulosissimum Brade; Arch. Inst. Biol. Vog. Rio Janeiro 1: 230. f.6. t.5(2) (1935) Polypodium glycyrrhiza D. C. Eaton; Amer. J. Sci. Arts, ser. 2, 22: 138 (1856) Trivial names: Licorice Fern Polypodium grandiceps Nichols.; Ill. Dict. Gard. 4: 592 (1888) Polypodium guilleminianum Fourn.; Mex. pl. 77, t. 2 (1872) Polypodium gyroflexum Christ; Bull. Boiss. 6: 994 (1898) Polypodium hahnii Fourn.; Mex. pl. 1: 79 (1872) Polypodium haitiense Urban; Symb. Ant. 9. 370 (1925) Polypodium hartwegianum Hook. in Benth.; Pl. Hartw. 54 (1840) Polypodium hassleri Christ; Bull. Boiss. II. 7. 923 (1907) Polypodium hesperium Maxon; Proc. Biol. Soc. Wash. 13: 200 (1900) Trivial names: Western Polypody Polypodium hirsutissimum Raddi; Opusc. sci. Bol. 3: 286 (1819) Polypodium hispidulum Bartlett; Proc. Amer. Acad. Arts 43: 48 (1907) Polypodium hoehnei Sampaio; Comm. Linh. telegr. Matto grosso ao Amazonas Publ. no. 33. 26. t. 4 (1916) Polypodium imbeanum Brade; Arch. Mus. Nac. Rio 34. 115 t. 1 f. 1 (1932) Polypodium insulare Brade; Bradea 1: 7, t. 3 (1969) Polypodium insularum (C. Morton) Sota; Rev. Mus. La Plata, Bot. 10: 152, f. 1, 4, 12, 15, 16 (1967) Polypodium interjectum Shivas; J. Linn. Soc. Bot. 58: 29 (1961) Trivial names: Gesägter Tüpfelfarn, Mittlerer Tüpfelfarn Polypodium intermedium Colla; Mem. Ac. Torino 39: 51, t. 74 (1837) [non Willd. (1810) ssp. intermedium Colla; Mem. Ac. Torino 39: 51, t. 74 (1837) [non Willd. (1810) ssp. masafueranum C. Chr. & Skottsb. Polypodium itatiayense Rosenst.; Hedwigia 64. 369 (1915) Polypodium kuhlmannii Sampaio; Comm. Linh. telegr. Mato Grosso ao Amazonas Publ. 33. 27 t. 5 (1916) Polypodium kunzeanum C. Chr.; Index Filic. 536 (1906) Polypodium laevigatum Cav.; Descr. Pl. 244 (1802) Polypodium lasiopus Kl.; Linn. 20: 393 (1847) Polypodium latissimum R.C.Moran & B. Ollg.; Nordic J. Bot. 15(2): 184 (1995) Polypodium lepidopteris (Langsd. & Fisch.) Martius; Reise 2. 554 (1828) Polypodium lepidotrichum (Fée) Maxon; Contr. U. S. Nat. Herb. 17. 591 (1916) Polypodium lepidum Brause; Notizbl. Bot. Gart. Berlin-Dahlem 8. 139 (1922) Polypodium leucosticton Kze.; Kl., Linn. 20: 380 (1847) Polypodium liebmannii C. Chr.; Ind. 539 (1906) Polypodium lindenianum Kunze; Farrnkräuter 2: 83 (1851) Polypodium longepinnulatum E. Fourn.; Mexic. Pl. 1: 77 (1872) Polypodium longipetiolatum Brade; Rodriguesia 3(9): 115 (1937) Polypodium loriceum L.; Sp. Pl. 1086 (1743) Polypodium loriciforme Rosenst.; Repert. Spec. Nov. Regni Veg. 22: 17 (1925) Polypodium macedoi Brade; Arq. Jard. Bot. Rio Janeiro 11: 30. t.99 11(3) (1951) Polypodium macrolepis Maxon; Contr. U.S. Natl. Herb. 17: 584 (1916) Polypodium madrense J. Sm. in Seem.; Bot. Voy. Herald 338 (1854) Polypodium maritimum Hieron.; Bot. Jahrb. Syst. 34: 527 (1904) Polypodium martensii Mett.; Pol. 61. n. 83 (1857) Polypodium martii Colla; Herb. Pedem. 6: 230 (1836) Polypodium masafuerae Phil.; Linn. 29: 107 (1857) Polypodium maxonii C. Chr.; Ind. 543 (1906) Polypodium megalolepis Maxon & Morton; Bull. Torrey Bot. Cl. 66: 39. f.1 (1939) Polypodium menae Mille; Revista Col. Nacion. Rocafuerte 9: 200 (1927) Polypodium mexiae Copel.; Univ. Calif. Publ. Bot. 17. 33 t. 8 (1932) Polypodium minarum Weath.; Contr. Gray Herb. 166: 78 (1947) Polypodium mindense Sod.; Cr. vasc. quit. 348 (1893) Polypodium minutissimum J. W. Moore; Bishop Mus. Bull. 102. 11 (1933) Polypodium mixtum Sod.; Cr. vasc. quit. 339 (1893) Polypodium monoides Weath.; Contr. Gray Herb. 166: 78 (1947) Polypodium monosorum Desv.; Berl. Mag. 5: 319 (1811) Polypodium montigenum Maxon; Field Mus. Nat. Hist., Bot. Ser. 17: 306 (1938) Polypodium moricandii Mett.; Pol. 87, n. 152, t. 1, f. 47-48 (1857) Polypodium moritzianum Link; Fil. Sp. 126 (1841) Polypodium mosenii C. Chr.; Bot. Tidsskr. 25: 78 (1902) Polypodium murorum Hook.; Icon. Pl. 1: t. 70 (1837) Polypodium myriolepis H. Christ; Bull. Herb. Boissier 4: 661 (1896) Polypodium otites L.; Sp. Pl. 2: 1085 (1753) Polypodium oxylepis C. Chr.; Ind. 550 (1906) Polypodium panorense C. Chr.; Dansk Bot. Ark. 6 no. 3. 97 (1929) Polypodium paraguayense Bak.; Journ. Bot. 301 (1878) Polypodium pellucidum Kaulf.; Enum. 101 (1824) f. opacum (Hillebr.) D. D. Palmer var. vulcanicum Skottsberg var. acuminatum D. D. Palmer; [ined. Polypodium picoti Carr.; Rév. Hort. 206, f. 62 (1886) Polypodium pinnatissimum R.C. Moran; Novon 2: 135 (1992) Polypodium platylepis Mett. ex Kuhn; Linn. 36: 137 (1869) Polypodium plebejum Schltdl. ex Cham.; Linnaea 5: 607 (1830) Polypodium plectolepidioides Rosenst.; Repert. Spec. Nov. Regni Veg. 10: 278 (1912) Polypodium pleopeltidis Fée; Cr. vasc. Br. 1: 86, t. 28 (1869) Polypodium plesiosorum Kunze; Linnaea 18: 313 (1845) Polypodium pleurosorum Kunze ex Mett.; Abh. Senckenberg. Vaturf. Ges. 2: 101 (1856) Polypodium ptilorhizon H. Christ; Bull. Herb. Boissier sér. 2, 5: 6 (1905) Polypodium puberulum Schltdl. & Cham.; Linnaea 5: 607 (1830) Polypodium pycnocarpum C. Chr.; Ind. 326 (1905), 557 (1906) var. buchtienii (Rosenst.) R. M. Tryon & Stolze; Fieldiana, Bot. 32: 136 (1993) Polypodium pyrrholepis (Fée) Maxon; Contr. U.S. Natl. Herb. 17: 593 (1916) Polypodium quitense Bak.; Journ. Bot. 165 (1877) Polypodium remotum Desv.; Mem. Soc. Linn. Paris 6: 232 (1827) Polypodium rhachipterygium Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd. ser. 5, 1: 191 (1849) Polypodium rhodopleuron Kunze; Linnaea 18: 315 (1845) Polypodium rimbachii Sod.; Cr. vasc. quit. 356 (1893) Polypodium rosei Maxon; Contr. U.S. Natl. Herb. 17: 594 (1916) Polypodium rupicolum Brade; Arch. Inst. Biol. Veg. Rio Janoiro 1: 228 (1935) Polypodium rusbyi Maxon; Contr. U. S. Nat. Herb. 17. 570 (1916) Polypodium ryanii Kaulf.; Enum. 92 (1824) Polypodium sanctae-rosae (Maxon) C. Chr.; Index Filic., Suppl. 1, 62 (1913) Polypodium saximontanum Windham; Contr. Univ. Michigan Herb. 19: 47 (1993) Polypodium schiffneri Christ; Denkschr. Akad. Wien 79. 33 (1907) Polypodium scouleri Hook. & Grev.; Ic. Fil. t. 56 (1828) Trivial names: Coast Polypody Polypodium scutulatum Sod.; Sert. Fl. Ecuad. II. 29 (1908) Polypodium segregatum Bak.; Syn. 510 (1874) Polypodium sessilifolium Desv.; Mém. Soc. Linn. P.,is 6: 238 (1827) Polypodium sibiricum Siplivinsky; Novosti Sist. Vyssh. Rast. 11: 329 (1974) Trivial names: Siberian Polypody Polypodium someyae Yatabe; Bot. Mag. Tokyo 5: 245, t. 27 (1891) Polypodium squamatum L.; Sp. 2: 1086 (1753) Polypodium steirolepis C. Chr.; Amer. Fern Journ. 7. 33 (1917) Polypodium subandinum Sod.; Cr. vasc. quit. 348 (1893) Polypodium subinaequale Christ; Ark. för Bot. 915. 2 (1910) Polypodium subpetiolatum Hook.; Benth., Pl. Hartw. 54 (1840) Polypodium subvestitum Maxon; Contr. U. S. Nat. Herb. 17. 566 (1916) Polypodium subviride Lellinger; Proc. Biol. Soc. Wash. 98(2): 389 (1985) Polypodium teresae Maxon; Contr. U.S. Nat. Herb. 8: 274. t. 61, f. 2 (1903) Polypodium tobagense C. Chr.; Amer. Fern Journ. 7. 35 (1917) Polypodium trinidadense Brade; Arch. Inst. Biol. Vog. Rio Janeiro 3: 4. t.3(2)9 4(7) p 6(7-9) (1936) Polypodium trinitensis Jenm.; Ferns Brit. W. Ind. Guiana 278 (1908) Polypodium triseriale Sw.; J. Bot. (Schrader) 1800(2): 26 (1801) var. gladiatum (Kuhn) Proctor; Amer. Fern J. 72(4): 114 (1982) Polypodium truncatulum Rosenst.; Fedde Repert. 9. 343 (1911) Polypodium tweedianum Hook.; Ic. pl. t. 86 (1837) Polypodium ursipes Moritz ex C. Chr.; Index Filic. 572 (1906) Polypodium vagans Mett.; Ann. sc. nat. V. 2: 255 (1864) Polypodium villagranii Copel.; Univ. Calif. Publ. Bot. 19: 292. t.44 (1941) Polypodium virginianum L.; Sp. Pl. 2: 1085 (1753) Trivial names: Rock Polypody Polypodium vulgare L.; Sp. PI.: 1085 (1753) Trivial names: Tüpfelfarn, Engelsüss var. eatonii Bak.; Phil. Trans. Roy. Soc. Lond. 168: 23 (1879) Polypodium wagneri Mett.; Ann. Sci. Nat. Bot. ser. 5, 1: 255 (1864) Polypodium wangii Ching; Bull. Fan Mem. Inst. Biol. ser. 2. 1: 307 (1949) Polypodium warmingii C. Chr.; Bot. Tidsskr. 25: 77 (1902) Polypodium x aspidiolepis Baker; Journ. Bot. 26 (1887) Remarks: Polypodium friedrichsthalianum x Polypodium thyssanolepis Polypodium x font-queri Rothm. in Cadev. & Font-Quer; Fl. Catalun. 6: 353 (1937) Remarks: Polypodium serratum x vulgare Polypodium x huancayanum Kunkel; Biota 6: 157, f. c, d (1966) Remarks: Polypodium lasiopus x mollendense Polypodium x mantoniae (Schidlay) Shivas; Brit. Fern Gaz. 10: 152 (1970) Remarks: Polypodium interjectum x vulgare, Polypodium x pinnatisectum (Brade) L.D. Gómez; Amer. Fern J. 66(1): 28 (1976) Remarks: polypodioides agg., Polypodium friedrichsthalianum x Polypodium furfuraceum Polypodium x rothmaleri Shivas; Brit. Fern Gaz. 10: 152 (1970) [nom. nov. Remarks: Polypodium australe x interjectum, Polypodium australe x vulgare ssp. prionodes Polypodium x schneideri G. Schneid.; Book Choice Ferns 3: 209 cum fig (1894) Remarks: Polypodium aureum x vulgare var. elegantissimum
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