Family Lomariopsidaceae, genus Polystichum;
world species list.


By Michael Hassler and Brian Swale.      

A list of the species of the genus Polystichum, giving current name, authority, synonyms, published references, geographic distribution and (sometimes) illustrations.
Please note: an asterisk * in the distribution list means that the species is exotic to the country named.

  Polystichum species J to Z plus hybrids.
  Polystichum species A to I Here.

005.00 Class Filicopsida
		Trivial names: Ferns, varens, helechos, fougeres, farnkraut.
		Statistics: 15 orders, 53 families, 310 genera, 9300-10700 species 
		[11501 species and 454 hybrids listed.
047.000 Order Dryopteridales
		Statistics: 3 families, 30 genera, 1421-1752 spp. 
		[1989 species and 158 hybrids listed]
049.0000 Family Dryopteridaceae Ching; Acta Phytotax. Sin. 10: 1 (1965) s. str.
		Statistics: 21 genera, 672-939 spp. 
		[1162 species and 147 hybrids listed]
		Revision date: 20.07.2001
		= Aspidiaceae Frank; Leunis, Syn. Pflanzenfam. ed. 2, 3: 1469 [nom. illeg.
		= Dryopteridaceae tribus Stenolepieae H.S. Kung; Proc. Int. Symp. Syst. Pteridol. 
		(eds. K.H. Shing & K.U. Kramer): 172 (1989)
049.0200 Tribus Dryopterideae

049.0217 Polystichum Roth; Tent. Fl. Germ. 3: 31, 69 (1799) [nom. cons. Statistics: 200-300 spp. nearly worldwide, 55 in trop. America [348 species and 57 hybrids listed]. This used to be a collective "omnibus genus", and reflecting that fact and that taxonomists still have to do much more work on these species, many currently listed species may better belong in other genera. China has too many (probably invalid) microspecies, and many of the Malesian species may turn out to be synonyms, too. Revision date: 20.07.2001 Trivial names: Schildfarn = Acropelta Nakai; Bull. Nat. Sci. Mus. Tokyo 33: 5 (1953) = Aetopteron Ehrhart; Beitr. 148 (1789), ex House, Amer. Fern Journ. 10. 88 (1920) = Cyrtogonellum Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 327 (1938) = Cyrtomidictyum Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 182 (1940) = Hemesteum Léveillé; Flore du Kouy-tcheou 496 (1915) [non Hemestheum Newman (1851) = Hemigonum J. Smith; Hist. Fil, 204 (1875) = Hypopeltis Michx.; Fl. Bor. Am. 2: 226 (1803), in obs., ex Bory, Bél., Voy. Bot. 2: 63 (1833) = Papuapteris Christensen; Brittonia 2: 300 (1937) = Plecosorus Fée; Mem. Fam. Foug. 5 (Gen. Fil.) 150 (1852) = Porpaea Presl ms. ex Ettingsh.; Farnkr. 174 (1865)
Polystichum jinfoshanense Ching & Z.Y.Liu; Bull. Bot. Res, Harbin 3(4): 29 (1983) Polystichum kadyrovii A. Askerov & A. Bobrov; Bot. Zhurn. 57: 1297 (1972) Polystichum kalambatitrense Tardieu; Notul. Syst. [Paris] 15. 165. f.1(1-4) (1956) Polystichum keysserianum Rosenst.; Fedde Repert. 10. 331 (1912) Polystichum kilimanjaricum Pichi-Serm.; Webbia 27: 445 (1972) publ. (1973) Polystichum kinabaluense C. Chr.; Gardens Bull. S. S. 7. 255 t. 52 (1934) Polystichum kiusiuense Tagawa; Acta Phytotax. Geobot. 16: 15 (1953) Polystichum kruckebergii W. H. Wagner; Amer. Fern J. 56: 4 (1966) Trivial names: Kruckeberg's Holly Fern Polystichum kunthianum B.K.Nayar & Geev.; Fern Fl. Malabar: 214 (1993) Polystichum kurokawae Tagawa; Journ. Jap. Bot. 26: 20 (1951) Polystichum kwakiutlii D.H. Wagner; Amer. Fern J. 80: 50, fig. 1 (1990) Polystichum kwangtungense Ching; Icon. Fil. Sin. 5: t.236 (1958) Polystichum lachenense (Hook.) Bedd.; Ferns br. Ind. t. 32 (1865) Trivial names: Aleutian Holly Fern Polystichum lanceolatum (Bak.) Diels; Engl. Jahrb. 29: 193 (1900) Polystichum lancilobum C. Chr.; Kungl. Svonska Vetens.-akad. Handl. ser. 3. 16(2): 40. t.9(1-4) (1937) Polystichum lanosum Fée; Cr. vasc. Br. 1: 126. t. 40, f. 2 (1869) Polystichum latilepis Ching & H.S.Kung; Acta Bot. Boreal-Occid. Sin. 9(4): 273-277 (1989) Polystichum lemmonii Underw.; Native Ferns ed. 6: 116 (1900) Trivial names: Shasta Fern Polystichum lentum (D. Don) Moore; Ind. Fil. 86 (1858) Polystichum lepidocaulon (Hook.) J. Sm.; Ferns br. and for. 286 (1866) Polystichum leveillei C. Chr.; Bull. Acad. int. Géogr. Bot. (1918) 143 Polystichum lindsaeifolium Scort.; Ridley, Journ. Mal. Br. R. As. Soc. 4. 61 (1926) Polystichum lineare (C. Chr.) Copel.; Philip. Journ. Sc. 78: 397 (1951) Polystichum liuii Ching; Bull. Bot. Res, Harbin 3(4): 28 (1983) Polystichum lonchitis (L.) Roth; Tent. Fl. Germ. 3: 71 (1799) Trivial names: Lanzen-Schildfarn, Northern Holly Fern Polystichum longecuspis Fée; Cr. vasc. Br. 1: 129. t. 41, f. 2 (1869) Polystichum longiaristatum Ching; Boufford & K. H. Shing, J. Arnold Arbor. 64(1): 33 (1983) Polystichum longidens Ching & S. K. Wu; Fl. Xizangica 1: 217 (1983) Polystichum longifrons Kurata; Sci. Rep. Yokosuka City Mus. 10: 31 (1964) Polystichum longipaleatum Christ; Not. syst. 1. 35 (1909) Polystichum longipes Ching & S. K. Wu; Fl. Xizangica 1: 213 (1983) Polystichum longissimum Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 16 (1984) Polystichum luctuosum (Kunze) T. Moore; Ind. 95 (1858) Polystichum machaerophyllum Slosson; Bull. Torr. Cl. 40. 688 t. 26 f. 4-5 (1913) Polystichum macleaii (Bak.) Diels; Nat. Pfl. 1(4): 190 (1899) (macleanii) Polystichum macrochlaenum Ching & S. K. Wu; Fl. Xizangica 1: 227 (1983) Polystichum maevaranense Tardieu; Notul. Syst. [Paris] 15: 166. f.l(5-9) (1956) Polystichum magnificum Ballard; Kew Bull. 195 7: 48b f. 1 (1957) Polystichum makinoi (Tagawa) Tagawa; Acta Phytotax. Geobot. 6: 258 (1936) Japan Polystichum manmeiense (Christ) Nakaike in Y.Otani; Rep. Cryptogamic Study Nepal (Misc. Publ. Nat. Sci. Mus. Tokyo): 141 (1982) Polystichum marionense Alston & Schelpe; Journ. S.Afr. Bot. 23: 106. i.l(a). t.34 (1957) Polystichum marquesense E. Brown; Bishop Mus. Bull. 89. 40 t. 6 (1931) Polystichum martini Christ; Bull. Ac. Géogr. Bot. Mans 263 (1902) Polystichum mashikoi Kurata; Journ. Geobot. 7: 116 (1958) Polystichum medogense Ching & S. K. Wu; Fl. Xizangica 1: 214 (1983) Polystichum mehrae Fraser-Jenk. & Khullar; Indian Fern J. 3(1-2): 121 (1986) Polystichum meiguense Ching & H.S.Kung; Acta Bot. Boreal-Occid. Sin. 9(4): 269 (1989) Polystichum melanostipes Ching & H.S.Kung; Bull. Bot. Res., Harbin 9(3)- 10 (1989) Polystichum michelii Christ; Bull. Géogr. Bot. Mans 16 (1910) Polystichum mickelii A.R. Sm.; Amer. Fern J. 70: 25 (1980) Polystichum microchlamys (Christ.) Matsumura; Index Pl. Jap. 1: 343 (1904) Trivial names: Attu Holly Fern Polystichum microsorium Fée; Cr. vasc. Br. 1: 125, t. 40, f. 1 (1869) Polystichum minimum (Y.T.Hsieh) comb. ined. Polystichum molliculum Christ; Bot. Gaz. 61. 354 (1911) Polystichum mollissimum Ching; Fl. Xizangica 1: 232 (1983) Polystichum moluccense Moore; Ind. 84 (1858) Polystichum monotis (Christ) C. Chr.; Ind. 83 (1905), 584 (1906) Polystichum montevidense Rosenst.; Hedwigia 46. 111 (1906) var. squamulosa (Hieron.) Hieron. var. nudicaule (Rosenst.) R. M. Tryon; Fieldiana, Bot. 27: 52 (1991) Polystichum monticola N.C.Anthony & Schelpe; Bothalia 15(3-4): 554 (1985) Polystichum moorei Christ; Ark. för Bot. 4(12): 3 (1905) ex W. R. B. Oliver Polystichum morii Hayata; Ic. Pl. Taiwan 7. 95 f (1918) Polystichum mucronatum (Sw.) Presl; Tent. 83 (1936) Polystichum mucronifolium (BI.) Nayar et Kaur; Comp. to Bedd. Handb. 49 (1974) Polystichum muenchii (H. Christ) C. Chr.; Index Filic. 585 (1906) Polystichum multifidum (Mett.) Moore; Ind. LXXXIV (1857) var. pearcei (Phil.) R.Rodr.; Gayana, Bot. 44(1-4): 51 (1987 publ. 1988) Polystichum munitum (Kaulf.) Presl; Tent. 83 (1836) Trivial names: Common Sword Fern Polystichum muricatum (L.) Fée; Mém. Foug. 5: 278 (1852) Polystichum muticum Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942) Polystichum myer-dreesii Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942) Polystichum nanchuanicum Ching; Bull. Fan Mem. Inst. Biol. ser. 2. 1: 310 (1949) Polystichum neolobatum Nakai; Bot. Mag. Tokyo 39: 118 (1925) Polystichum nepalense (Spreng.) C. Chr.; Ind. 84 (1905), 585 (1906) var. subbipinnatum C. Chr.; Contr. U.S. Nation Herb. 26: 284 (1931) Polystichum nigrum Ching & H.S.Kung; Bull. Bot. Res., Harbin 9(3): 8 (1989) Polystichum ningshenense Ching & Hsu; Fl. Tsinlingensis 2: 223 (1974) Polystichum nudum Copel.; Phil. Journ. Sci. 1. Suppl. II. 145 (1906) Polystichum obai Tagawa; Acta Phytotax. 2. 194 (1933) Polystichum obliquum (D. Don) Moore; Ind. Fil. 87 (1858) Polystichum obtusipinnum Ching & H.S.Kung; Bull. Bot. Res., Harbin 9(3): 6 (1989) Polystichum obtusum J. Sm.; Journ. Bot. 3: 412 (1841), nomen, ex Presl, Epim. 53 (1849) Polystichum ohmurae Kurata; Hokuriku Journ. Bot. 5: 79 (1956) var. fujipedis Nameg. & Sa. Kurata; Sci. Rep. Yokosuka City Mus. 15: 46 (1969) Polystichum oligocarpum Ching ex H.S. Kung & L.B. Zhang in L.B. Zhang & H.S. Kung (1995) Polystichum omeiense (Ching ex Y.T.Hsieh) comb. ined. Polystichum opacum Rosenst.; Hedwigia 46. 112 (1906) Polystichum oppositum Copel.; Philip. Journ. Sc. 56: 474. t.5 (1935) Polystichum orbiculatum (Desv.) J. Rémy & Fée in Gay; Fl. Chil. 6: 515 (1853) var. boboense (Hieron.) R. M. Tryon; Fieldiana, Bot. 27: 54 (1991) Polystichum ordinatum (Kunze) Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd. ser. 5, 1: 276 (1849) Polystichum oreodoxa H.S.Kung & L.B.Zhang ex Ching; Acta Phytotax. Sin. 33(3): 309 (1995) Polystichum otomasui Kurata; J. Geobot. 12: 67 (1963) Polystichum otophorum (Franch.) Bedd.; Handb. Suppl. 42 (1892) Polystichum ovatopaleaceum (Kodama) Kurata; Sci. Rep. Yokosuka City Mus. 10: 35 (1964) Polystichum paleatum Copel.; Occ. Pap. Bishop Mus. 14: 58. t.9 (1938) Polystichum papuanum C. Chr.; Brittonia 2: 299 (1937) Polystichum papyrifolium Alderw.; Bull. Buit. II no. 28. 41 (1918) Polystichum paradeltodon L.L.Xiang; Acta Phytotax. Sin. 32(3): 265 (1994) Polystichum parasinense C.Y.Yang; Fl. Xinjiangensis 1: 302 (1992 publ. 1993) Polystichum parvifoliolatum W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 49 (1992) Polystichum parvipinnulum Tagawa; Acta Phytotax. Geobot. 6: 160 (1937) Polystichum pflanzii Hieron.; Engl. Jahrb. 49. 179 (1912) Polystichum pianmaense W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 51 (1992) Polystichum piceopaleaceum Tagawa; Acta Phytotax. Geobot. 5: 255 (1936) Polystichum pilosum Copel.; Journ. Arnold Arbor. 30: 439 (1949) Polystichum plaschnickianum (Kze.) Moore; Ind. 100 (1858) Polystichum platychlamys Ching; Fl. Tsinlingensis 2: 224 (1974) Polystichum platylepis Fée; Cr. vasc. Br. 1: 129. t. 41, f. 1 (1869) Polystichum platyphyllum (Willd.) C. Presl; Tent. Pterid. 84 (1836) var. klotschii Rosenst. Polystichum plicatum (Poepp. ex Kunze) Hicken ex Hosseus; Trab. Inst. Bot. Parmac. [Buenos Aires] 33: 9 (1915) Polystichum polyblepharum (Roem. ex Kunze) C. Presl; Epim. 56 (1849-51) f. scabiosum (Sa.Kurata) Seriz.; J. Phytogeogr. Taxon. 29(1): 24 (1981) Polystichum polystichiforme (Fée) Maxon; Contr. U. S. Nat. Herb. 18. 35 (1909) Polystichum prescottianum (Wall. ex Mett.) Moore; lnd. Fil. 101 (1858) var. castaneum Clarke; Trans Linn. Soc. Lond. Il. Bot, 1: 509. 1880 Polystichum prionolepis Hayata; Ic. Fl. Formosa 4. 197 f. 134 (1914) Polystichum proliferum (R. Br.) C. Presl; Tent. 83 (1836) Trivial names: Mother shield fern Polystichum prolificans Alderw.; Bull. Buit. III. 2. 170 (1920) Polystichum pseudodeltodon Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 17 (1984) [non Tagawa (1937) Polystichum pseudo-makinoi Tagawa; Acta Phytotax. Geobot. 5: 257 (1936) Polystichum pseudomicrophyllum Nakaike; Bull. Nat. Sci. Mus., Tokyo, B 9(3): 103 (1983) Polystichum pseudosetosum Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 19 (1984) Polystichum pseudotsus-simense Ching; Bull. Fan Mem. Inst. 2. 190 (1931) Polystichum pulogense Copel.; Philip. Journ. Sc. 81: 19 (1952) Polystichum pultenii Nakaike; Bull. Nat. Sci. Mus., Tokyo, B 43(3): 129 (1978) Polystichum punctiferum C. Chr.; Contr. U.S. Nation Herb. 26: 288. t. 17 (1931) Polystichum pungens (Kaulf.) Presl; Tent. 83 (1836) Polystichum pycnolepis (Kunze ex Kl.) Moore; Ind. 92 (1858) Polystichum pygmaeum Copel.; Univ. Calif. Publ. Bot. 19: 300. t.56 (1941) Polystichum qamdoense Ching & S. K. Wu; Fl. Xizangica 1: 227 (1983) Polystichum rachichlaena Fée; Mém. Foug. 8: 100 (1857) Polystichum rapense E. Brown; Bishop Mus. Bull. 89. 39 t. 4 A (1931) Polystichum remotum Fée; Cr. vasc. Br. 1: 125, t. 39. f. 1 (1869) Polystichum retroso-paleaceum (Kodama) Tagawa; Journ. Jap. Bot. 13: 187 (1937) Polystichum rhizophorum (Jenm.) Maxon; Contr. U. S. Nat. Herb. 18. 36 t. 7 (1909) Polystichum rhizophyllum (Sw.) Presl; Tent. 82 (1836) Polystichum richardii (Hook.) J. Smith (1875) Trivial names: Common shield fern, Pikopiko, Tutoke (Maori) Polystichum rigens Tagawa; Acta Phytotax. Geobot. 6: 91 (1937) Polystichum rochaleanum Glaz. ex Fée; Cr. vasc. Br. 2: 69, t. 99, f. 2 (1872) Polystichum rupicola W.M.Chu ex Ching; Acta Bot. Yunnan., Suppl. 5: 51 (1992) Polystichum salicifolium (Ching & Y.T.Hsieh) comb. ined. Polystichum salwinense Ching & H.S.Kung; Bull. Bot. Res., Harbin 9(3): 7 (1989) Polystichum scariosum (Roxb.) C. Morton; Contrib. U.S. Nation. Herb. 38: 359 (1974) Polystichum scopulinum (D. C. Eaton) Maxon; Fern Bull. 8: 29 (1900) Trivial names: Crag Holly Fern, Rock Sword Fern Polystichum semifertile (Clarke) Ching; Lingnan Sc. Journ. 15: 398 (1938) Polystichum setiferum (Forsskal) Moore ex Woynar; Mitt. Naturwiss. Vereines Steiermark 49: 181 (1913) Trivial names: Borstiger Schildfarn Polystichum setigerum (Presl) Presl; Tent. Pterid. 83 (1836) Trivial names: Alaska Sword Fern Polystichum setulosum Ching; Bull. Fan Mem. Inst. 2. 188 t. 3 (1931) Polystichum shandongense J.X.LI & Y. Wei; Acta Phytotax. Sin. 22(2): 164 (1984) Polystichum shensiense Christ; Bull. Géogr. Bot. Mans 113 (1906) Polystichum shimurae Sa.Kurata ex Seriz.; J. Jap. Bot. 54(5): 139 (1979) Polystichum shizuokaense Nakaike; New Fl. Japan, Pterid.: 842 (1999) [nom. nov. Polystichum silaense Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 71 (1941) Polystichum silvaticum (Col.) Diels; Nat. Pfl. 1(4): 192 (1899) Polystichum simile (Ching ex Y.T.Hsieh) comb. ined. Polystichum sinkiangense Ching ex C.Y. Yang; Bull. Bot. Res., Harbin 8(2): 55 (1988) Polystichum sinotsussimense Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 18 (1984) Polystichum smithii Mickel & Beitel; Pterid. Fl. Oaxaca (Mem. New York Bot. Gard. 46): 315 (1988) Polystichum solitarium (Maxon) Underw.; Maxon, Contr. U. S. Nat. Herb. 10. 493 (1908) Polystichum sozanense H.S.Kung & L.B.Zhang ex Ching; Acta Phytotax. Sin. 33(3): 309 (1995) Polystichum speciosissimum (A. Braun ex Kunze) Copel.; Gen. Fil. (Ann. Cryptog. Phytopathol. 5) 109 (1947) Polystichum spongiosum Maxon; Amer. Fern Journ. 19. 47 (1929) Polystichum squarrosum (D. Don) Fée; Gen. Fil. 278 (1850-52) Polystichum stenophyllum Christ; Bull. Soc. Bot. France 52., Mém. 1: 27 (1905) Polystichum stimulans (Kunze ex Mett.) Bedd.; Ferns Brit. India t. 31 (1865) Polystichum stokesii E. Brown; Bishop Mus. Bull. 89. 40 t. 5 (1931) Polystichum stuebelii Hieron.; Hedwigia 46. 355 t. 8 f. 21 (1907) Polystichum subacutidens L.L.Xiang ex Ching; Acta Phytotax. Sin. 32(3): 261 (1994) Polystichum subattenuatum Ching & W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 51 (1992) Polystichum subdeltodon Ching; Bull. Fan Mem. Inst. Biol. ser. 2. 1: 308 (1949) Polystichum subintegerrimum (Hook. & Arn.) R.Rodr.; Gayana, Bot. 44(1-4): 48 (1987 publ. 1988) Polystichum submarginale (Baker) L.L.Xiang ex Ching; Acta Phytotax. Sin. 32(3): 263 (1994) Polystichum submite (Christ) Diels; Engl. Jahrb. 29: 192 (1900) Polystichum subtripteron Tzvelev; Novosti Sist. Vyssh. Rast. 26: 7 (1989) Polystichum subulatum L.B. Zhang ex Ching; Acta Bot. Yunnan. 16(2): 133 (1994) Polystichum tagawanum Kurata; Journ. Geobot. 7: 41 (1958) Polystichum taizhongense H.S. Kung (1997) Polystichum takakii Nakaike; Bot. Exped. Papua New Guinea, Nation. Sci. Mus. (Tokyo), 11 1 (1975) Polystichum talamancanum Barrington; Ann. Missouri Bot. Gard. 76: 369 (1989) Polystichum tangmaiense H.S.Kung & Y.Tateishi; Acta Phytotax. Sin. 32(3): 268 (1994) Polystichum tenuium (Ching) comb. ined. Polystichum tetragonum Fée; 8 mém. 99 (1857) Polystichum thomsonii (Hook. f.) Bedd.; Ferns Brit. India 1: t. 126 (1865) Polystichum tialooshanense Ching; Ching & Wang, Acta Phytotax. Sin. 9: 366 (1964) Polystichum tibeticum Ching; Fl. Xizangica 1: 219 (1983) Polystichum tijucense Fée; Cr. vasc. Br. 1: 127. t. 38, f. 1 (1869) Polystichum tortuosum Nakaike; Bull. Nat. Sci. Mus., Tokyo, B 2(3): 102 (1976) Polystichum transkeiense W.Jacobsen; J. S. African Bot. 44(2): 169 (1978) Polystichum transvaalense N.C.Anthony; Contr. Bolus Herb. 10: 146 (1982) Polystichum trapezoides Presl; Tent. 83 (1836) Polystichum tridens (Hook.) Fée; 11 mém. 74 (1866) Polystichum tripteron (Kunze) C. Presl; Epim. 55 (1851) Polystichum tsaratananense Tardieu; Mém. Inst. Sc. Madagascar ser. B. 7: 44 (1956) Polystichum tsingkanshanense Ching ex K.H. Shing & J.R Cheng; Jiangxi Sci. 8(3): 46 (1990) Polystichum underwoodii Maxon; Contr. U. S. Nat. Herb. 13. 38 t. 9 (1909) Polystichum variiforme (Hayata) Tagawa; Journ. Jap. Bot. 13: 186 (1937) Polystichum vestitum (Forst.) Presl; Tent. 83 (1836) Trivial names: Prickly shield fern, Punui (Maori) Polystichum volkensii (Hier.) C. Chr.; Ind. 97 (1905), 586 (1906) Polystichum vulcanicum Moore; Ind. 108 (1858) Polystichum wattii (Bedd.) C. Chr.; Ind. 98 (1905), 589 (1906) Polystichum whiteleggii Watts; Proc. Linn. Soc. N. S. Wales 89. p. 257-62 (1914) Polystichum wilesianum Proctor; Brit. Fern Gaz. 10: 24, t. 3 (1968) Polystichum wilsonii Christ; Bot. Gaz. 61. 353 (1911) Polystichum wolfii Hieron.; Hedwigia 46. 366 t. 8 t 23, 24 (1907) Polystichum woodsioides Christ; Bot. Gaz. 61. 354 (1911) Polystichum woronowii Fomin; Moniteur Jard. bot. Tiflis livr. 18. 21. c. tab (1911) Polystichum wrightii (Bak.) C. Chr. apud Maxon; Contr. U. S. Nat. Herb. 16. 50 (1912) Polystichum wulingshanense S.F.Wu in Keys Vasc. Pl. Wuling Mount. (ed. W.T.Wang et al.): 568 (1995) Polystichum x amboversum Kurata; Sci. Rep. Yokosuka City Mus. 10: 36 (1964) Remarks: Polystichum ovatopaleaceum x retrorsopaleaceum Polystichum x anceps Kurata; Sci. Rep. Yokosuka City Mus. 10: 32 (1964) Remarks: Polystichum longifrons x polyblepharum Polystichum x bicknellii (Christ) Hahne; Allg. Bot. Zeitschr. 10. 103 (1904) Remarks: Polystichum aculeatum (L.) Roth x P. setiferum (Forssk.) Moore ex Woynar Polystichum x bootii Gandog.; Fl. Eur. 27: 168 (1891) Remarks: Dryopteris cristata x spinulosa Polystichum x carthusianum Sanio; Verh. Bot. Ver. Brand. 25: 84 (1884) Remarks: Dryopteris filix-mas x spinulosa Polystichum x dmitrievae A. Askerov Polystichum x eberlei D.E.Meyer; Nova Hedwigia 1: 107 (1959) Remarks: Polystichum braunii x lonchitis Polystichum x flemingii Fraser-Jenkins; Aspects Plant Sciences 13: 277 (1991) Remarks: Polystichum discretum x yunnanense Polystichum x fominii A. Askerov & A. Bobrov; Bot. Zhurn. 57: 1298 (1972) Remarks: Polystichum aculeatum x woronowii Polystichum x fujisanense Seriz.; J. Jap. Bot. 45: 263 (1970) Remarks: Polystichum braunii x ohmurae Polystichum x gemmilachenense Miyam. & T.Nakam.; J. Jap. Bot. 58(5): 149 (1983) Remarks: Polystichum lachenense x stenophyllum Polystichum x hagenahii Cody; Amer. Fern J. 58: 30 (1968) Remarks: Polystichum acrostichoides x Polystichum lonchitis Polystichum x hakonense Kurata; Journ. Geobot. 9: 99 (1961) Remarks: Polystichum polyblepharum var. intermedium x pseudo-makinoii Polystichum x hitoyoshiense Kurata; Sci. Rep. Yokosuka City Mus. 15: 47 (1969) Remarks: Polystichum otomasui x pseudomakinoi Polystichum x hokurikuense Kurata; Sci. Rep. Yokosuka City Mus. 10: 39 (1964) Remarks: Polystichum longifrons x retrosopaleaceum Polystichum x iidanum Kurata; Sci. Rep. Yokosuka City Mus. 10: 33 (1964) Remarks: Polystichum longifrons x makinoi Polystichum x illyricum (Borb.) Hahne; Allg. Bot. Zeitschr. 10. 103 (1904) Remarks: Polystichum aculeatum (L.) Roth x Polystichum lonchitis (L.) Roth Polystichum x inayatii Fraser-Jenkins; Aspects Plant Sciences 13: 277 (1991) Remarks: Polystichum piceopaleaceum x yunnanense Polystichum x izuense Kurata; Journ. Geobot. 7: 42 (1958) Remarks: Polystichum makinoi x tagawanum Polystichum x jamunae Fraser-Jenkins; Aspects Plant Sciences 13: 277 (1991) Remarks: Polystichum mehrae x squamosum Polystichum x jitaroi Kurata; Sci. Rep. Yokosuka City Mus. 10: 33 (1964) Remarks: Polystichum polyblepharum x pseudomakinoi Polystichum x kai-montanum Seriz.; J. Jap. Bot. 54(5): 143 (1979) Remarks: Polystichum ohmurae x ovatopaleaceum var. coraiense Polystichum x kasayamense Kurata; Sci. Rep. Yokosuka City Mus. 15: 47 (1969) Remarks: Polystichum braunii x polyblepharum Polystichum x kiyozumianum Kurata; Journ. Geobot. 9: 99 (1961) Remarks: Polystichum pseudo-makinoi x tagawanum Polystichum x kumamontanum Kurata; J. Geobot. 12: 68 (1963) Remarks: Polystichum otomasui x polyblepharum Polystichum x kunioi Kurata; Sci. Rep. Yokosuka City Mus. 15: 48 (1969) Remarks: Polystichum braunii x makinoi Polystichum x kuratae Seriz.; J. Jap. Bot. 54(5): 143 (1979) Remarks: Polystichum polyblepharum x pseudomakinoi Polystichum x lonchitiforme (Halácsy) Becherer; Boll. Soc. Ticin. Sc. Nat. 36: 17 (1941) Remarks: Polystichum lonchitis x setiferum Polystichum x luerssenii (Doerfl.) Hahne; Allg. Bot. Zeitschr. 10. 103 (1904) Remarks: Polystichum aculeatum (L.) Roth x Polystichum braunii (Spenn.) Fée Polystichum x maderense Johnson (1866) Remarks: Polystichum falcinellum x Polystichum setiferum Polystichum x meyeri Sleep & Reichst.; Bauhinia 3: 303, f. 7, 13, 15, 17 (1967) Remarks: Polystichum braunii (Spenn.) Fée x Polystichum lonchitis (L.) Roth Polystichum x microlepis Kurata; Sci. Rep. Yokosuka City Mus, 10: 37 (1964) Remarks: Polystichum makinoi x retrosopaleaceum Polystichum x midoriense T.Oka & Ohtani; Sci. Rep. Yokosuka City Mus. 27: 57 (1980) Remarks: Polystichum fibrillosopaleaceum x ovatopaleaceum Polystichum x minamitanii Sa.Kurata ex Seriz.; J. Jap. Bot. 54(5): 141 (1979) Remarks: Polystichum makinoi x otomasui Polystichum x miuranum Kurata; Sci. Rep. Yokosuka City Mus. 15: 46 (1969) Remarks: Polystichum fimbrillosopaleaceum x polyblepharum Polystichum x namegatae Kurata; Hokuriku Journ. Bot. 6: 113 (1957) Remarks: Polystichum makinoi x pseudo-makinoi Polystichum x ohtanii Kurata; Sci. Rep. Yokosuka City Mus. 15: 46 (1969) Remarks: Polystichum fimbrillosopaleaceum x longifrons Polystichum x okanum Kurata; Journ. Geobot. 8: 16 (1959) Remarks: Polystichum retrosopaleaceum x tagawanum Polystichum x ongataense Kurata; Hokuriku Journ. Bot. 6: 114 (1957) Remarks: Polystichum retroso-paleaceum var. ovato-paleaceum x pseudo-makinoi Polystichum x potteri Barrington; Rhodora 88(855): 298 (1986) Remarks: Polystichum acrostichoides x Polystichum braunii Polystichum x pseudoparvipinnulum Miyam. & T.Nakam.; J. Jap. Bot. 58(5): 146 (1983) Remarks: Polystichum parvipinnulum x piceopaleaceum Polystichum x remotum (A.Br.) P.Fourn.; Quatre Fl. France 17 (1934) [non Fée Remarks: Polystichum filix-mas x spinulosum Polystichum x safarovii A. Askerov & A. Bobrov; Bot. Zhurn. 57: 1298 (1972) Remarks: Polystichum braunii (Spenn.) Fée x Polystichum woronowii Fomin (or P. setiferum) Polystichum x sarukurense Seriz.; J. Jap. Bot. 54(5): 144 (1979) Remarks: Polystichum braunii x microchlamys var. azumiense Polystichum x shin-tashiroi Kurata; J. Geobot. 1 1: 38 (1962) Remarks: Polystichum microchlamys x retrosopaleaceum Polystichum x silviamontanum Miyarn. & T.Nakain.; J. Jap. Bot. 58(5): 147 (1983) Remarks: Polystichum piceopaleaceum x wilsonii Polystichum x stewartii Fraser-Jenkins; Aspects Plant Sciences 13: 276 (1991) Remarks: Polystichum lonchitis x yunnanense Polystichum x suginoi Kurata; J. Geobot. 12: 68 (1963) Remarks: Polystichum x tagawanum Polystichum x suyamanum Sa.Kurata ex Seriz.; J. Jap. Bot. 54(5): 142 (1979) Remarks: Polystichum igaense x longifrons Polystichum x takaosanense Kurata; Journ. Geobot. 8: 17 (1959) Remarks: Polystichum polyblepharum var. intermedium x retroso-paleaceum var. ovato-paleaceum Polystichum x tare-bhirense Nakaike & Gurung; Bull. Nat. Sci. Mus., Tokyo, B 14(3): 83 (1988) Remarks: Polystichum piceopaleaceum x squarrosum. Status doubtful, possible syn. of P. yunnanense Polystichum x tetsuyamense Sa.Kurata ex Seriz.; J. Jap. Bot. 54(5): 141 (1979) Remarks: Polystichum otomasui x ovato-paleaceum Polystichum x titibuense Kurata; J. Geobot. 12: 68 (1963) Remarks: Polystichum x retrosopalaeceum var. coraiense Polystichum x tokyoense Sa.Kurata ex Seriz.; J. Jap. Bot. 54(5): 140 (1979) Remarks: Polystichum ovatopaleaceum x shimurae Polystichum x utsumii (Kurata) Kurata; Sci. Rep. Yokosuka City Mus. 10: 37 (1964) Remarks: Polystichum pseudomakinoi x retrosopaleaceum Polystichum x wirtgenii Hahne; Allg. Bot. Zeitschr. 10. 103 (1904) Remarks: Polystichum braunii (Spenn.) Fée x Polystichum setiferum (Forssk.) Moore ex Woynar Polystichum x yokohamense T.Oka & Ohtani; Sci. Rep. Yokosuka City Mus. 27: 57 (1980) Remarks: Polystichum fibrillosopaleaceum x tagawanum Polystichum xichouense (S. K. Wu & Mitsuta) comb. ined. Polystichum xiphophyllum (Bak.) Diels; Nat. Pfl. 1(4): 189 (1906) Polystichum yadongense Ching & S. K. Wu; Fl. Xizangica 1: 220 (1983) Polystichum yaeyamense (Makino) Makino; Bot. Mag. Tokyo 23. 144 (1909) Polystichum yuanum Ching; Icon. Fil. Sin. 5: t.235 (1958) Polystichum yungense Rosenst.; Fedde Repert. 11. 55 (1912) Polystichum yunnanense Christ; Not. syst. 1. 34 (1909) Polystichum zambesiacum Schelpe; Bol. Soc. Brot. Ser. 2, 41: 215 (1967)
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