Checklist of World Ferns.

Brian Swale B.Sc.(NZ), M.A.(Forestry)(Oxon)

References: and other notes.

To go directly to the other notes about fern distribution, and taxonomy, Click Here.

  1. Brownsey, Patrick J, and John C. Smith-Dodsworth. 1989. New Zealand Ferns and Allied Plants. David Bateman. pp. 168. ISBN 1 86953 003 9.
  2. Holttum, R.E. 1966. Flora of Malaya, Volume II, Ferns of Malaya. 2nd edition. Government Printing Office, Singapore, pp. 653.
  3. Grounds, Roger. 1974. Ferns. Pelham Books. pp. 264. ISBN 0 7207 0687 4.
  4. Hoshizaki, Barbara Joe. 1975. Fern Growers Manual. Alfred A. Knopf, New York. pp. 256 + xiii, xi. ISBN 0 394 73774 1 (paperback edition, 1979).
  5. Jones, David L. 1987. Encyclopedia of Ferns. Timber Press, Portland Oregon. pp. 433. ISBN 0 88192 054 1.
  6. Jones, David L. and S.C. Clemescha. 1976. Australian Ferns and Fern Allies. A.H.and A.W. Reed. pp. 294. ISBN 0 589 07197 1.
  7. Perl, Philip. 1977. Ferns. Time-Life Books Inc. pp. 159. LCC 7 83154.
  8. The Royal Botanic Gardens, Kew. (M. Jackson, 9/3/1996), at   direct to the databases here
  9. Johnson, A. 1977. The Ferns of Singapore Island; a Student's Guide. Singapore University Press. pp. xii + 126. No ISBN.
  10. Missouri Botanic Gardens Tropical Database; W3 Tropicos at   here
  11. N.Z. Journal of Botany. Royal Society of New Zealand.   here
  12. Fern Gazette 1996. 15 (3):91-100
  13. Prodromus Florae Novae Hollandiae et Insulae Van-Diemen, London. 1810.
  14. Davidse, G; M. Sousa S; & Knapp, S. (Eds.).  1995.   Lista Anotada, from Flora Mesoamericana 1: i - xxi, 1 - 470. ( here
  15. Chinnery, Louis E. & George Rogers. Checklist of Barbadian ferns, based on sheets in the Barbados National Herbarium - BAR. See the latest list at Barbados. or contact Louis at just here.
  16. Gordon McPherson, email and Chris Freeland, email.   Missouri Botanic Gardens Checklist of Lope-Okanda Reserve, Gabon, Africa. See this (being constantly updated) at here.
  17. Andean Botanical Information System at with many pages recording their ongoing work click here
  18. Information from the following texts was available, and used as appropriate for the species and infraspecific taxa.
  19. Taylor, T.M.C, 1970. Pacific Northwest Ferns and Their Allies. University of Toronto Press. pp. i + 247. ISBN 0- 8020-5227-4.
  20. GULF OF GUINEA CONSERVATION GROUP - GULF OF GUINEA ISLANDS' BIODIVERSITY NETWORK, website references in "Bibliografia relevante para o estudo dos pteridófitos de São Tomé e Príncipe" at citations as at 28 August 2000, with links to all their fern pages at
    1. ALSTON, A. H. G., 1944. Pteridophyta. In A. Exell, Catalogue of the Vascular Plants of S.Tomé (with Príncipe and Annobon), pp. 57-99. London.
    2. ALSTON, A. H. G., 1956. Pteridophyta. In A. Exell, Supplement to the Catalogue of the Vascular Plants of S.Tomé (with Príncipe and Annobon), pp. 7-9. London.
    3. ALSTON, A. H. G., 1958. Pteridophyta. In A. Exell & A. Rozeira, Aditamento à Flora das ilhas de São Tomé e Príncipe. Conferência Internacional dos Africanistas Ocidentais 3: pp.78- 81.
    4. ALSTON, A. H. G. (1959) Pteridophyta. In A. Exell, Additions to the Flora of S.Tomé and Príncipe. Bulletin de l’Institut Française d’Afrique Noire, série A, 21: pp. 440-448.
    5. ALSTON, A. H. G., 1959. The Ferns and Fern-allies of West Tropical Africa. London.
    6. BENL, G., 1978. The Pteridophyta of Fernando Po I. Acta Botanica Barcinonensia 31: pp. 1-31.
    7. BENL, G., 1980. The Pteridophyta of Fernando Po II. Acta Botanica Barcinonensia 32: 1-pp. 34.
    8. BENL, G., 1982. The Pteridophyta of Fernando Po III. Acta Botanica Barcinonensia 33: pp. 1-46.
    9. BENL, G., 1988. The Pteridophyta of Bioko (Fernando Po) IV. Acta Botanica Barcinonensia 38: pp. 1-69.
    10. BENL, G., 1991. The Pteridophyta of Bioko (Fernando Po) V. Acta Botanica Barcinonensia 40: pp.1-106.
    11. FIGUEIREDO, E., 1998. The Pteridophytes of São Tomé and Príncipe (Gulf of Guinea). Bulletin of the Natural History Museum, London (Botany) 28 (1): pp. 41-66.
    12. TARDIEU-BLOT, M.-L., 1953. Les Pteridophytes de l’Afrique Intertropicale Française. Mémoires de l’Institut Français d’Afrique Noire 28.
    13. TARDIEU-BLOT, M.-L., 1964. Ptéridophytes. In A. Aubréville (ed.) Flore du Gabon 8. Paris: Muséum National d’Histoire Naturelle.
    14. TARDIEU-BLOT, M.-L., 1964. Ptéridophytes. In A. Aubréville (ed.) Flore du Cameroun 3. Paris: Muséum National d’Histoire Naturelle.
  21. The Iwokrama Forest Research Centre, Guyana, South America. List of plant species identified by the Smithsonian Institute's "Biological Diversity of the Guianas" Program staff, as at 25 June 1999.
  22. The Kaieteur Plant Checklist from Smithsonian National Museum of Natural History: Biological Diversity of the Guianas - Guyana, Surinam, French Guiana; Kaieteur Falls Plant Checklist, August 2000, (compiled by Carol Kelloff, 1994; reviewed by D.B. Lellinger, 1997) at SNMNH

Distribution of ferns.

Information given in these pages on the distribution of ferns has come from the texts where I have access to them, information (if any) in the title where I have not, and from specific information in databases I have interrogated through the internet.

In the frequent case where several species are treated as synonyms and there are several names listed under the one preferred name, often some of the synonyms have associated information on the geographical distribution of plants to which that name has been applied. In such cases I have combined the location names of occurrence into one list.


I am in the process of learning a lot more detail about taxonomy.

It was my intention initially to avoid becoming too deeply involved in this process, but it has come clear to me that if these pages are to be of use to a wider audience, then I cannot duck the issue.

Because I have no access to the important specimens, have insufficient expertise, and too little time, I have to rely on the expert opinions of people who work in this field all the time; many bring a lifetime of careful, patient and conscientious work to the topic. I try to interpret the results of their work as accurately as I can, and am very grateful for their caring toil.

And I have just decided that this page is becoming too large, so to go to the separate page on taxonomy, Click here.


The first (botanic) name given to a plant is the basionym.   I have begun to underline these names where they are among the synonyms, as they often are. In such cases they are usually in another genus.

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