Family Lomariopsidaceae, genus Polystichum;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Polystichum, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

  Polystichum species A to I.
  Polystichum species J to Z plus hybrids Here.


005.00 Class Filicopsida
		Trivial names: Ferns, varens, helechos,  fougeres, farnkraut.
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
047.000 Order Dryopteridales
		Statistics: 3 families, 30 genera, 1421-1752 spp. 
		[1989 species and 158 hybrids listed]
049.0000 Family Dryopteridaceae Ching; Acta Phytotax. Sin. 10: 1 (1965) s. str.
		Statistics: 21 genera, 672-939 spp. 
		[1162 species and 147 hybrids listed]
		Revision date: 20.07.2001
		= Aspidiaceae Frank; Leunis, Syn. Pflanzenfam. ed. 2, 3: 1469 [nom. illeg.
		= Dryopteridaceae tribus Stenolepieae H.S. Kung; Proc. Int. Symp. Syst. Pteridol. 
		(eds. K.H. Shing & K.U. Kramer): 172 (1989)
049.0200 Tribus Dryopterideae

049.0217 Polystichum Roth; Tent. Fl. Germ. 3: 31, 69 (1799) [nom. cons. Statistics: 200-300 spp. nearly worldwide, 55 in trop. America [348 species and 57 hybrids listed]. This used to be a collective "omnibus genus", and reflecting that fact and that taxonomists still have to do much more work on these species, many currently listed species may better belong in other genera. China has too many (probably invalid) microspecies, and many of the Malesian species may turn out to be synonyms, too. Revision date: 20.07.2001 Trivial names: Schildfarn = Acropelta Nakai; Bull. Nat. Sci. Mus. Tokyo 33: 5 (1953) = Aetopteron Ehrhart; Beitr. 148 (1789), ex House, Amer. Fern Journ. 10. 88 (1920) = Cyrtogonellum Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 327 (1938) = Cyrtomidictyum Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 182 (1940) = Hemesteum Léveillé; Flore du Kouy-tcheou 496 (1915) [non Hemestheum Newman (1851) = Hemigonum J. Smith; Hist. Fil, 204 (1875) = Hypopeltis Michx.; Fl. Bor. Am. 2: 226 (1803), in obs., ex Bory, Bél., Voy. Bot. 2: 63 (1833) = Papuapteris Christensen; Brittonia 2: 300 (1937) = Plecosorus Fée; Mem. Fam. Foug. 5 (Gen. Fil.) 150 (1852) = Porpaea Presl ms. ex Ettingsh.; Farnkr. 174 (1865) Polystichum acanthophyllum (Franch.) Christ; Bull. Soc. Brot. France 52. Mem. 1.: 30 (1905) Polystichum acrostichoides (Michx.) Schott; Gen. Fil., pl. 9 (1834) Trivial names: Christmas Fern Polystichum aculeatum (L.) Roth; Tent. Fl. Germ. 3: 79 (1799) Trivial names: Dorniger Schildfarn, Gelappter Schildfarn var. batjanense Christ; Farnkr. 238 (1897) var. samoensis Luerss.; Fil. Graeff. 174 (1871) Polystichum aculeolatum Fée; Cr. vasc. Br. 1: 128 t. 38. f. 2 (1869) Polystichum acutidens Christ; Bull. Ac. Géogr. Bot. Mans 259 (1902) Polystichum acutipinnulum Ching & K. H. Shing; Wuyi Sci. J. 1: 9(1981) Polystichum adungense H.S.Kung & L.B.Zhang ex Ching & Fraser-Jenk.; Acta Phytotax. Sin. 32(4): 370 (1994) Polystichum alatawshanicum C.Y.Yang; Fl. Xinjiangensis 1: 302 (1992 publ. 1993) Polystichum alcicorne (Bak.) Diels; Engl. Jahrb. 29: 194 (1900) Polystichum alfarii (H. Christ) Barrington; Rhodora 94: 328 (1992) Polystichum alpinum Rosenst.; Fedde Repert. 12. 171 (1913) Polystichum alticola Schelpe & N.C.Anthony; Contr. Bolus Herb. 10: 144 (1982) Polystichum ambiguum Maxon; Contr. U. S. Nat. Herb. 16. 49 t. 27 (1912) Polystichum andersonii Hopkins; Amer. Fern J. 3: 116, pl. 9 (1913) Trivial names: Anderson's Sword Fern Polystichum andinum Phil.; Linn. 29: 108 (1857) Polystichum anomalum (Hook. & Arn.) J. Sm.; Ferns Brit. For. 151 (1866) Polystichum aquifolium Underw. & Maxon; Bull. Torr. Cl. 29: 584 (1902) Polystichum aquilinum Copel.; Philip. Journ. Sc. 81: 20 (1952) Polystichum archboldii Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942) Polystichum atkinsonii Bedd.; Ferns Brit. India Suppl. 14. t 362 (1876) Polystichum attenuatum Tag. & lwatsuki; Acta Phytotax. Geobot. 23: 113, f. 9 (1968) Polystichum auriculum Ching; Bull. Fan Mem. Inst. Biol. ser. 2. 1: 309 (1949) Polystichum australiense Tindale; S.Austr. Nat. 28: 31. 1.1 (1954) Polystichum australium Copel.; Occ. Pap. Bishop Mus. 14: 57. t.8 (1938) Polystichum bakerianum (Atkins.) Diels; Nat. Pfl. 1(4): 191 (1899) Polystichum bamlerianum Rosenst.; Fedde Repert. 10. 330 (1912) Polystichum baoxingense Ching & H.S.Kung; Acta Bot. Boreal-Occid. Sin. 9(4): 271 (1989) Polystichum basipinnatum (Bak.) Diels; Nat. Pfl. 1(4): 189 (1899) Polystichum biaristatum (BI.) Moore; Ind. Fil. 86 (1856) Polystichum bifidum Ching; Icon. Fil. Sin. 5: t.242 (1958) Polystichum bissectum C. Chr.; Ind. Suppl. 64 (1913) Polystichum blepharistomum Copel.; Phil. Journ. Sci. 1. Suppl. II. 145 (1906) Polystichum bolanicum Rosenst.; Fedde Repert. 12. 170 (1913) Polystichum bonapartii Rosenst.; Fedde Repert. 7. 297 (1909) Polystichum bonseyi W.H.Wagner & R.W.Hobdy; Contr. Univ. Michigan Herb. 19: 75 (1993) Polystichum bradei Rosenst.; Hedwigia 66. 365 (1915) Polystichum brassii Copel.; Philip. Journ. Sc. 78: 394. t.2 (1951) Polystichum braunii (Spenner) Fée; Mém. Foug. 5: 278 (1852) Trivial names: Weicher Schildfarn, Zarter Schildfarn, Brauns Schildfarn var. clarkei Christ; Bull. Boiss. 7: 25 (1899) Polystichum bulbiferum Barrington; Ann. Missouri Bot. Gard. 76(2): 371 (1989) Polystichum caducum (Ching) comb. ined. Polystichum calderonense Proctor; Mem. New York Bot. Gard. 53: 273 (1989) Polystichum californicum (D. C. Eaton) Diels; Engler & Prantl, Natürl. Pflanzenfamilien 1(4): 191 (1899) Trivial names: California Sword Fern Polystichum capillipes (Bak.) Diels; Nat. Pfl. 1(4): 191 (1899) Polystichum carvifolium Diels; Engl. Jahrb. 29: 194 (1900) Polystichum caudatum Fée; Gen. 278 (1850-52) Polystichum caudescens Dutra; An. Prim. Reun. Sul-Amer. Bot. 2: 49 (1940) Polystichum cheilanthoides Copel.; Univ. Calif. Publ . Bot. 18: 221 (1942) Polystichum chilense (Christ) Diels; Nat. Pfl. 1(4): 192 (1899) var. dusenii (C. Chr.) Looser ex R.Rodr. Gayana; Bot. 44(1-4): 48 (1987 publ. 1988) Polystichum chingae Ching; lcon. Fil. Sin. 4: t.182 (1937) Polystichum christianae (Jenm.) Underw. & Maxon; Maxon, Contr. U. S. Nat. Herb. 18: 30 t, 2 (1909) Polystichum christii Ching; Bull. Fan Mem. Hist. 2. 192 t. 7 (1931) Polystichum chunii Ching; Sinensia 1. 2 (1929), Ic. Fil. Sin. t. 13 Polystichum concinnum Lellinger ex Barrington; Ann. Missouri Bot. Gard. 76: 367 (1989) Polystichum conjunctum (Ching) comb. ined. Polystichum copelandii Christ; Phil. Journ. Sci. Bot. 2. 157 (1907) Polystichum coursii Tardieu; Mém. Inst. Sc. Madagascar ser. B. 7: 42. f.2(I-8) (1957) Polystichum craspedocarpium L.L.Xiang ex Ching & W.M.Chu; Acta Phytotax. Sin. 32(3): 259 (1994) Polystichum craspedosorum (Maxim.) Diels; Nat. Pfl. 1(4): 189 (1899) Polystichum crinigerum (C. Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 495 (1938) Polystichum crinulosum (Desv.) J.P.Roux; Novon 6(2): 202 (1996) Polystichum cuneatum Nakaike; Bull. Nat. Sci. Mus., Tokyo, B 4(3): 128 (1978) Polystichum cyclolobum C. Chr.; Lév., Cat. Pl. Yun-Nan 111 (1916) Polystichum cystostegia (Hook.) Armstr.; Tr. New Zeal. Inst. 13: 364 (1881) (cystostigium) Trivial names: Alpine shield fern Polystichum daguanense L.L.Xiang ex Ching; Acta Phytotax. Sin. 32(3): 263 (1994) Polystichum daymanense Nakaike; Bot. Exped. Papua New Guinea, Nation. Sci. Mus. (Tokyo), 102 (1975) Polystichum decoratum Maxon; Contr. U. S. Nat. Herb. 13. 20 t. 3 (1909) Polystichum decorum Ching & S. K. Wu; Fl. Xizangica 1: 229 (1983) Polystichum deflexum W.M.Chu ex Ching; Acta Bot. Yunnan., Suppl. 5: 47 (1992) Polystichum deltodon (Bak.) Diels; Nat. Pfl. 1(4): 191 (1899) Polystichum deminuens Maxon; Contr. U. S. Nat. Herb. 24. 53 t. 19 (1922) Polystichum dendrophilum Alderw.; Bull. Buit. III. 5. 221 (1922) Polystichum devonum Christ; Bot. Gaz. 61. 353 (1911) Polystichum diaphanum (Zoll. & Moritz) Moore; Ind. 90 (1858) Polystichum dielsii Christ; Bull. Géogr. Bot. Mans 238 (1906) Polystichum diffundens H.S.Kung & L.B.Zhang; Acta Bot. Yunnan. 17(1): 27 (1995) Polystichum discretum (D. Don.) Diels; Nat. Pfl. 1(4): 199 (1899) Polystichum distans E. Fourn.; Mexic. Pl. 1: 91 (1872) Polystichum dracomontanum Schelpe & N.C.Anthony; Contr. Bolus Herb. 10: 145 (1982) Polystichum drepanum (Sw.) Presl; Tent. 84 (1836) Polystichum dudleyi Maxon; Journ. Wash. Acad. Sci. 8. 620 (1918) Trivial names: Dudley's Holly Fern, Woodrustic fern. Polystichum echinatum (Gmel.) C. Chr.; Ind. 83. 905, 581 (1906) Polystichum edanoi Copel.; Philip. Journ. Sc. 83: 97. t.2 (1954) Polystichum ekmani Maxon; Proc. Biol. Soc. Wash. 46. 108 (1933) Polystichum elmeri Copel.; Philip. Journ. Sc. 81: 21 (1952) Polystichum eriorachis Alderw.; Bull. Buit. III. 5. 220 (1922) Polystichum erosum Ching & Shing; Acta Phytotax. Sin. 10: 303 (1965) [nom. nov. Polystichum erythrosorum A.R. Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 225 (1975) Polystichum exauriforme H.S.Kung & L.B.Zhang; Acta Bot. Yunnan. 17(1): 27 (1995) Polystichum excellens Ching; Icon. Fil. Sin. 5: t.234 (1958) Polystichum excelsior Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 16 (1984) Polystichum eximium (Mett. ex Kuhn) C. Chr.; Bull. Dept. Biol. Coll. Sci. Sunyatsen Univ. no. 6: 8 (1933) Polystichum falcilobum (Ching ex Y.T.Hsieh) comb. ined. Polystichum falcilobum Ching; Fl. Xizangica 1: 236 (1983) Polystichum falcinellum (Sw.) Presl; Tent. 83 (1836) Polystichum fallax Tindale; Proc. Linn. Soc. N.S.W. 80: 54. t.2 (1955) Polystichum fibrilloso-paleaceum (Kodama) Tagawa (1949) var. marginale Serizawa (1971) Polystichum fibrillosum Ching; Bull. Fan Mem. lnst. 2. 189 t. 4 (1931) Polystichum filiorum Mickel; R.J.Johns (ed.), Holtt. Mem. Vol. p. 127. Fig. 4 a (1997) Polystichum fimbriatum Christ; Bull. Géogr. Bot. Mans 237 (1906) Polystichum flaccidum Alderw.; Bull. Buit. III. 6. 220 (1922) Polystichum formosanum Rosenst.; Hedwigia 66. 338 (1915) Polystichum formosum Tindale; S.Austr. Nat. 28: 32 (1954) Polystichum fournieri A. R. Sm.; Amer. Fern J. 70(1): 27 (1980) [nom. nov. Polystichum frigidicola H.S.Kung & L.B.Zhang; Acta Bot. Yunnan. 17(1): 28 (1995) Polystichum fuentesii Espinosa; Rev. Chil. Nat. Hist. 36. 239 (1932) Polystichum furfuraceum A.R. Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 226 (1975) Polystichum fuscopaleaceum Alston; Bol. Soc. Broter. ser. 2. 30: 22 (1956) Polystichum fuscum Copel.; Philip. Journ. Sc. 81: 20 (1952) Polystichum gemmiparum C. Chr.; Gardens Bull. S. S. 7. 256 t. 54 (1934) Polystichum giganteum Fée; Cr. vasc. Br. 1: 127, t. 39, f. 2 (1869) Polystichum gongboense Ching & S. K. Wu; Fl. Xizangica 1: 228 (1983) Polystichum grandifrons C. Chr.; Ind. Suppl. III. 163 (1934) Polystichum grossidentatum Ching; Icon. Fil. Sin. 6: t.239 (1958) Polystichum guadalupense Fée; 11 mém. 74, t. 19, f.2 (1866) Polystichum habaense Ching & H.S.Kung; Bull. Bot. Res., Harbin 9(3): 4 (1989) Polystichum habbemense Nakaike; Bull. Nat. Sci. Mus., Tokyo, B 2(2): 51 (1976) Polystichum haleakalense Brackenr.; Expl. Exp. 16: 204, t. 28 (1854) Polystichum hancockii (Hance) Diels; Nat. Pfl. 1(4): 191 (1899) Polystichum harpophyllum (Zenker ex Kunze) Sledge; Bot. J. Linn. Soc. 84(1): 7 (1982) Polystichum harrisii Maxon; Contr. U. S. Nat. Herb. 13. 32 t. 4 B (1909) Polystichum hartwegii (Klotzsch) Hieron.; Hedwigia 46: 355 (1907) Polystichum hecatopterum Diels; Engl. Jahrb. 29: 193 (1900) Polystichum herbaceum Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 20 (1984) Polystichum hillebrandii Carr.; Seem. Fl. vit. 358 (1873) Polystichum holttumii C. Chr.; Gardens Bull. S. S. 7. 256 t. 53 (1934) Polystichum hottense C. Chr.; Kungl. Svenska Vetens.-akad. Handl. ser. 3. 16(2): 41. t.12(1-3) (1937) Polystichum huae H.S.Kung & L.B.Zhang; Acta Phytotax. Sin. 33(3): 311 (1995) Polystichum ichangense Christ; Bull. Soc. Fr. 52, Mem. 1: 28 (1905) Polystichum igaense Tagawa; Acta Phytotax. Geobot. 15: 15 (1953) Polystichum imbricans (D. C. Eat.) D. H. Wagner; Pteridologia 1: 50 (1979) Trivial names: Narrow-leaved Sword Fern ssp. imbricans (D. C. Eat.) D. H. Wagner; Pteridologia 1: 50 (1979) ssp. curtum (Ewan) D. H. Wagner; Pteridologia 1: 51 (1979) Polystichum inadae Kurata; J. Geobot. 11: 6 (1962) Polystichum inaense (Tagawa) Tagawa; Journ. Jap. Bot. 14: 709 (1938) Polystichum inaequale (Christ) comb. ined. Polystichum incisopinnulum H.S.Kung & L.B.Zhang; Acta Bot. Yunnan. 17(1): 25 (1995) Polystichum indochinense Tardieu & C. Chr.; Notul. Syst. [Paris] 7: 82 (1938) Polystichum integrilimbum Ching & H.S.Kung; Acta Bot. Boreal-Occid. Sin. 9(4): 275 (1989) Polystichum integripinnulum Ching; Fl. Xizangica 1: 210 (1983)
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For each species, information on its distribution by country and sometimes by province, region or state within a country; conservation status (where applicable), and all synonyms with publishing source (other latin names by which the species has been known) that we could find, and where applicable whether or not the given name has been verified; are available on the CDROM version of this page.

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